simgenome.eg

; insert_intron decreased to 0.0016 from 0.003
; extend_coding decreased to 0.9948 from 0.996
; end_coding decreased to 0.0004 from 0.001
; conserved increased to 0.15 from 0.085

;; The grammar.
(grammar
 (parametric)
 (name simgenome)
 (update-rates 0)  ;; turned off to prevent accidentally changing the grammar
 (update-rules 0)

 (meta
  (simgenome

   ;; This block can be used to patch in external feature-generators.
   ;; Here are a few examples:

   ;; columns annotated with any non-period character in the "#=GC GSIM" row will be replaced by the output of a call to gsimulator
   (gsimulator
    (row GSIM)
    )

   ;; columns annotated with any non-period character in the "#=GC GSIM100" row will be replaced by the output of a call to gsimulator, with the root sequence length constrained to be 100
   ;; This is the only external generator that is actually used in the current version of this model.
   (gsimulator
    (row GSIM100)
    (args --rootseqlength 100)
    )
   
   ;; columns annotated with any non-period character in the "#=GC EXT" row will be replaced by the output of the following Perl one-liner (the tree file is passed in as an argument but discarded)
   ;; more robustly, an external program name could be specified here, instead of "perl".
   (external
    (row EXT)
    (executable perl -e 'print"# STOCKHOLM 1.0\nA GGGGG\nB GGGGG\n//\n"')
    )

   ;; Below are descriptions of the various submodels in this grammar.
   ;; This info is not (as of 4/1/2008) used by any program, but sticking it in the "meta" block means that xrate won't delete it.

   (submodel-meta
    (submodel neutral)
    (brief-description Single-nucleotide strand-symmetric model of neutral evolution with gaps as 5th character.  Distribution and (base) rates from caf1screen/training/null_nuc_symmetric_gapsub_trained.eg.)
   )

   (submodel-meta
    (submodel conserved)
    (brief-description Single-nucleotide strand-symmetric ungapped model of evolution of conserved regions.  Distribution and rates from caf1screen/training/null_nuc_symmetric_trained.eg.)
   )

   (submodel-meta
    (submodel coding)
    (brief-description Coding sequence (codons).  UTRs modeled as conserved, splice sites as conserved and introns as neutral, albeit with an optionally different evolutionary rate scale, with optional conserved within.  Distribution and rates of coding sequence from simultaneous-mutations empirically-estimated model of Kosiol, Holmes and Goldman 2007.  Rates modified to allow any codon to mutate to all gaps (frame-preserving) at a fixed rate.)
   )

   (submodel-meta
    (submodel pseudogene)
    (brief-description Unprocessed pseudogenes.  Coding region only.  No UTRs or introns.  Codons generated from distribution of Kosiol, Holmes and Goldman 2007, then evolve as triplets of nucleotides, each of which evolves independently under neutral model.)
   )

   (submodel-meta
    (submodel ncrna)
    (brief-description Structural evolution of ncRNAs with gaps as 5th character.  Distribution and rates from GapSub (ncRna_v24.eg).)
   )

   (submodel-meta
    (submodel transposon)
    (brief-description DNA transposons with TIRs.  TIR distribution and rates identical to base-pairs in ncrna.  Inner sequence emitted as neutral, with exception of optional pseudogenes in inner region.)
   )

  )
 )




;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;
;; User-modified parameters ;;
;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;

 ;; Priors over length of simulated genome and frequencies of features.
 (const-pgroup

  ;; Parameters estimated (crudely) from Drosophila melanogaster.
  ;; Sources:
  ;;  FlyBase 4.3 annotation.
  ;;  "The transposable elements of the Drosophila melanogaster euchromatin: a genomics perspective" (2002), Genome Biology.
  ;;  "Evolution of genes and genomes on the Drosophila phylogeny" (2007), Nature 450, 203-218.
  ;;  "Minos as a Genetic and Genomic Tool in Drosophila melanogaster" (2005), Genetics 171(2), 571-581.
  ;;  "Minos, a new transposable element from Drosophila hydei, is a member of the Tc1-like family of transposons" (1991), Nucleic Acids Res., 19(23):6646.

  ;; Some numbers:
  ;; 130Mb euchromatic DNA
  ;; ~14,000 protein-coding genes (~31Mb exonic, ~54Mb intronic)
  ;; ~64,000 exons and ~48,000 introns
  ;; Mean exon length is ~500nt, standard deviation is ~600nt
  ;; Mean intron length is ~1,140nt, standard deviation is ~4,200nt
  ;; ~700 RNA genes
  ;; ~50 pseudogenes
  ;; 72 full-length and 332 partial transposons in the TIR and FB classes (which have inverted repeats)
  ;; ~6,000 transposons and ~20,000 "repeat regions"
  ;; ~1,400 "protein binding site" features (major overlap with "regulatory region" and "enhancer" features)
  ;; ~30% of intergenic sequence predicted conserved by PHASTCONS

  ;; Some calculations:
  ;; Approx ~45Mb (35%) of sequence is intergenic. This corresponds to ~450,000 GSIM100 states (each of which generates ~100 bases)
  ;; Features of interest:
  ;;  14,000 "coding"
  ;;      50 "pseudogene"
  ;;   1,400 "conserved" (protein binding sites)  ... but PHASTCONS puts this much higher ... we use PHASTCONS's estimate for introns, but not intergenic DNA (for which we have a better model).
  ;;     700 "ncrna"
  ;;     400 "transposon (including full-length AND partial TIR and foldback elements)

  ;;  16,550 total => roughly 0.037 per GSIM100 state

  ;; Mean size of Mercator syntenic block is 25kb. Assuming 35% of this is intergenic, this is 8,750nt of intergenic sequence, or ~88 GSIM100 states.
  ;; Hence, probabilities in NEUTRAL pgroup should be:
  ;;    insert_feature ~ 0.037
  ;; extend_intergenic ~ 0.952
  ;;        end_genome ~ 0.011

  ;; Relative proportions for FEATURE pgroup:
  ;;  0.85  "coding"
  ;;  0.003 "pseudogene"
  ;;  0.085 "conserved"
  ;;  0.04  "ncrna"
  ;;  0.024 "transposon"

  ;; Length distribution of exons:
  ;; Mean is ~500 (~166 codons), sd is ~600 (~200 codons)
  ;; Sum of r geometric variables with (stopping) parameter p is negative binomial distribution with parameters {r,p}.
  ;; Mean of negative binomial is r(1-p)/p = 166
  ;; Variance is r(1-p)/(p^2) = 200^2
  ;; Thus p = 166/200^2 ~= .004
  ;;  and r = 166*p/(1-p) ~= 0.67  !!???

  ;; That r~1 is confirmed by the following crude gridsearch over negative binomial parameter space:
  ;; cat exon.gff | perl -ane 'print $F[4]-$F[3],"\n"' | linbins 50 | perl -e 'while(&<>){($n,$c)=split;$c{$n}=$c}for($p=.001;$p<=.005;$p+=.001){for($r=1;$r<=5;++$r){$ll=0;while(($n,$c)=each%c){$ll+=$c*($r*log($p)+$n*log(1-$p)+logfac($n+$r)-logfac($n)-logfac($r))}print"p= $p r= $r ll= $ll\n"}}sub logfac{my$n=shift;my$ll=0;for my$i(2..$n){$ll+=log($i)}return $ll}'

  ;; There are roughly 0.75 introns per exon (&=47676/64113)
  ;;       and roughly 4.4    exons per gene (&=64113/14449)

  ;; Thus,
  ;;  NUMBER_OF_CODING_STATES ~ 1
  ;;            extend_coding ~ .996
  ;;            insert_intron ~ .003
  ;;               end_coding ~ .001

  ;; Gap rate in coding regions is chosen heuristically.

  ;; Average length of protein binding site is ~20nt
  ;;  extend_conserved ~ 0.95
  ;;     end_conserved ~ 0.05

  ;; PHASTCONS reports the slowest-substituting 10% of regions:
  ;;  conserved_rate_scale ~ 0.1

  ;; PHASTCONS says that 23% of intronic DNA is conserved
  ;; Mean length of an intron is 1,140nt; 23% of this is ~260nt
  ;; That is, roughly 13 "conserved" features per 1000nt of intron

  ;; So, INTRON params are as follows:
  ;;  insert_conserved ~ 0.013
  ;;  extend_intron    ~ 0.986
  ;;     end_intron    ~ 0.001

  ;; Transposon TIR lengths and gene counts: hard to find comprehensive statistics.
  ;; Take the "minos" element as a canonical example.
  ;; Total length is 1751nt; TIR length is 255nt. Contains one gene of length 1036nt.

  ;; So for minos, TRANSPOSON params should be as follows:
  ;;      extend_tir ~ 0.996
  ;;         end_tir ~ 0.004

  ;;    extend_inner ~ 0.996
  ;; transposon_gene ~ 0.002
  ;;       end_inner ~ 0.002

   ;; NEUTRAL
   ((insert_feature 0.045)    ;; Prior over all features.
    (extend_intergenic 0.944) ;; Geometric parameter for length distribution of simulated genome. (Mean intergenic distance is approx 100/extend_intergenic bases; grep for GSIM100 to see where the factor of 100 comes from)
    (end_genome 0.011))

   ;; FEATURE
   ((coding .888)             ;; Priors over particular features.
    (pseudogene .003)
    (conserved .15)
    (ncrna 0)
    (transposon .024))

 )

 ;; number of coding states
 ;; This effectively corresponds to the "shape" parameter
 ;; in the gamma distribution over exon lengths.
 ;; (Technically, it's a "negative binomial" distribution, not a gamma.
 ;;  The negative binomial is the discrete analogue of the gamma,
 ;;  obtained in this case as a convolution of identical geometric distrib'ns.)

 ;; Despite IH's intuition that this should be a peaked distribution (&>1), empirically a geometric distribution (&=1) seems to be a better fit.
 (&define NUMBER_OF_CODING_STATES 1)

 ;; Length of features.
 (const-pgroup

   ;; CONSERVED
   ((extend_conserved 0.95) ;; Length distribution of conserved regions.
    (end_conserved 0.05))

   ;; CODING
   ((insert_intron 0.0016)     ;; Frequency of introns per codon.
    (extend_coding 0.9948)    ;; Length distribution of coding regions.
                ;; (this is the "scale" parameter in the gamma distribution)
    (end_coding 0.0004))

   ;; INTRON
   ((insert_conserved 0.013)  ;; Prior over conserved regions within introns.
    (extend_intron 0.986)     ;; Length distribution of introns.
    (end_intron 0.001))

   ;; TRANSPOSON
   ((extend_tir 0.996)       ;; Length distribution of terminal inverted repeats (TIRs).
    (end_tir 0.004))
   ((transposon_gene 0.002)  ;; Prior over genes in transposons.
    (extend_inner 0.996)     ;; Length distribution of inner region.
    (end_inner 0.002))

 )

 ;; Rates of mutation of features.
 (const-rate

   ;; NEUTRAL
   (neutral_rate_scale 1)       

   ;; INTRON
   (intron_rate_scale 1)      

   ;; CONSERVED
   (conserved_rate_scale 0.1)

   ;; CODING
   (coding_rate_scale 1)       
   (coding_gap_rate .1)        ;; Rate of (any codon) -> (all gaps) mutation.

   ;; NCRNA
   (ncrna_rate_scale 1)

 )







;!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!
;! WARNING: The below should only be modified by expert users !
;! See http://biowiki.org/XrateFormat for format docs...      !
;!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!






