diff --git a/packages/platform-browser/test/testing_public_spec.ts b/packages/platform-browser/test/testing_public_spec.ts
index 4a9824ee82f75..c1bd7e47c0bfd 100644
--- a/packages/platform-browser/test/testing_public_spec.ts
+++ b/packages/platform-browser/test/testing_public_spec.ts
@@ -860,7 +860,7 @@ describe('public testing API', () => {
          })));
 
       itPromise.then(() => done.fail('Expected test to fail, but it did not'), (err) => {
-        expect(err.message).toEqual('Uncaught (in promise): baz');
+        expect(err.message).toEqual('baz');
         done();
       });
       restoreJasmineIt();
diff --git a/packages/zone.js/lib/common/promise.ts b/packages/zone.js/lib/common/promise.ts
index 215646a16459b..8f73655ed1275 100644
--- a/packages/zone.js/lib/common/promise.ts
+++ b/packages/zone.js/lib/common/promise.ts
@@ -23,7 +23,7 @@ Zone.__load_patch('ZoneAwarePromise', (global: any, Zone: ZoneType, api: _ZonePr
   const __symbol__ = api.symbol;
   const _uncaughtPromiseErrors: UncaughtPromiseError[] = [];
   const isDisableWrappingUncaughtPromiseRejection =
-      global[__symbol__('DISABLE_WRAPPING_UNCAUGHT_PROMISE_REJECTION')] === true;
+      global[__symbol__('DISABLE_WRAPPING_UNCAUGHT_PROMISE_REJECTION')] !== false;
   const symbolPromise = __symbol__('Promise');
   const symbolThen = __symbol__('then');
   const creationTrace = '__creationTrace__';
diff --git a/packages/zone.js/lib/zone.configurations.api.ts b/packages/zone.js/lib/zone.configurations.api.ts
index 6a602b8cfff57..93dd691870c37 100644
--- a/packages/zone.js/lib/zone.configurations.api.ts
+++ b/packages/zone.js/lib/zone.configurations.api.ts
@@ -576,11 +576,11 @@ interface ZoneGlobalConfigurations {
   /**
    * Disable wrapping uncaught promise rejection.
    *
-   * By default, `zone.js` wraps the uncaught promise rejection in a new `Error` object
-   * which contains additional information such as a value of the rejection and a stack trace.
+   * By default, `zone.js` throws the original error occurs in the uncaught promise rejection.
    *
-   * If you set `__zone_symbol__DISABLE_WRAPPING_UNCAUGHT_PROMISE_REJECTION = true;` before
-   * importing `zone.js`, `zone.js` will not wrap the uncaught promise rejection.
+   * If you set `__zone_symbol__DISABLE_WRAPPING_UNCAUGHT_PROMISE_REJECTION = false;` before
+   * importing `zone.js`, `zone.js` will wrap the uncaught promise rejection in a new `Error` object
+   * which contains additional information such as a value of the rejection and a stack trace.
    */
   __zone_symbol__DISABLE_WRAPPING_UNCAUGHT_PROMISE_REJECTION?: boolean;
 }
diff --git a/packages/zone.js/test/browser-zone-setup.ts b/packages/zone.js/test/browser-zone-setup.ts
index 264d797b7ac97..26c0cd2c6b268 100644
--- a/packages/zone.js/test/browser-zone-setup.ts
+++ b/packages/zone.js/test/browser-zone-setup.ts
@@ -9,6 +9,7 @@ if (typeof window !== 'undefined') {
   const zoneSymbol = (window as any).Zone.__symbol__;
   (window as any)['__Zone_enable_cross_context_check'] = true;
   (window as any)[zoneSymbol('fakeAsyncAutoFakeAsyncWhenClockPatched')] = true;
+  (window as any)[zoneSymbol('DISABLE_WRAPPING_UNCAUGHT_PROMISE_REJECTION')] = false;
 }
 import '../lib/common/to-string';
 import '../lib/browser/api-util';
diff --git a/packages/zone.js/test/browser_disable_wrap_uncaught_promise_rejection_setup.ts b/packages/zone.js/test/browser_disable_wrap_uncaught_promise_rejection_setup.ts
index 7665ce669189d..f2aef91cde2b3 100644
--- a/packages/zone.js/test/browser_disable_wrap_uncaught_promise_rejection_setup.ts
+++ b/packages/zone.js/test/browser_disable_wrap_uncaught_promise_rejection_setup.ts
@@ -5,5 +5,4 @@
  * Use of this source code is governed by an MIT-style license that can be
  * found in the LICENSE file at https://angular.io/license
  */
-
-(window as any)['__zone_symbol__DISABLE_WRAPPING_UNCAUGHT_PROMISE_REJECTION'] = true;
+// This is an empty karma test entry file just to keep the bazel macro happy.
diff --git a/packages/zone.js/test/browser_symbol_setup.ts b/packages/zone.js/test/browser_symbol_setup.ts
index 81cbcc163817d..b73003c47d2aa 100644
--- a/packages/zone.js/test/browser_symbol_setup.ts
+++ b/packages/zone.js/test/browser_symbol_setup.ts
@@ -1,3 +1,4 @@
 (window as any).global = window;
 // Change default symbol prefix for testing to ensure no hard-coded references.
 (window as any)['__Zone_symbol_prefix'] = '_test__';
+(window as any)['_test__DISABLE_WRAPPING_UNCAUGHT_PROMISE_REJECTION'] = false;
diff --git a/packages/zone.js/test/node-env-setup.ts b/packages/zone.js/test/node-env-setup.ts
index 77f6606835cd3..0bf818bbc9228 100644
--- a/packages/zone.js/test/node-env-setup.ts
+++ b/packages/zone.js/test/node-env-setup.ts
@@ -1,2 +1,3 @@
 // Change default symbol prefix for testing to ensure no hard-coded references.
 (global as any)['__Zone_symbol_prefix'] = '__zone_symbol_test__';
+(global as any)['__zone_symbol_test__DISABLE_WRAPPING_UNCAUGHT_PROMISE_REJECTION'] = false;