diff --git a/components/formats-bsd/src/loci/formats/tiff/TiffParser.java b/components/formats-bsd/src/loci/formats/tiff/TiffParser.java
index bfc4228e132..3aa9e2f07a0 100644
--- a/components/formats-bsd/src/loci/formats/tiff/TiffParser.java
+++ b/components/formats-bsd/src/loci/formats/tiff/TiffParser.java
@@ -725,6 +725,14 @@ else if (stripByteCounts[countIndex] < 0 && countIndex > 0) {
     in.seek(stripOffset);
     in.read(tile);
 
+    // reverse bits in each byte if FillOrder == 2
+
+    if (ifd.getIFDIntValue(IFD.FILL_ORDER) == 2) {
+      for (int i=0; i<tile.length; i++) {
+        tile[i] = (byte) (Integer.reverse(tile[i]) >> 24);
+      }
+    }
+
     codecOptions.maxBytes = (int) Math.max(size, tile.length);
     codecOptions.ycbcr =
       ifd.getPhotometricInterpretation() == PhotoInterp.Y_CB_CR &&
@@ -874,6 +882,7 @@ public byte[] getSamples(IFD ifd, byte[] buf, int x, int y,
       photoInterp != PhotoInterp.WHITE_IS_ZERO &&
       photoInterp != PhotoInterp.CMYK && photoInterp != PhotoInterp.Y_CB_CR &&
       compression == TiffCompression.UNCOMPRESSED &&
+      ifd.getIFDIntValue(IFD.FILL_ORDER) != 2 &&
       numTileRows * numTileCols == 1 && stripOffsets != null && stripByteCounts != null &&
       in.length() >= stripOffsets[0] + stripByteCounts[0])
     {