;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;
;; Nonterminal declarations ;;
;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;

 (nonterminal (name START))
 (nonterminal (name NEUTRAL)) ;; gsimulator-generated neutral regions (rates also used to evolve pseudogenes and transposons)
 (nonterminal (name INSERT_FEATURE)) ;; bifurcation to generate feature as (FEATURE NEUTRAL)
 (nonterminal (name FEATURE)) ;; generic genomic feature
 (nonterminal (name CODING)) ;; protein-coding gene
 (nonterminal (name PSEUDOGENE)) ;; unprocessed pseudogene
 (nonterminal (name NCRNA)) ;; ncRNA
 (nonterminal (name TRANSPOSON)) ;; transposon
 (nonterminal (name CONSERVED)) ;; conserved region (ungapped neutral strand-symmetric evolution)

 ;; CODING
 (nonterminal (name FWD_START_CODING))
 (nonterminal (name REV_START_CODING))

 ;; INTRON
 (nonterminal (name FWD_INSERT_INTRON))
 (nonterminal (name REV_INSERT_INTRON))
 (nonterminal (name START_INTRON))
 (nonterminal (name INTRON))

 ;; PSEUDOGENE
 (nonterminal (name FWD_PSEUDOGENE))
 (nonterminal (name REV_PSEUDOGENE))

 ;; NCRNA
 (nonterminal (name FWD_NCRNA_PFOLD_S))
 (nonterminal (name REV_NCRNA_PFOLD_S))
 (nonterminal (name FWD_NCRNA_PFOLD_F))
 (nonterminal (name REV_NCRNA_PFOLD_F))
 (nonterminal (name FWD_NCRNA_PFOLD_L))
 (nonterminal (name REV_NCRNA_PFOLD_L))
 (nonterminal (name FWD_NCRNA_PFOLD_B))
 (nonterminal (name REV_NCRNA_PFOLD_B))
 (nonterminal (name FWD_NCRNA_PFOLD_U))
 (nonterminal (name REV_NCRNA_PFOLD_U))

 ;; TRANSPOSON
 (nonterminal (name FWD_TRANSPOSON))
 (nonterminal (name REV_TRANSPOSON))
 (nonterminal (name FWD_TRANSPOSON_TIR))
 (nonterminal (name REV_TRANSPOSON_TIR))
 (nonterminal (name FWD_TRANSPOSON_INNER))
 (nonterminal (name REV_TRANSPOSON_INNER))
 (nonterminal (name FWD_TRANSPOSON_INSERT_PSEUDOGENE))
 (nonterminal (name REV_TRANSPOSON_INSERT_PSEUDOGENE))


;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;
;; Tranformations (NEUTRAL and FEATURE) ;;
;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;

 ;; state START
 (transform (from (START)) (to (NEUTRAL)) (prob 1))

 ;; state NEUTRAL  .. generates 100 nucleotides of ancestral sequence, which is then mutated by GSIMULATOR
 (transform (from (NEUTRAL)) (to (X NEUTRAL*))
 	    (annotate (row GSIM100) (column X) (label 1)))
 (transform (from (NEUTRAL*)) (to (INSERT_FEATURE)) (prob insert_feature))
 (transform (from (NEUTRAL*)) (to (NEUTRAL)) (prob extend_intergenic))
 (transform (from (NEUTRAL*)) (to ()) (prob end_genome))

 ;; state INSERT_FEATURE
 (transform (from (INSERT_FEATURE)) (to (FEATURE NEUTRAL)))

 ;; state FEATURE
 (transform (from (FEATURE)) (to (CONSERVED)) (prob conserved))
 (transform (from (FEATURE)) (to (CODING)) (prob coding))
 (transform (from (FEATURE)) (to (PSEUDOGENE)) (prob pseudogene))
 (transform (from (FEATURE)) (to (NCRNA)) (prob ncrna))
 (transform (from (FEATURE)) (to (TRANSPOSON)) (prob transposon))


;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;
;; Tranformations (CONSERVED) ;;
;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;

 ;; state CONSERVED
 (transform (from (CONSERVED)) (to (CONS CONSERVED*))
  (annotate (row CONSERVED) (column CONS) (label C)))
 (transform (from (CONSERVED*)) (to (CONSERVED)) (prob extend_conserved))
 (transform (from (CONSERVED*)) (to ()) (prob end_conserved))


;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;
;; Transformations (CODING) ;;
;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;

 ;; state CODING
 (transform (from (CODING)) (to (FWD_START_CODING)) (prob 0.5))
 (transform (from (CODING)) (to (REV_START_CODING)) (prob 0.5))

 ;; state FWD_START_CODING, REV_START_CODING
 (transform (from (FWD_START_CODING)) (to (CONSERVED FWD_CODING_1))) ;; start coding region with conserved sequence
 (transform (from (REV_START_CODING)) (to (CONSERVED REV_CODING_1)))

 ;; {FWD,REV}_CODING_* states
 (&foreach-integer
  NUM
  (1 NUMBER_OF_CODING_STATES)

  (nonterminal (name (&. FWD_CODING_ NUM)))
  (nonterminal (name (&. REV_CODING_ NUM)))

  ;; state FWD_CODING_[NUM]
  (transform (from ((&. FWD_CODING_ NUM)))
	     (to (COD1 COD2 COD3 (&. FWD_CODING_ NUM *)))
	     (gaps-ok)
	     (annotate (row CODING) (column COD1) (label 0))
	     (annotate (row CODING) (column COD2) (label 1))
	     (annotate (row CODING) (column COD3) (label 2))
	     (annotate (row STRAND) (column COD1) (label +))
	     (annotate (row STRAND) (column COD2) (label +))
	     (annotate (row STRAND) (column COD3) (label +)))

  (transform (from ((&. FWD_CODING_ NUM *)))
	     (to ((&. FWD_CODING_ NUM)))
	     (prob extend_coding))

  (transform (from ((&. FWD_CODING_ NUM *)))
	     (to (FWD_INSERT_INTRON))
	     (prob insert_intron))

  (transform (from ((&. FWD_CODING_ NUM *)))
	     (to ((&?
		   (&= NUM NUMBER_OF_CODING_STATES)
		   CONSERVED
		   (&. FWD_CODING_ (&+ NUM 1)))))
	     (prob end_coding))

  ;; state REV_CODING_[NUM]
  (transform (from ((&. REV_CODING_ NUM)))
	     (to (~COD3 ~COD2 ~COD1 (&. REV_CODING_ NUM *)))
	     (gaps-ok)
	     (annotate (row CODING) (column COD3) (label 2))
	     (annotate (row CODING) (column COD2) (label 1))
	     (annotate (row CODING) (column COD1) (label 0))
	     (annotate (row STRAND) (column COD3) (label -))
	     (annotate (row STRAND) (column COD2) (label -))
	     (annotate (row STRAND) (column COD1) (label -)))

  (transform (from ((&. REV_CODING_ NUM *)))
	     (to ((&. REV_CODING_ NUM)))
	     (prob extend_coding))

  (transform (from ((&. REV_CODING_ NUM *)))
	     (to (REV_INSERT_INTRON))
	     (prob insert_intron))

  (transform (from ((&. REV_CODING_ NUM *)))
	     (to ((&?
		   (&= NUM NUMBER_OF_CODING_STATES)
		   CONSERVED
		   (&. REV_CODING_ (&+ NUM 1)))))
	     (prob end_coding))

  ;; end of coding state generation macro
  )

 ;; state FWD_INSERT_INTRON, REV_INSERT_INTRON
 (transform (from (FWD_INSERT_INTRON)) (to (START_INTRON FWD_CODING_1)))
 (transform (from (REV_INSERT_INTRON)) (to (START_INTRON REV_CODING_1)))

 ;; state START_INTRON
 (transform (from (START_INTRON)) (to (CONSERVED INTRON))) ;; start intronic region with conserved sequence

 ;; state INTRON
 (transform (from (INTRON)) (to (XINTRON INTRON*))
  (annotate (row CODING) (column XINTRON) (label I))
  (annotate (row STRAND) (column XINTRON) (label _)))
 (transform (from (INTRON*)) (to (INTRON)) (prob extend_intron))
 (transform (from (INTRON*)) (to (INSERT_CONSERVED)) (prob insert_conserved)) ;; insert conserved sequence into middle of intron
 (transform (from (INTRON*)) (to (CONSERVED)) (prob end_intron)) ;; end intron with conserved sequence

 ;; state INSERT_CONSERVED
 (transform (from (INSERT_CONSERVED)) (to (CONSERVED INTRON))) 


;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;
;; Transformations (PSEUDOGENE) ;;
;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;

 ;; state CODING
 (transform (from (PSEUDOGENE)) (to (FWD_PSEUDOGENE)) (prob 0.5))
 (transform (from (PSEUDOGENE)) (to (REV_PSEUDOGENE)) (prob 0.5))

 ;; state FWD_PSEUDOGENE
 (transform (from (FWD_PSEUDOGENE)) (to (PG1 PG2 PG3 FWD_PSEUDOGENE*))
  (annotate (row PSEUDOGENE) (column PG1) (label 0))
  (annotate (row PSEUDOGENE) (column PG2) (label 1))
  (annotate (row PSEUDOGENE) (column PG3) (label 2))
  (annotate (row STRAND) (column PG1) (label +))
  (annotate (row STRAND) (column PG2) (label +))
  (annotate (row STRAND) (column PG3) (label +)))
 (transform (from (FWD_PSEUDOGENE*)) (to (FWD_PSEUDOGENE)) (prob extend_coding))
 (transform (from (FWD_PSEUDOGENE*)) (to ()) (prob end_coding))

 ;; state REV_PSEUDOGENE
 (transform (from (REV_PSEUDOGENE)) (to (~PG3 ~PG2 ~PG1 REV_PSEUDOGENE*)) (gaps-ok)
  (annotate (row PSEUDOGENE) (column PG3) (label 2))
  (annotate (row PSEUDOGENE) (column PG2) (label 1))
  (annotate (row PSEUDOGENE) (column PG1) (label 0))
  (annotate (row STRAND) (column PG3) (label -))
  (annotate (row STRAND) (column PG2) (label -))
  (annotate (row STRAND) (column PG1) (label -)))
 (transform (from (REV_PSEUDOGENE*)) (to (REV_PSEUDOGENE)) (prob extend_coding))
 (transform (from (REV_PSEUDOGENE*)) (to ()) (prob end_coding))

;;;;;;;;;;;;;;;;;;;;;;;;;;;;;
;; Transformations (NCRNA) ;;
;;;;;;;;;;;;;;;;;;;;;;;;;;;;;

 ;; state NCRNA
 (transform (from (NCRNA)) (to (FWD_NCRNA_PFOLD_S)) (prob 0.5))
 (transform (from (NCRNA)) (to (REV_NCRNA_PFOLD_S)) (prob 0.5))

 ;; state 10: FWD_NCRNA_PFOLD_S = start state for FWD_NCRNA_PFOLD_ subgrammar (fwd)
 (transform (from (FWD_NCRNA_PFOLD_S)) (to (FWD_NCRNA_PFOLD_L)) (prob Trans_S2L))
 (transform (from (FWD_NCRNA_PFOLD_S)) (to (FWD_NCRNA_PFOLD_B)) (prob Trans_S2B))

 ;; state 11: REV_NCRNA_PFOLD_S = start state for REV_NCRNA_PFOLD_ subgrammar (rev)
 (transform (from (REV_NCRNA_PFOLD_S)) (to (REV_NCRNA_PFOLD_L)) (prob Trans_S2L))
 (transform (from (REV_NCRNA_PFOLD_S)) (to (REV_NCRNA_PFOLD_B)) (prob Trans_S2B))

 ;; state 12: FWD_NCRNA_PFOLD_F = emit state for paired bases (fwd)
 (transform (from (FWD_NCRNA_PFOLD_F)) (to (LNUC FWD_NCRNA_PFOLD_F* RNUC))
  (annotate (row NCRNA) (column LNUC) (label <))
  (annotate (row NCRNA) (column RNUC) (label >))
  (annotate (row STRAND) (column LNUC) (label +))
  (annotate (row STRAND) (column RNUC) (label +)))
 (transform (from (FWD_NCRNA_PFOLD_F*)) (to (FWD_NCRNA_PFOLD_F)) (prob Trans_F2F))
 (transform (from (FWD_NCRNA_PFOLD_F*)) (to (FWD_NCRNA_PFOLD_B)) (prob Trans_F2B))

 ;; state 13: REV_NCRNA_PFOLD_F = emit state for paired bases (rev)
 (transform (from (REV_NCRNA_PFOLD_F)) (to (~RNUC REV_NCRNA_PFOLD_F* ~LNUC))
  (annotate (row NCRNA) (column RNUC) (label <))
  (annotate (row NCRNA) (column LNUC) (label >))
  (annotate (row STRAND) (column RNUC) (label -))
  (annotate (row STRAND) (column LNUC) (label -)))
 (transform (from (REV_NCRNA_PFOLD_F*)) (to (REV_NCRNA_PFOLD_F)) (prob Trans_F2F))
 (transform (from (REV_NCRNA_PFOLD_F*)) (to (REV_NCRNA_PFOLD_B)) (prob Trans_F2B))

 ;; state 14: FWD_NCRNA_PFOLD_L = transition between paired and unpaired sites (fwd)
 (transform (from (FWD_NCRNA_PFOLD_L)) (to (FWD_NCRNA_PFOLD_F)) (prob Trans_L2F))
 (transform (from (FWD_NCRNA_PFOLD_L)) (to (FWD_NCRNA_PFOLD_U)) (prob Trans_L2U))

 ;; state 15: REV_NCRNA_PFOLD_L = transition between paired and unpaired sites (rev)
 (transform (from (REV_NCRNA_PFOLD_L)) (to (REV_NCRNA_PFOLD_F)) (prob Trans_L2F))
 (transform (from (REV_NCRNA_PFOLD_L)) (to (REV_NCRNA_PFOLD_U)) (prob Trans_L2U))

 ;; state 16: FWD_NCRNA_PFOLD_B = bifurcation (fwd)
 (transform (from (FWD_NCRNA_PFOLD_B)) (to (FWD_NCRNA_PFOLD_L FWD_NCRNA_PFOLD_S)))

 ;; state 17: REV_NCRNA_PFOLD_B = bifurcation (rev)
 (transform (from (REV_NCRNA_PFOLD_B)) (to (REV_NCRNA_PFOLD_S REV_NCRNA_PFOLD_L)))

 ;; state 18: FWD_NCRNA_PFOLD_U = emit state for unpaired bases (fwd)
 (transform (from (FWD_NCRNA_PFOLD_U)) (to (FWD_NCRNA_PFOLD_U* NUC))
  (annotate (row NCRNA) (column NUC) (label _))
  (annotate (row STRAND) (column NUC) (label +)))
 (transform (from (FWD_NCRNA_PFOLD_U*)) (to ()) (prob 1))

 ;; state 19: REV_NCRNA_PFOLD_U = emit state for unpaired bases (rev)
 (transform (from (REV_NCRNA_PFOLD_U)) (to (~NUC REV_NCRNA_PFOLD_U*))
  (annotate (row NCRNA) (column NUC) (label _))
  (annotate (row STRAND) (column NUC) (label -)))
 (transform (from (REV_NCRNA_PFOLD_U*)) (to ()) (prob 1))

 
;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;
;; Transformations (TRANSPOSON) ;;
;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;;

 ;; state TRANSPOSON
 (transform (from (TRANSPOSON)) (to (FWD_TRANSPOSON_TIR)) (prob 0.5))
 (transform (from (TRANSPOSON)) (to (REV_TRANSPOSON_TIR)) (prob 0.5))

 ;; state 12: FWD_TRANSPOSON_TIR = emit state for paired bases (fwd)
 (transform (from (FWD_TRANSPOSON_TIR)) (to (LTIR FWD_TRANSPOSON_TIR* RTIR))
  (annotate (row TRANSPOSON) (column LTIR) (label <))
  (annotate (row TRANSPOSON) (column RTIR) (label >))
  (annotate (row STRAND) (column LTIR) (label +))
  (annotate (row STRAND) (column RTIR) (label +)))
 (transform (from (FWD_TRANSPOSON_TIR*)) (to (FWD_TRANSPOSON_TIR)) (prob extend_tir))
 (transform (from (FWD_TRANSPOSON_TIR*)) (to (FWD_TRANSPOSON_INNER)) (prob end_tir))

 ;; state 13: REV_TRANSPOSON_TIR = emit state for paired bases (rev)
 (transform (from (REV_TRANSPOSON_TIR)) (to (~RTIR REV_TRANSPOSON_TIR* ~LTIR))
  (annotate (row TRANSPOSON) (column RTIR) (label <))
  (annotate (row TRANSPOSON) (column LTIR) (label >))
  (annotate (row STRAND) (column RTIR) (label -))
  (annotate (row STRAND) (column LTIR) (label -)))
 (transform (from (REV_TRANSPOSON_TIR*)) (to (REV_TRANSPOSON_TIR)) (prob extend_tir))
 (transform (from (REV_TRANSPOSON_TIR*)) (to (REV_TRANSPOSON_INNER)) (prob end_tir))

 ;; state 18: FWD_TRANSPOSON_INNER = emit state for unpaired bases (fwd)
 (transform (from (FWD_TRANSPOSON_INNER)) (to (FWD_TRANSPOSON_INNER* X))
  (annotate (row TRANSPOSON) (column X) (label _))
  (annotate (row STRAND) (column X) (label +)))
 (transform (from (FWD_TRANSPOSON_INNER*)) (to (FWD_TRANSPOSON_INNER)) (prob extend_inner))
 (transform (from (FWD_TRANSPOSON_INNER*)) (to (FWD_TRANSPOSON_INSERT_PSEUDOGENE)) (prob transposon_gene))
 (transform (from (FWD_TRANSPOSON_INNER*)) (to ()) (prob end_inner))

 (transform (from (FWD_TRANSPOSON_INSERT_PSEUDOGENE)) (to (FWD_PSEUDOGENE FWD_TRANSPOSON_INNER)))

 ;; state 19: REV_TRANSPOSON_INNER = emit state for unpaired bases (rev)
 (transform (from (REV_TRANSPOSON_INNER)) (to (~X REV_TRANSPOSON_INNER*))
  (annotate (row TRANSPOSON) (column X) (label _))
  (annotate (row STRAND) (column X) (label -)))
 (transform (from (REV_TRANSPOSON_INNER*)) (to (REV_TRANSPOSON_INNER)) (prob extend_inner))
 (transform (from (REV_TRANSPOSON_INNER*)) (to (REV_TRANSPOSON_INSERT_PSEUDOGENE)) (prob transposon_gene))
 (transform (from (REV_TRANSPOSON_INNER*)) (to ()) (prob end_inner))

 (transform (from (REV_TRANSPOSON_INSERT_PSEUDOGENE)) (to (REV_PSEUDOGENE REV_TRANSPOSON_INNER)))





;; parameters

;;;;;;;;;;;;;
;; NEUTRAL ;;
;;;;;;;;;;;;;

;; Gapsub neutral model
;; Taken from caf1screen/training/null_nuc_symmetric_gapsub_trained.eg.
;; NB: Some parameters renamed for consistency:
;;  (X_g2gap_c2gap 0.929805) -> (X_c2gap_g2gap 0.929805)
;;  (X_gap2g_gap2c 0.197373) -> (X_gap2c_gap2g 0.197373)

 ;; constant probability parameters
 (const-pgroup
  ((X_at 0.0947323)
   (X_gc 0.0828704)
   (X_gap 0.82302))
  )  ;; end pgroup

 ;; constant rate parameters
 (const-rate
  (X_a2c_t2g 0.125521)
  (X_a2g_t2c 0.302289)
  (X_a2t_t2a 0.168872)
  (X_c2a_g2t 0.204476)
  (X_c2g_g2c 0.186986)
  (X_c2t_g2a 0.483303)
  (X_a2gap_t2gap 0.971307)
  (X_c2gap_g2gap 0.929805)
  (X_gap2a_gap2t 0.311866)
  (X_gap2c_gap2g 0.197373)
  )  ;; end rate

;;;;;;;;;;;;;;;
;; CONSERVED ;;
;;;;;;;;;;;;;;;

;; Ungapped neutral model
;; Taken from caf1screen/training/null_nuc_symmetric_trained.eg.

 ;; constant probability parameters
 (const-pgroup
  ((CONS_at 0.595016)
   (CONS_gc 0.405283))
  )  ;; end pgroup

 ;; constant rate parameters
 (const-rate
  (CONS_a2c_t2g 0.176654)
  (CONS_a2g_t2c 0.352086)
  (CONS_a2t_t2a 0.247414)
  (CONS_c2a_g2t 0.265908)
  (CONS_c2g_g2c 0.2111)
  (CONS_c2t_g2a 0.529976)
  )  ;; end rate



;;;;;;;;;;;;
;; CODING ;;
;;;;;;;;;;;;

 ;; constant probability parameters
 (const-pgroup
  ((COD1_COD2_COD3_aaa 0.0310987)
   (COD1_COD2_COD3_aac 0.0203335)
   (COD1_COD2_COD3_aag 0.0267003)
   (COD1_COD2_COD3_aat 0.0222817)
   (COD1_COD2_COD3_aca 0.0131208)
   (COD1_COD2_COD3_acc 0.0178863)
   (COD1_COD2_COD3_acg 0.010687)
   (COD1_COD2_COD3_act 0.0126563)
   (COD1_COD2_COD3_aga 0.00975258)
   (COD1_COD2_COD3_agc 0.0137065)
   (COD1_COD2_COD3_agg 0.00679175)
   (COD1_COD2_COD3_agt 0.0103158)
   (COD1_COD2_COD3_ata 0.0128152)
   (COD1_COD2_COD3_atc 0.0237653)
   (COD1_COD2_COD3_atg 0.0211823)
   (COD1_COD2_COD3_att 0.0247745)
   (COD1_COD2_COD3_caa 0.0171696)
   (COD1_COD2_COD3_cac 0.0110409)
   (COD1_COD2_COD3_cag 0.020732)
   (COD1_COD2_COD3_cat 0.0106722)
   (COD1_COD2_COD3_cca 0.0111718)
   (COD1_COD2_COD3_ccc 0.0104092)
   (COD1_COD2_COD3_ccg 0.0121998)
   (COD1_COD2_COD3_cct 0.0104308)
   (COD1_COD2_COD3_cga 0.00480916)
   (COD1_COD2_COD3_cgc 0.014609)
   (COD1_COD2_COD3_cgg 0.00816122)
   (COD1_COD2_COD3_cgt 0.008496)
   (COD1_COD2_COD3_cta 0.0073604)
   (COD1_COD2_COD3_ctc 0.0168046)
   (COD1_COD2_COD3_ctg 0.0284979)
   (COD1_COD2_COD3_ctt 0.0151767)
   (COD1_COD2_COD3_gaa 0.0345301)
   (COD1_COD2_COD3_gac 0.0252951)
   (COD1_COD2_COD3_gag 0.0306069)
   (COD1_COD2_COD3_gat 0.0285968)
   (COD1_COD2_COD3_gca 0.0165274)
   (COD1_COD2_COD3_gcc 0.0268302)
   (COD1_COD2_COD3_gcg 0.0183002)
   (COD1_COD2_COD3_gct 0.0189193)
   (COD1_COD2_COD3_gga 0.0163905)
   (COD1_COD2_COD3_ggc 0.0236667)
   (COD1_COD2_COD3_ggg 0.0102233)
   (COD1_COD2_COD3_ggt 0.0168863)
   (COD1_COD2_COD3_gta 0.0109267)
   (COD1_COD2_COD3_gtc 0.0184274)
   (COD1_COD2_COD3_gtg 0.0226397)
   (COD1_COD2_COD3_gtt 0.0190245)
   (COD1_COD2_COD3_taa 0)
   (COD1_COD2_COD3_tac 0.0167001)
   (COD1_COD2_COD3_tag 0)
   (COD1_COD2_COD3_tat 0.0172412)
   (COD1_COD2_COD3_tca 0.0103732)
   (COD1_COD2_COD3_tcc 0.0107613)
   (COD1_COD2_COD3_tcg 0.00872276)
   (COD1_COD2_COD3_tct 0.0118006)
   (COD1_COD2_COD3_tga 0)
   (COD1_COD2_COD3_tgc 0.00741674)
   (COD1_COD2_COD3_tgg 0.0127443)
   (COD1_COD2_COD3_tgt 0.00644323)
   (COD1_COD2_COD3_tta 0.0161985)
   (COD1_COD2_COD3_ttc 0.0213592)
   (COD1_COD2_COD3_ttg 0.0157228)
   (COD1_COD2_COD3_ttt 0.0214185))
  )  ;; end const-pgroup

 ;; constant rate parameters
 (const-rate
  (COD1_COD2_COD3_aaa2aac 0.00841396)
  (COD1_COD2_COD3_aaa2aag 0.345319)
  (COD1_COD2_COD3_aaa2aat 0.0462309)
  (COD1_COD2_COD3_aaa2aca 0.019998)
  (COD1_COD2_COD3_aaa2acc 0.00160079)
  (COD1_COD2_COD3_aaa2acg 0.00213285)
  (COD1_COD2_COD3_aaa2act 0.0111177)
  (COD1_COD2_COD3_aaa2aga 0.0566806)
  (COD1_COD2_COD3_aaa2agc 0.00458139)
  (COD1_COD2_COD3_aaa2agg 0.0126826)
  (COD1_COD2_COD3_aaa2agt 0.0133409)
  (COD1_COD2_COD3_aaa2ata 0.0085551)
  (COD1_COD2_COD3_aaa2atc 0.000547065)
  (COD1_COD2_COD3_aaa2atg 0.00583521)
  (COD1_COD2_COD3_aaa2att 0.00548611)
  (COD1_COD2_COD3_aaa2caa 0.0552455)
  (COD1_COD2_COD3_aaa2cac 0.00158533)
  (COD1_COD2_COD3_aaa2cag 0.00591667)
  (COD1_COD2_COD3_aaa2cat 0.00959834)
  (COD1_COD2_COD3_aaa2cca 0.0116462)
  (COD1_COD2_COD3_aaa2ccc 0.000128673)
  (COD1_COD2_COD3_aaa2ccg 4.6997e-06)
  (COD1_COD2_COD3_aaa2cct 0.00257895)
  (COD1_COD2_COD3_aaa2cga 0.0147514)
  (COD1_COD2_COD3_aaa2cgc 0.00432825)
  (COD1_COD2_COD3_aaa2cgg 9.04199e-05)
  (COD1_COD2_COD3_aaa2cgt 0.0150301)
  (COD1_COD2_COD3_aaa2cta 0.00319272)
  (COD1_COD2_COD3_aaa2ctc 3.61912e-05)
  (COD1_COD2_COD3_aaa2ctg 8.47079e-07)
  (COD1_COD2_COD3_aaa2ctt 0.000924525)
  (COD1_COD2_COD3_aaa2gaa 0.0780207)
  (COD1_COD2_COD3_aaa2gac 0.00134517)
  (COD1_COD2_COD3_aaa2gag 0.00487629)
  (COD1_COD2_COD3_aaa2gat 0.017301)
  (COD1_COD2_COD3_aaa2gca 0.0152611)
  (COD1_COD2_COD3_aaa2gcc 0.000147295)
  (COD1_COD2_COD3_aaa2gcg 8.66722e-06)
  (COD1_COD2_COD3_aaa2gct 0.0040891)
  (COD1_COD2_COD3_aaa2gga 0.0117028)
  (COD1_COD2_COD3_aaa2ggc 6.80993e-05)
  (COD1_COD2_COD3_aaa2ggg 1.19552e-05)
  (COD1_COD2_COD3_aaa2ggt 0.000822897)
  (COD1_COD2_COD3_aaa2gta 0.00773705)
  (COD1_COD2_COD3_aaa2gtc 3.56683e-05)
  (COD1_COD2_COD3_aaa2gtg 4.45853e-06)
  (COD1_COD2_COD3_aaa2gtt 0.00383382)
  (COD1_COD2_COD3_aaa2taa 0)
  (COD1_COD2_COD3_aaa2tac 0.00071687)
  (COD1_COD2_COD3_aaa2tag 0)
  (COD1_COD2_COD3_aaa2tat 0.00805444)
  (COD1_COD2_COD3_aaa2tca 0.0155171)
  (COD1_COD2_COD3_aaa2tcc 0.000224967)
  (COD1_COD2_COD3_aaa2tcg 4.07435e-06)
  (COD1_COD2_COD3_aaa2tct 0.00402443)
  (COD1_COD2_COD3_aaa2tga 0)
  (COD1_COD2_COD3_aaa2tgc 3.15281e-05)
  (COD1_COD2_COD3_aaa2tgg 0.000649672)
  (COD1_COD2_COD3_aaa2tgt 0.000641172)
  (COD1_COD2_COD3_aaa2tta 0.00992305)
  (COD1_COD2_COD3_aaa2ttc 0.000128406)
  (COD1_COD2_COD3_aaa2ttg 7.84428e-05)
  (COD1_COD2_COD3_aaa2ttt 0.00317506)
  (COD1_COD2_COD3_aac2aaa 0.0128775)
  (COD1_COD2_COD3_aac2aag 0.0277379)
  (COD1_COD2_COD3_aac2aat 0.392836)
  (COD1_COD2_COD3_aac2aca 0.00341003)
  (COD1_COD2_COD3_aac2acc 0.0240803)
  (COD1_COD2_COD3_aac2acg 0.00878343)
  (COD1_COD2_COD3_aac2act 0.00374972)
  (COD1_COD2_COD3_aac2aga 0.0026699)
  (COD1_COD2_COD3_aac2agc 0.0565629)
  (COD1_COD2_COD3_aac2agg 0.00407778)
  (COD1_COD2_COD3_aac2agt 0.00604937)
  (COD1_COD2_COD3_aac2ata 0.000267162)
  (COD1_COD2_COD3_aac2atc 0.00417213)
  (COD1_COD2_COD3_aac2atg 0.00464212)
  (COD1_COD2_COD3_aac2att 0.000629284)
  (COD1_COD2_COD3_aac2caa 0.00648355)
  (COD1_COD2_COD3_aac2cac 0.024332)
  (COD1_COD2_COD3_aac2cag 0.0204324)
  (COD1_COD2_COD3_aac2cat 0.00492043)
  (COD1_COD2_COD3_aac2cca 0.000828047)
  (COD1_COD2_COD3_aac2ccc 0.00553961)
  (COD1_COD2_COD3_aac2ccg 0.00116456)
  (COD1_COD2_COD3_aac2cct 9.61688e-06)
  (COD1_COD2_COD3_aac2cga 7.96481e-05)
  (COD1_COD2_COD3_aac2cgc 0.0126125)
  (COD1_COD2_COD3_aac2cgg 0.000531737)
  (COD1_COD2_COD3_aac2cgt 2.68262e-05)
  (COD1_COD2_COD3_aac2cta 0.000107082)
  (COD1_COD2_COD3_aac2ctc 0.0031575)
  (COD1_COD2_COD3_aac2ctg 0.000995011)
  (COD1_COD2_COD3_aac2ctt 5.08966e-06)
  (COD1_COD2_COD3_aac2gaa 0.00668434)
  (COD1_COD2_COD3_aac2gac 0.0690603)
  (COD1_COD2_COD3_aac2gag 0.0114144)
  (COD1_COD2_COD3_aac2gat 0.0106279)
  (COD1_COD2_COD3_aac2gca 0.000684814)
  (COD1_COD2_COD3_aac2gcc 0.0151872)
  (COD1_COD2_COD3_aac2gcg 0.00125335)
  (COD1_COD2_COD3_aac2gct 2.48052e-05)
  (COD1_COD2_COD3_aac2gga 0.00459729)
  (COD1_COD2_COD3_aac2ggc 0.0262962)
  (COD1_COD2_COD3_aac2ggg 0.00300587)
  (COD1_COD2_COD3_aac2ggt 9.95651e-05)
  (COD1_COD2_COD3_aac2gta 2.58406e-05)
  (COD1_COD2_COD3_aac2gtc 0.00396902)
  (COD1_COD2_COD3_aac2gtg 0.000307101)
  (COD1_COD2_COD3_aac2gtt 5.7024e-06)
  (COD1_COD2_COD3_aac2taa 0)
  (COD1_COD2_COD3_aac2tac 0.00814991)
  (COD1_COD2_COD3_aac2tag 0)
  (COD1_COD2_COD3_aac2tat 0.000803731)
  (COD1_COD2_COD3_aac2tca 0.00194772)
  (COD1_COD2_COD3_aac2tcc 0.0120402)
  (COD1_COD2_COD3_aac2tcg 0.00633254)
  (COD1_COD2_COD3_aac2tct 3.98793e-05)
  (COD1_COD2_COD3_aac2tga 0)
  (COD1_COD2_COD3_aac2tgc 0.0024146)
  (COD1_COD2_COD3_aac2tgg 0.000686715)
  (COD1_COD2_COD3_aac2tgt 5.6161e-06)
  (COD1_COD2_COD3_aac2tta 4.56825e-05)
  (COD1_COD2_COD3_aac2ttc 0.00306691)
  (COD1_COD2_COD3_aac2ttg 0.000954478)
  (COD1_COD2_COD3_aac2ttt 1.69064e-06)
  (COD1_COD2_COD3_aag2aaa 0.402204)
  (COD1_COD2_COD3_aag2aac 0.021109)
  (COD1_COD2_COD3_aag2aat 0.0116947)
  (COD1_COD2_COD3_aag2aca 0.00336308)
  (COD1_COD2_COD3_aag2acc 0.0111486)
  (COD1_COD2_COD3_aag2acg 0.0122337)
  (COD1_COD2_COD3_aag2act 0.00368531)
  (COD1_COD2_COD3_aag2aga 0.0116785)
  (COD1_COD2_COD3_aag2agc 0.0127267)
  (COD1_COD2_COD3_aag2agg 0.0496866)
  (COD1_COD2_COD3_aag2agt 0.00401329)
  (COD1_COD2_COD3_aag2ata 0.000721857)
  (COD1_COD2_COD3_aag2atc 0.00276789)
  (COD1_COD2_COD3_aag2atg 0.0105399)
  (COD1_COD2_COD3_aag2att 0.00102654)
  (COD1_COD2_COD3_aag2caa 0.00792156)
  (COD1_COD2_COD3_aag2cac 0.00631063)
  (COD1_COD2_COD3_aag2cag 0.047399)
  (COD1_COD2_COD3_aag2cat 0.00346483)
  (COD1_COD2_COD3_aag2cca 0.000134045)
  (COD1_COD2_COD3_aag2ccc 0.00316188)
  (COD1_COD2_COD3_aag2ccg 0.00685323)
  (COD1_COD2_COD3_aag2cct 0.00230183)
  (COD1_COD2_COD3_aag2cga 0.000165716)
  (COD1_COD2_COD3_aag2cgc 0.0397473)
  (COD1_COD2_COD3_aag2cgg 0.0312717)
  (COD1_COD2_COD3_aag2cgt 0.00476272)
  (COD1_COD2_COD3_aag2cta 5.20021e-06)
  (COD1_COD2_COD3_aag2ctc 0.000376255)
  (COD1_COD2_COD3_aag2ctg 0.00379154)
  (COD1_COD2_COD3_aag2ctt 0.000597406)
  (COD1_COD2_COD3_aag2gaa 0.0106648)
  (COD1_COD2_COD3_aag2gac 0.0107912)
  (COD1_COD2_COD3_aag2gag 0.0478944)
  (COD1_COD2_COD3_aag2gat 0.00538066)
  (COD1_COD2_COD3_aag2gca 7.21821e-05)
  (COD1_COD2_COD3_aag2gcc 0.0100337)
  (COD1_COD2_COD3_aag2gcg 0.0138882)
  (COD1_COD2_COD3_aag2gct 0.00508337)
  (COD1_COD2_COD3_aag2gga 2.91529e-05)
  (COD1_COD2_COD3_aag2ggc 0.00417213)
  (COD1_COD2_COD3_aag2ggg 0.00760941)
  (COD1_COD2_COD3_aag2ggt 0.000365713)
  (COD1_COD2_COD3_aag2gta 5.86678e-06)
  (COD1_COD2_COD3_aag2gtc 0.00103439)
  (COD1_COD2_COD3_aag2gtg 0.00706057)
  (COD1_COD2_COD3_aag2gtt 0.00137248)
  (COD1_COD2_COD3_aag2taa 0)
  (COD1_COD2_COD3_aag2tac 0.00235346)
  (COD1_COD2_COD3_aag2tag 0)
  (COD1_COD2_COD3_aag2tat 0.00125683)
  (COD1_COD2_COD3_aag2tca 4.61279e-05)
  (COD1_COD2_COD3_aag2tcc 0.00270904)
  (COD1_COD2_COD3_aag2tcg 0.00670754)
  (COD1_COD2_COD3_aag2tct 0.00184649)
  (COD1_COD2_COD3_aag2tga 0)
  (COD1_COD2_COD3_aag2tgc 0.000258419)
  (COD1_COD2_COD3_aag2tgg 0.00134144)
  (COD1_COD2_COD3_aag2tgt 7.416e-05)
  (COD1_COD2_COD3_aag2tta 7.06406e-07)
  (COD1_COD2_COD3_aag2ttc 0.000931601)
  (COD1_COD2_COD3_aag2ttg 0.00524806)
  (COD1_COD2_COD3_aag2ttt 0.000280638)
  (COD1_COD2_COD3_aat2aaa 0.0645249)
  (COD1_COD2_COD3_aat2aac 0.358489)
  (COD1_COD2_COD3_aat2aag 0.0140139)
  (COD1_COD2_COD3_aat2aca 0.0163224)
  (COD1_COD2_COD3_aat2acc 0.00120814)
  (COD1_COD2_COD3_aat2acg 0.00239792)
  (COD1_COD2_COD3_aat2act 0.0260423)
  (COD1_COD2_COD3_aat2aga 0.00913107)
  (COD1_COD2_COD3_aat2agc 0.00882615)
  (COD1_COD2_COD3_aat2agg 0.0030861)
  (COD1_COD2_COD3_aat2agt 0.0512608)
  (COD1_COD2_COD3_aat2ata 0.00328475)
  (COD1_COD2_COD3_aat2atc 0.000341459)
  (COD1_COD2_COD3_aat2atg 0.00442533)
  (COD1_COD2_COD3_aat2att 0.00592898)
  (COD1_COD2_COD3_aat2caa 0.0302274)
  (COD1_COD2_COD3_aat2cac 0.00208318)
  (COD1_COD2_COD3_aat2cag 0.00428943)
  (COD1_COD2_COD3_aat2cat 0.0287359)
  (COD1_COD2_COD3_aat2cca 0.0010731)
  (COD1_COD2_COD3_aat2ccc 1.59501e-06)
  (COD1_COD2_COD3_aat2ccg 3.30269e-05)
  (COD1_COD2_COD3_aat2cct 0.00821229)
  (COD1_COD2_COD3_aat2cga 4.80868e-05)
  (COD1_COD2_COD3_aat2cgc 8.51789e-07)
  (COD1_COD2_COD3_aat2cgg 7.86566e-06)
  (COD1_COD2_COD3_aat2cgt 0.0057949)
  (COD1_COD2_COD3_aat2cta 0.000144464)
  (COD1_COD2_COD3_aat2ctc 2.03284e-07)
  (COD1_COD2_COD3_aat2ctg 4.01825e-06)
  (COD1_COD2_COD3_aat2ctt 0.00393615)
  (COD1_COD2_COD3_aat2gaa 0.0337025)
  (COD1_COD2_COD3_aat2gac 0.00422451)
  (COD1_COD2_COD3_aat2gag 0.00457821)
  (COD1_COD2_COD3_aat2gat 0.10097)
  (COD1_COD2_COD3_aat2gca 0.00139841)
  (COD1_COD2_COD3_aat2gcc 1.4256e-06)
  (COD1_COD2_COD3_aat2gcg 6.88111e-05)
  (COD1_COD2_COD3_aat2gct 0.012166)
  (COD1_COD2_COD3_aat2gga 0.014916)
  (COD1_COD2_COD3_aat2ggc 2.64019e-05)
  (COD1_COD2_COD3_aat2ggg 0.00199416)
  (COD1_COD2_COD3_aat2ggt 0.022328)
  (COD1_COD2_COD3_aat2gta 9.65075e-05)
  (COD1_COD2_COD3_aat2gtc 3.97925e-07)
  (COD1_COD2_COD3_aat2gtg 1.46474e-05)
  (COD1_COD2_COD3_aat2gtt 0.00606196)
  (COD1_COD2_COD3_aat2taa 0)
  (COD1_COD2_COD3_aat2tac 0.000713895)
  (COD1_COD2_COD3_aat2tag 0)
  (COD1_COD2_COD3_aat2tat 0.0128954)
  (COD1_COD2_COD3_aat2tca 0.00904289)
  (COD1_COD2_COD3_aat2tcc 9.85304e-06)
  (COD1_COD2_COD3_aat2tcg 0.00035522)
  (COD1_COD2_COD3_aat2tct 0.0209196)
  (COD1_COD2_COD3_aat2tga 0)
  (COD1_COD2_COD3_aat2tgc 1.33475e-06)
  (COD1_COD2_COD3_aat2tgg 0.000953817)
  (COD1_COD2_COD3_aat2tgt 0.00231945)
  (COD1_COD2_COD3_aat2tta 0.00440086)
  (COD1_COD2_COD3_aat2ttc 2.43272e-06)
  (COD1_COD2_COD3_aat2ttg 0.00122415)
  (COD1_COD2_COD3_aat2ttt 0.00564426)
  (COD1_COD2_COD3_aca2aaa 0.0473661)
  (COD1_COD2_COD3_aca2aac 0.00528456)
  (COD1_COD2_COD3_aca2aag 0.00684374)
  (COD1_COD2_COD3_aca2aat 0.0277186)
  (COD1_COD2_COD3_aca2acc 0.149789)
  (COD1_COD2_COD3_aca2acg 0.207906)
  (COD1_COD2_COD3_aca2act 0.40416)
  (COD1_COD2_COD3_aca2aga 0.012833)
  (COD1_COD2_COD3_aca2agc 0.00604518)
  (COD1_COD2_COD3_aca2agg 0.00347686)
  (COD1_COD2_COD3_aca2agt 0.024723)
  (COD1_COD2_COD3_aca2ata 0.028976)
  (COD1_COD2_COD3_aca2atc 0.00116214)
  (COD1_COD2_COD3_aca2atg 0.0180858)
  (COD1_COD2_COD3_aca2att 0.00732985)
  (COD1_COD2_COD3_aca2caa 0.0285177)
  (COD1_COD2_COD3_aca2cac 0.000241615)
  (COD1_COD2_COD3_aca2cag 2.65303e-05)
  (COD1_COD2_COD3_aca2cat 0.000879518)
  (COD1_COD2_COD3_aca2cca 0.011367)
  (COD1_COD2_COD3_aca2ccc 0.000272964)
  (COD1_COD2_COD3_aca2ccg 0.00018311)
  (COD1_COD2_COD3_aca2cct 0.0038418)
  (COD1_COD2_COD3_aca2cga 0.00173482)
  (COD1_COD2_COD3_aca2cgc 6.11107e-08)
  (COD1_COD2_COD3_aca2cgg 2.19086e-07)
  (COD1_COD2_COD3_aca2cgt 5.00222e-06)
  (COD1_COD2_COD3_aca2cta 0.00695372)
  (COD1_COD2_COD3_aca2ctc 2.01191e-06)
  (COD1_COD2_COD3_aca2ctg 1.82459e-06)
  (COD1_COD2_COD3_aca2ctt 1.30825e-05)
  (COD1_COD2_COD3_aca2gaa 0.0402463)
  (COD1_COD2_COD3_aca2gac 8.48927e-05)
  (COD1_COD2_COD3_aca2gag 2.57512e-05)
  (COD1_COD2_COD3_aca2gat 0.00265882)
  (COD1_COD2_COD3_aca2gca 0.0452167)
  (COD1_COD2_COD3_aca2gcc 0.000526967)
  (COD1_COD2_COD3_aca2gcg 0.000911165)
  (COD1_COD2_COD3_aca2gct 0.00612956)
  (COD1_COD2_COD3_aca2gga 0.00765172)
  (COD1_COD2_COD3_aca2ggc 5.85024e-07)
  (COD1_COD2_COD3_aca2ggg 6.20559e-06)
  (COD1_COD2_COD3_aca2ggt 9.19958e-06)
  (COD1_COD2_COD3_aca2gta 0.0423939)
  (COD1_COD2_COD3_aca2gtc 4.65132e-05)
  (COD1_COD2_COD3_aca2gtg 0.000258957)
  (COD1_COD2_COD3_aca2gtt 0.00107087)
  (COD1_COD2_COD3_aca2taa 0)
  (COD1_COD2_COD3_aca2tac 0.000891795)
  (COD1_COD2_COD3_aca2tag 0)
  (COD1_COD2_COD3_aca2tat 0.00358702)
  (COD1_COD2_COD3_aca2tca 0.0592106)
  (COD1_COD2_COD3_aca2tcc 0.00441002)
  (COD1_COD2_COD3_aca2tcg 0.00305206)
  (COD1_COD2_COD3_aca2tct 0.0238148)
  (COD1_COD2_COD3_aca2tga 0)
  (COD1_COD2_COD3_aca2tgc 0.000454321)
  (COD1_COD2_COD3_aca2tgg 0.000770456)
  (COD1_COD2_COD3_aca2tgt 0.00267917)
  (COD1_COD2_COD3_aca2tta 0.0141211)
  (COD1_COD2_COD3_aca2ttc 0.000224811)
  (COD1_COD2_COD3_aca2ttg 0.000174922)
  (COD1_COD2_COD3_aca2ttt 0.00171331)
  (COD1_COD2_COD3_acc2aaa 0.00278135)
  (COD1_COD2_COD3_acc2aac 0.0273749)
  (COD1_COD2_COD3_acc2aag 0.0166539)
  (COD1_COD2_COD3_acc2aat 0.00150502)
  (COD1_COD2_COD3_acc2aca 0.10988)
  (COD1_COD2_COD3_acc2acg 0.304181)
  (COD1_COD2_COD3_acc2act 0.177145)
  (COD1_COD2_COD3_acc2aga 0.000211655)
  (COD1_COD2_COD3_acc2agc 0.0643909)
  (COD1_COD2_COD3_acc2agg 0.00095514)
  (COD1_COD2_COD3_acc2agt 0.0028457)
  (COD1_COD2_COD3_acc2ata 0.000213867)
  (COD1_COD2_COD3_acc2atc 0.0253477)
  (COD1_COD2_COD3_acc2atg 0.00918821)
  (COD1_COD2_COD3_acc2att 0.00085606)
  (COD1_COD2_COD3_acc2caa 0.00114218)
  (COD1_COD2_COD3_acc2cac 0.0107018)
  (COD1_COD2_COD3_acc2cag 0.0128063)
  (COD1_COD2_COD3_acc2cat 1.24975e-05)
  (COD1_COD2_COD3_acc2cca 0.000303293)
  (COD1_COD2_COD3_acc2ccc 0.009068)
  (COD1_COD2_COD3_acc2ccg 0.0064343)
  (COD1_COD2_COD3_acc2cct 0.000460025)
  (COD1_COD2_COD3_acc2cga 9.65696e-06)
  (COD1_COD2_COD3_acc2cgc 0.0216423)
  (COD1_COD2_COD3_acc2cgg 0.000144665)
  (COD1_COD2_COD3_acc2cgt 9.00402e-06)
  (COD1_COD2_COD3_acc2cta 4.80178e-06)
  (COD1_COD2_COD3_acc2ctc 0.0150823)
  (COD1_COD2_COD3_acc2ctg 0.00209912)
  (COD1_COD2_COD3_acc2ctt 6.44673e-06)
  (COD1_COD2_COD3_acc2gaa 0.00378891)
  (COD1_COD2_COD3_acc2gac 0.0256304)
  (COD1_COD2_COD3_acc2gag 0.0110256)
  (COD1_COD2_COD3_acc2gat 1.65708e-05)
  (COD1_COD2_COD3_acc2gca 0.00127702)
  (COD1_COD2_COD3_acc2gcc 0.0579118)
  (COD1_COD2_COD3_acc2gcg 0.0133317)
  (COD1_COD2_COD3_acc2gct 0.00227172)
  (COD1_COD2_COD3_acc2gga 1.4089e-06)
  (COD1_COD2_COD3_acc2ggc 0.0122846)
  (COD1_COD2_COD3_acc2ggg 4.53332e-06)
  (COD1_COD2_COD3_acc2ggt 2.20315e-06)
  (COD1_COD2_COD3_acc2gta 2.15493e-05)
  (COD1_COD2_COD3_acc2gtc 0.0527385)
  (COD1_COD2_COD3_acc2gtg 0.00202902)
  (COD1_COD2_COD3_acc2gtt 5.75031e-05)
  (COD1_COD2_COD3_acc2taa 0)
  (COD1_COD2_COD3_acc2tac 0.00657406)
  (COD1_COD2_COD3_acc2tag 0)
  (COD1_COD2_COD3_acc2tat 1.5928e-06)
  (COD1_COD2_COD3_acc2tca 0.00155702)
  (COD1_COD2_COD3_acc2tcc 0.0412346)
  (COD1_COD2_COD3_acc2tcg 0.0209051)
  (COD1_COD2_COD3_acc2tct 0.00232428)
  (COD1_COD2_COD3_acc2tga 0)
  (COD1_COD2_COD3_acc2tgc 0.0059785)
  (COD1_COD2_COD3_acc2tgg 0.000481893)
  (COD1_COD2_COD3_acc2tgt 4.72911e-06)
  (COD1_COD2_COD3_acc2tta 7.32331e-07)
  (COD1_COD2_COD3_acc2ttc 0.00793255)
  (COD1_COD2_COD3_acc2ttg 4.65455e-05)
  (COD1_COD2_COD3_acc2ttt 1.66044e-06)
  (COD1_COD2_COD3_acg2aaa 0.00620652)
  (COD1_COD2_COD3_acg2aac 0.0167117)
  (COD1_COD2_COD3_acg2aag 0.0305645)
  (COD1_COD2_COD3_acg2aat 0.0049995)
  (COD1_COD2_COD3_acg2aca 0.25543)
  (COD1_COD2_COD3_acg2acc 0.509446)
  (COD1_COD2_COD3_acg2act 0.153361)
  (COD1_COD2_COD3_acg2aga 0.000857248)
  (COD1_COD2_COD3_acg2agc 0.0393634)
  (COD1_COD2_COD3_acg2agg 0.00519738)
  (COD1_COD2_COD3_acg2agt 0.00581099)
  (COD1_COD2_COD3_acg2ata 0.00197902)
  (COD1_COD2_COD3_acg2atc 0.0074116)
  (COD1_COD2_COD3_acg2atg 0.0291775)
  (COD1_COD2_COD3_acg2att 0.00224666)
  (COD1_COD2_COD3_acg2caa 4.88934e-05)
  (COD1_COD2_COD3_acg2cac 0.00113901)
  (COD1_COD2_COD3_acg2cag 0.0366511)
  (COD1_COD2_COD3_acg2cat 0.000821757)
  (COD1_COD2_COD3_acg2cca 0.000932893)
  (COD1_COD2_COD3_acg2ccc 0.00453086)
  (COD1_COD2_COD3_acg2ccg 0.0117434)
  (COD1_COD2_COD3_acg2cct 0.000831498)
  (COD1_COD2_COD3_acg2cga 1.35721e-05)
  (COD1_COD2_COD3_acg2cgc 0.000208452)
  (COD1_COD2_COD3_acg2cgg 0.0243658)
  (COD1_COD2_COD3_acg2cgt 3.88426e-05)
  (COD1_COD2_COD3_acg2cta 2.879e-06)
  (COD1_COD2_COD3_acg2ctc 9.55707e-06)
  (COD1_COD2_COD3_acg2ctg 0.0209051)
  (COD1_COD2_COD3_acg2ctt 4.44003e-06)
  (COD1_COD2_COD3_acg2gaa 0.000119724)
  (COD1_COD2_COD3_acg2gac 0.00240791)
  (COD1_COD2_COD3_acg2gag 0.0443169)
  (COD1_COD2_COD3_acg2gat 0.00121823)
  (COD1_COD2_COD3_acg2gca 0.00526263)
  (COD1_COD2_COD3_acg2gcc 0.0133779)
  (COD1_COD2_COD3_acg2gcg 0.0524469)
  (COD1_COD2_COD3_acg2gct 0.00205307)
  (COD1_COD2_COD3_acg2gga 2.9888e-06)
  (COD1_COD2_COD3_acg2ggc 0.000151962)
  (COD1_COD2_COD3_acg2ggg 0.00732477)
  (COD1_COD2_COD3_acg2ggt 6.70189e-06)
  (COD1_COD2_COD3_acg2gta 2.7504e-05)
  (COD1_COD2_COD3_acg2gtc 0.000189174)
  (COD1_COD2_COD3_acg2gtg 0.0699758)
  (COD1_COD2_COD3_acg2gtt 9.19367e-05)
  (COD1_COD2_COD3_acg2taa 0)
  (COD1_COD2_COD3_acg2tac 0.0029686)
  (COD1_COD2_COD3_acg2tag 0)
  (COD1_COD2_COD3_acg2tat 0.000726374)
  (COD1_COD2_COD3_acg2tca 0.00535833)
  (COD1_COD2_COD3_acg2tcc 0.0168163)
  (COD1_COD2_COD3_acg2tcg 0.0429261)
  (COD1_COD2_COD3_acg2tct 0.00426275)
  (COD1_COD2_COD3_acg2tga 0)
  (COD1_COD2_COD3_acg2tgc 0.00128501)
  (COD1_COD2_COD3_acg2tgg 0.00462286)
  (COD1_COD2_COD3_acg2tgt 0.000347428)
  (COD1_COD2_COD3_acg2tta 9.1166e-07)
  (COD1_COD2_COD3_acg2ttc 0.000779048)
  (COD1_COD2_COD3_acg2ttg 0.00922651)
  (COD1_COD2_COD3_acg2ttt 0.000172514)
  (COD1_COD2_COD3_act2aaa 0.0273181)
  (COD1_COD2_COD3_act2aac 0.00602009)
  (COD1_COD2_COD3_act2aag 0.00777469)
  (COD1_COD2_COD3_act2aat 0.0458479)
  (COD1_COD2_COD3_act2aca 0.418994)
  (COD1_COD2_COD3_act2acc 0.250347)
  (COD1_COD2_COD3_act2acg 0.129498)
  (COD1_COD2_COD3_act2aga 0.00422158)
  (COD1_COD2_COD3_act2agc 0.00853733)
  (COD1_COD2_COD3_act2agg 0.00210203)
  (COD1_COD2_COD3_act2agt 0.0509773)
  (COD1_COD2_COD3_act2ata 0.00451205)
  (COD1_COD2_COD3_act2atc 0.00180348)
  (COD1_COD2_COD3_act2atg 0.0088937)
  (COD1_COD2_COD3_act2att 0.0313802)
  (COD1_COD2_COD3_act2caa 0.00642985)
  (COD1_COD2_COD3_act2cac 3.35792e-06)
  (COD1_COD2_COD3_act2cag 0.00060365)
  (COD1_COD2_COD3_act2cat 0.00959169)
  (COD1_COD2_COD3_act2cca 0.00385782)
  (COD1_COD2_COD3_act2ccc 0.000560499)
  (COD1_COD2_COD3_act2ccg 0.000140125)
  (COD1_COD2_COD3_act2cct 0.010957)
  (COD1_COD2_COD3_act2cga 4.71602e-06)
  (COD1_COD2_COD3_act2cgc 6.90396e-08)
  (COD1_COD2_COD3_act2cgg 2.02581e-07)
  (COD1_COD2_COD3_act2cgt 0.00292773)
  (COD1_COD2_COD3_act2cta 3.23472e-05)
  (COD1_COD2_COD3_act2ctc 1.41575e-06)
  (COD1_COD2_COD3_act2ctg 2.1608e-06)
  (COD1_COD2_COD3_act2ctt 0.0126125)
  (COD1_COD2_COD3_act2gaa 0.00868656)
  (COD1_COD2_COD3_act2gac 4.77192e-06)
  (COD1_COD2_COD3_act2gag 0.00102867)
  (COD1_COD2_COD3_act2gat 0.0265711)
  (COD1_COD2_COD3_act2gca 0.00630626)
  (COD1_COD2_COD3_act2gcc 0.00103439)
  (COD1_COD2_COD3_act2gcg 0.000436721)
  (COD1_COD2_COD3_act2gct 0.0446251)
  (COD1_COD2_COD3_act2gga 4.06045e-05)
  (COD1_COD2_COD3_act2ggc 6.71086e-07)
  (COD1_COD2_COD3_act2ggg 5.762e-06)
  (COD1_COD2_COD3_act2ggt 0.00642539)
  (COD1_COD2_COD3_act2gta 9.83988e-05)
  (COD1_COD2_COD3_act2gtc 1.95158e-05)
  (COD1_COD2_COD3_act2gtg 8.24569e-05)
  (COD1_COD2_COD3_act2gtt 0.0551689)
  (COD1_COD2_COD3_act2taa 0)
  (COD1_COD2_COD3_act2tac 1.00253e-05)
  (COD1_COD2_COD3_act2tag 0)
  (COD1_COD2_COD3_act2tat 0.0103444)
  (COD1_COD2_COD3_act2tca 0.0238313)
  (COD1_COD2_COD3_act2tcc 0.00524442)
  (COD1_COD2_COD3_act2tcg 0.00174326)
  (COD1_COD2_COD3_act2tct 0.0553989)
  (COD1_COD2_COD3_act2tga 0)
  (COD1_COD2_COD3_act2tgc 1.07523e-05)
  (COD1_COD2_COD3_act2tgg 0.000458433)
  (COD1_COD2_COD3_act2tgt 0.00702152)
  (COD1_COD2_COD3_act2tta 0.000202471)
  (COD1_COD2_COD3_act2ttc 2.90305e-06)
  (COD1_COD2_COD3_act2ttg 0.000245839)
  (COD1_COD2_COD3_act2ttt 0.01065)
  (COD1_COD2_COD3_aga2aaa 0.180867)
  (COD1_COD2_COD3_aga2aac 0.00556655)
  (COD1_COD2_COD3_aga2aag 0.0319952)
  (COD1_COD2_COD3_aga2aat 0.0208762)
  (COD1_COD2_COD3_aga2aca 0.017277)
  (COD1_COD2_COD3_aga2acc 0.000388446)
  (COD1_COD2_COD3_aga2acg 0.000939382)
  (COD1_COD2_COD3_aga2act 0.00548231)
  (COD1_COD2_COD3_aga2agc 0.00553577)
  (COD1_COD2_COD3_aga2agg 0.298127)
  (COD1_COD2_COD3_aga2agt 0.0169567)
  (COD1_COD2_COD3_aga2ata 0.009999)
  (COD1_COD2_COD3_aga2atc 0.000293165)
  (COD1_COD2_COD3_aga2atg 0.00647906)
  (COD1_COD2_COD3_aga2att 0.00287345)
  (COD1_COD2_COD3_aga2caa 0.0213296)
  (COD1_COD2_COD3_aga2cac 0.000281808)
  (COD1_COD2_COD3_aga2cag 6.33159e-06)
  (COD1_COD2_COD3_aga2cat 0.00438867)
  (COD1_COD2_COD3_aga2cca 0.0055128)
  (COD1_COD2_COD3_aga2ccc 4.13982e-06)
  (COD1_COD2_COD3_aga2ccg 7.8431e-08)
  (COD1_COD2_COD3_aga2cct 0.000119558)
  (COD1_COD2_COD3_aga2cga 0.15368)
  (COD1_COD2_COD3_aga2cgc 0.016266)
  (COD1_COD2_COD3_aga2cgg 0.0197364)
  (COD1_COD2_COD3_aga2cgt 0.148239)
  (COD1_COD2_COD3_aga2cta 0.00301422)
  (COD1_COD2_COD3_aga2ctc 1.19546e-06)
  (COD1_COD2_COD3_aga2ctg 5.97287e-08)
  (COD1_COD2_COD3_aga2ctt 7.18327e-05)
  (COD1_COD2_COD3_aga2gaa 0.0230834)
  (COD1_COD2_COD3_aga2gac 1.81962e-05)
  (COD1_COD2_COD3_aga2gag 1.38285e-05)
  (COD1_COD2_COD3_aga2gat 0.00087042)
  (COD1_COD2_COD3_aga2gca 0.00763054)
  (COD1_COD2_COD3_aga2gcc 9.54336e-07)
  (COD1_COD2_COD3_aga2gcg 1.63297e-07)
  (COD1_COD2_COD3_aga2gct 0.000261301)
  (COD1_COD2_COD3_aga2gga 0.0153885)
  (COD1_COD2_COD3_aga2ggc 1.89427e-05)
  (COD1_COD2_COD3_aga2ggg 0.000710932)
  (COD1_COD2_COD3_aga2ggt 0.000636744)
  (COD1_COD2_COD3_aga2gta 0.00612532)
  (COD1_COD2_COD3_aga2gtc 4.0322e-06)
  (COD1_COD2_COD3_aga2gtg 1.3562e-06)
  (COD1_COD2_COD3_aga2gtt 0.000611229)
  (COD1_COD2_COD3_aga2taa 0)
  (COD1_COD2_COD3_aga2tac 0.00101662)
  (COD1_COD2_COD3_aga2tag 0)
  (COD1_COD2_COD3_aga2tat 0.00731463)
  (COD1_COD2_COD3_aga2tca 0.0113513)
  (COD1_COD2_COD3_aga2tcc 2.16992e-05)
  (COD1_COD2_COD3_aga2tcg 7.84893e-07)
  (COD1_COD2_COD3_aga2tct 0.000564789)
  (COD1_COD2_COD3_aga2tga 0)
  (COD1_COD2_COD3_aga2tgc 0.000228899)
  (COD1_COD2_COD3_aga2tgg 0.00185162)
  (COD1_COD2_COD3_aga2tgt 0.000930311)
  (COD1_COD2_COD3_aga2tta 0.00686274)
  (COD1_COD2_COD3_aga2ttc 0.000268647)
  (COD1_COD2_COD3_aga2ttg 2.66963e-05)
  (COD1_COD2_COD3_aga2ttt 0.00301004)
  (COD1_COD2_COD3_agc2aaa 0.0103948)
  (COD1_COD2_COD3_agc2aac 0.0839108)
  (COD1_COD2_COD3_agc2aag 0.0248089)
  (COD1_COD2_COD3_agc2aat 0.0143481)
  (COD1_COD2_COD3_agc2aca 0.00578687)
  (COD1_COD2_COD3_agc2acc 0.0840855)
  (COD1_COD2_COD3_agc2acg 0.0306919)
  (COD1_COD2_COD3_agc2act 0.00788322)
  (COD1_COD2_COD3_agc2aga 0.00393888)
  (COD1_COD2_COD3_agc2agg 0.00874091)
  (COD1_COD2_COD3_agc2agt 0.294839)
  (COD1_COD2_COD3_agc2ata 0.0002762)
  (COD1_COD2_COD3_agc2atc 0.00567171)
  (COD1_COD2_COD3_agc2atg 0.00645217)
  (COD1_COD2_COD3_agc2att 0.00077099)
  (COD1_COD2_COD3_agc2caa 0.00399663)
  (COD1_COD2_COD3_agc2cac 0.0141603)
  (COD1_COD2_COD3_agc2cag 0.0294212)
  (COD1_COD2_COD3_agc2cat 5.4439e-05)
  (COD1_COD2_COD3_agc2cca 5.82251e-05)
  (COD1_COD2_COD3_agc2ccc 0.0139267)
  (COD1_COD2_COD3_agc2ccg 0.00127702)
  (COD1_COD2_COD3_agc2cct 1.33112e-05)
  (COD1_COD2_COD3_agc2cga 0.00170501)
  (COD1_COD2_COD3_agc2cgc 0.0193034)
  (COD1_COD2_COD3_agc2cgg 0.0111023)
  (COD1_COD2_COD3_agc2cgt 0.00329387)
  (COD1_COD2_COD3_agc2cta 2.03445e-05)
  (COD1_COD2_COD3_agc2ctc 0.00504825)
  (COD1_COD2_COD3_agc2ctg 0.000493384)
  (COD1_COD2_COD3_agc2ctt 5.35753e-06)
  (COD1_COD2_COD3_agc2gaa 0.01065)
  (COD1_COD2_COD3_agc2gac 0.0507306)
  (COD1_COD2_COD3_agc2gag 0.0251901)
  (COD1_COD2_COD3_agc2gat 0.000357691)
  (COD1_COD2_COD3_agc2gca 0.00104665)
  (COD1_COD2_COD3_agc2gcc 0.0594162)
  (COD1_COD2_COD3_agc2gcg 0.00366239)
  (COD1_COD2_COD3_agc2gct 0.000100675)
  (COD1_COD2_COD3_agc2gga 0.00489661)
  (COD1_COD2_COD3_agc2ggc 0.0566414)
  (COD1_COD2_COD3_agc2ggg 0.0117597)
  (COD1_COD2_COD3_agc2ggt 0.00550516)
  (COD1_COD2_COD3_agc2gta 6.84746e-06)
  (COD1_COD2_COD3_agc2gtc 0.00702639)
  (COD1_COD2_COD3_agc2gtg 9.42597e-05)
  (COD1_COD2_COD3_agc2gtt 4.6997e-06)
  (COD1_COD2_COD3_agc2taa 0)
  (COD1_COD2_COD3_agc2tac 0.0057949)
  (COD1_COD2_COD3_agc2tag 0)
  (COD1_COD2_COD3_agc2tat 2.36619e-06)
  (COD1_COD2_COD3_agc2tca 0.0108889)
  (COD1_COD2_COD3_agc2tcc 0.117278)
  (COD1_COD2_COD3_agc2tcg 0.0856739)
  (COD1_COD2_COD3_agc2tct 0.00111789)
  (COD1_COD2_COD3_agc2tga 0)
  (COD1_COD2_COD3_agc2tgc 0.0101808)
  (COD1_COD2_COD3_agc2tgg 0.00213137)
  (COD1_COD2_COD3_agc2tgt 0.00159857)
  (COD1_COD2_COD3_agc2tta 6.79074e-06)
  (COD1_COD2_COD3_agc2ttc 0.00355977)
  (COD1_COD2_COD3_agc2ttg 9.07967e-05)
  (COD1_COD2_COD3_agc2ttt 2.22463e-06)
  (COD1_COD2_COD3_agg2aaa 0.0581128)
  (COD1_COD2_COD3_agg2aac 0.0122082)
  (COD1_COD2_COD3_agg2aag 0.195467)
  (COD1_COD2_COD3_agg2aat 0.0101245)
  (COD1_COD2_COD3_agg2aca 0.00671685)
  (COD1_COD2_COD3_agg2acc 0.00251714)
  (COD1_COD2_COD3_agg2acg 0.00817821)
  (COD1_COD2_COD3_agg2act 0.00391981)
  (COD1_COD2_COD3_agg2aga 0.428094)
  (COD1_COD2_COD3_agg2agc 0.0176401)
  (COD1_COD2_COD3_agg2agt 0.0152399)
  (COD1_COD2_COD3_agg2ata 0.00406931)
  (COD1_COD2_COD3_agg2atc 0.00148225)
  (COD1_COD2_COD3_agg2atg 0.0191967)
  (COD1_COD2_COD3_agg2att 0.00231624)
  (COD1_COD2_COD3_agg2caa 6.45153e-05)
  (COD1_COD2_COD3_agg2cac 0.00186708)
  (COD1_COD2_COD3_agg2cag 0.0238148)
  (COD1_COD2_COD3_agg2cat 0.00150398)
  (COD1_COD2_COD3_agg2cca 0.000103291)
  (COD1_COD2_COD3_agg2ccc 0.000346466)
  (COD1_COD2_COD3_agg2ccg 0.00334217)
  (COD1_COD2_COD3_agg2cct 0.000630594)
  (COD1_COD2_COD3_agg2cga 0.0735567)
  (COD1_COD2_COD3_agg2cgc 0.0597052)
  (COD1_COD2_COD3_agg2cgg 0.151249)
  (COD1_COD2_COD3_agg2cgt 0.0577514)
  (COD1_COD2_COD3_agg2cta 7.38485e-06)
  (COD1_COD2_COD3_agg2ctc 1.6073e-05)
  (COD1_COD2_COD3_agg2ctg 0.00115012)
  (COD1_COD2_COD3_agg2ctt 0.000144966)
  (COD1_COD2_COD3_agg2gaa 0.000152595)
  (COD1_COD2_COD3_agg2gac 0.000227633)
  (COD1_COD2_COD3_agg2gag 0.0366003)
  (COD1_COD2_COD3_agg2gat 0.000450246)
  (COD1_COD2_COD3_agg2gca 7.39034e-05)
  (COD1_COD2_COD3_agg2gcc 4.89952e-05)
  (COD1_COD2_COD3_agg2gcg 0.00343613)
  (COD1_COD2_COD3_agg2gct 0.000859032)
  (COD1_COD2_COD3_agg2gga 0.00271468)
  (COD1_COD2_COD3_agg2ggc 0.000289932)
  (COD1_COD2_COD3_agg2ggg 0.0159423)
  (COD1_COD2_COD3_agg2ggt 0.00080095)
  (COD1_COD2_COD3_agg2gta 2.29523e-05)
  (COD1_COD2_COD3_agg2gtc 3.78591e-05)
  (COD1_COD2_COD3_agg2gtg 0.00711461)
  (COD1_COD2_COD3_agg2gtt 0.000985402)
  (COD1_COD2_COD3_agg2taa 0)
  (COD1_COD2_COD3_agg2tac 0.00450268)
  (COD1_COD2_COD3_agg2tag 0)
  (COD1_COD2_COD3_agg2tat 0.00345763)
  (COD1_COD2_COD3_agg2tca 9.61069e-05)
  (COD1_COD2_COD3_agg2tcc 0.000308595)
  (COD1_COD2_COD3_agg2tcg 0.0044871)
  (COD1_COD2_COD3_agg2tct 0.000755647)
  (COD1_COD2_COD3_agg2tga 0)
  (COD1_COD2_COD3_agg2tgc 0.000522964)
  (COD1_COD2_COD3_agg2tgg 0.00633254)
  (COD1_COD2_COD3_agg2tgt 0.000787736)
  (COD1_COD2_COD3_agg2tta 3.58897e-06)
  (COD1_COD2_COD3_agg2ttc 0.00121233)
  (COD1_COD2_COD3_agg2ttg 0.00797113)
  (COD1_COD2_COD3_agg2ttt 0.00178483)
  (COD1_COD2_COD3_agt2aaa 0.0402463)
  (COD1_COD2_COD3_agt2aac 0.0119239)
  (COD1_COD2_COD3_agt2aag 0.0103876)
  (COD1_COD2_COD3_agt2aat 0.110798)
  (COD1_COD2_COD3_agt2aca 0.0314674)
  (COD1_COD2_COD3_agt2acc 0.00493409)
  (COD1_COD2_COD3_agt2acg 0.00602009)
  (COD1_COD2_COD3_agt2act 0.0625867)
  (COD1_COD2_COD3_agt2aga 0.0160309)
  (COD1_COD2_COD3_agt2agc 0.391749)
  (COD1_COD2_COD3_agt2agg 0.0100337)
  (COD1_COD2_COD3_agt2ata 0.00346003)
  (COD1_COD2_COD3_agt2atc 0.000445589)
  (COD1_COD2_COD3_agt2atg 0.00649255)
  (COD1_COD2_COD3_agt2att 0.0088937)
  (COD1_COD2_COD3_agt2caa 0.020221)
  (COD1_COD2_COD3_agt2cac 5.06502e-06)
  (COD1_COD2_COD3_agt2cag 0.00172642)
  (COD1_COD2_COD3_agt2cat 0.0138018)
  (COD1_COD2_COD3_agt2cca 0.000234683)
  (COD1_COD2_COD3_agt2ccc 3.45711e-06)
  (COD1_COD2_COD3_agt2ccg 1.25583e-05)
  (COD1_COD2_COD3_agt2cct 0.0107912)
  (COD1_COD2_COD3_agt2cga 0.00255405)
  (COD1_COD2_COD3_agt2cgc 0.000264949)
  (COD1_COD2_COD3_agt2cgg 0.000203879)
  (COD1_COD2_COD3_agt2cgt 0.0065604)
  (COD1_COD2_COD3_agt2cta 6.98684e-05)
  (COD1_COD2_COD3_agt2ctc 5.80983e-07)
  (COD1_COD2_COD3_agt2ctg 2.52025e-06)
  (COD1_COD2_COD3_agt2ctt 0.00489661)
  (COD1_COD2_COD3_agt2gaa 0.0287559)
  (COD1_COD2_COD3_agt2gac 1.08873e-05)
  (COD1_COD2_COD3_agt2gag 0.00395255)
  (COD1_COD2_COD3_agt2gat 0.0486978)
  (COD1_COD2_COD3_agt2gca 0.00208751)
  (COD1_COD2_COD3_agt2gcc 1.23682e-05)
  (COD1_COD2_COD3_agt2gcg 0.000116531)
  (COD1_COD2_COD3_agt2gct 0.0439498)
  (COD1_COD2_COD3_agt2gga 0.0193302)
  (COD1_COD2_COD3_agt2ggc 0.00127968)
  (COD1_COD2_COD3_agt2ggg 0.00700694)
  (COD1_COD2_COD3_agt2ggt 0.0402463)
  (COD1_COD2_COD3_agt2gta 6.86205e-05)
  (COD1_COD2_COD3_agt2gtc 1.01718e-06)
  (COD1_COD2_COD3_agt2gtg 3.03909e-05)
  (COD1_COD2_COD3_agt2gtt 0.00953864)
  (COD1_COD2_COD3_agt2taa 0)
  (COD1_COD2_COD3_agt2tac 2.57499e-06)
  (COD1_COD2_COD3_agt2tag 0)
  (COD1_COD2_COD3_agt2tat 0.00873486)
  (COD1_COD2_COD3_agt2tca 0.0618535)
  (COD1_COD2_COD3_agt2tcc 0.000900491)
  (COD1_COD2_COD3_agt2tcg 0.00594544)
  (COD1_COD2_COD3_agt2tct 0.143587)
  (COD1_COD2_COD3_agt2tga 0)
  (COD1_COD2_COD3_agt2tgc 0.00134797)
  (COD1_COD2_COD3_agt2tgg 0.0017725)
  (COD1_COD2_COD3_agt2tgt 0.0129402)
  (COD1_COD2_COD3_agt2tta 0.000177241)
  (COD1_COD2_COD3_agt2ttc 1.01085e-06)
  (COD1_COD2_COD3_agt2ttg 0.000145671)
  (COD1_COD2_COD3_agt2ttt 0.00483255)
  (COD1_COD2_COD3_ata2aaa 0.0207607)
  (COD1_COD2_COD3_ata2aac 0.000423603)
  (COD1_COD2_COD3_ata2aag 0.00150398)
  (COD1_COD2_COD3_ata2aat 0.00571116)
  (COD1_COD2_COD3_ata2aca 0.0296669)
  (COD1_COD2_COD3_ata2acc 0.000298289)
  (COD1_COD2_COD3_ata2acg 0.00165036)
  (COD1_COD2_COD3_ata2act 0.00445611)
  (COD1_COD2_COD3_ata2aga 0.00760414)
  (COD1_COD2_COD3_ata2agc 0.000295204)
  (COD1_COD2_COD3_ata2agg 0.00215514)
  (COD1_COD2_COD3_ata2agt 0.00278328)
  (COD1_COD2_COD3_ata2atc 0.137547)
  (COD1_COD2_COD3_ata2atg 0.0472677)
  (COD1_COD2_COD3_ata2att 0.40669)
  (COD1_COD2_COD3_ata2caa 0.00669825)
  (COD1_COD2_COD3_ata2cac 3.06008e-06)
  (COD1_COD2_COD3_ata2cag 1.15955e-06)
  (COD1_COD2_COD3_ata2cat 0.000427734)
  (COD1_COD2_COD3_ata2cca 0.00536577)
  (COD1_COD2_COD3_ata2ccc 2.64725e-07)
  (COD1_COD2_COD3_ata2ccg 4.58365e-07)
  (COD1_COD2_COD3_ata2cct 2.12379e-05)
  (COD1_COD2_COD3_ata2cga 0.00058633)
  (COD1_COD2_COD3_ata2cgc 3.70726e-09)
  (COD1_COD2_COD3_ata2cgg 1.38648e-08)
  (COD1_COD2_COD3_ata2cgt 3.78291e-07)
  (COD1_COD2_COD3_ata2cta 0.0323969)
  (COD1_COD2_COD3_ata2ctc 0.00228752)
  (COD1_COD2_COD3_ata2ctg 0.00139357)
  (COD1_COD2_COD3_ata2ctt 0.019117)
  (COD1_COD2_COD3_ata2gaa 0.0124821)
  (COD1_COD2_COD3_ata2gac 8.90382e-08)
  (COD1_COD2_COD3_ata2gag 3.37425e-06)
  (COD1_COD2_COD3_ata2gat 3.2776e-05)
  (COD1_COD2_COD3_ata2gca 0.0162547)
  (COD1_COD2_COD3_ata2gcc 3.45934e-07)
  (COD1_COD2_COD3_ata2gcg 1.4385e-06)
  (COD1_COD2_COD3_ata2gct 3.38849e-05)
  (COD1_COD2_COD3_ata2gga 0.00318608)
  (COD1_COD2_COD3_ata2ggc 1.7586e-08)
  (COD1_COD2_COD3_ata2ggg 6.23546e-07)
  (COD1_COD2_COD3_ata2ggt 2.09154e-06)
  (COD1_COD2_COD3_ata2gta 0.123108)
  (COD1_COD2_COD3_ata2gtc 0.00284176)
  (COD1_COD2_COD3_ata2gtg 0.0206173)
  (COD1_COD2_COD3_ata2gtt 0.0490365)
  (COD1_COD2_COD3_ata2taa 0)
  (COD1_COD2_COD3_ata2tac 0.000424779)
  (COD1_COD2_COD3_ata2tag 0)
  (COD1_COD2_COD3_ata2tat 0.00988187)
  (COD1_COD2_COD3_ata2tca 0.00418371)
  (COD1_COD2_COD3_ata2tcc 2.6381e-07)
  (COD1_COD2_COD3_ata2tcg 1.53954e-07)
  (COD1_COD2_COD3_ata2tct 5.45496e-06)
  (COD1_COD2_COD3_ata2tga 0)
  (COD1_COD2_COD3_ata2tgc 8.97953e-05)
  (COD1_COD2_COD3_ata2tgg 0.00216713)
  (COD1_COD2_COD3_ata2tgt 0.000448377)
  (COD1_COD2_COD3_ata2tta 0.0835625)
  (COD1_COD2_COD3_ata2ttc 0.00138779)
  (COD1_COD2_COD3_ata2ttg 0.0167581)
  (COD1_COD2_COD3_ata2ttt 0.0230355)
  (COD1_COD2_COD3_atc2aaa 0.000715877)
  (COD1_COD2_COD3_atc2aac 0.00356718)
  (COD1_COD2_COD3_atc2aag 0.00310972)
  (COD1_COD2_COD3_atc2aat 0.000320142)
  (COD1_COD2_COD3_atc2aca 0.000641617)
  (COD1_COD2_COD3_atc2acc 0.0190773)
  (COD1_COD2_COD3_atc2acg 0.00333061)
  (COD1_COD2_COD3_atc2act 0.000960451)
  (COD1_COD2_COD3_atc2aga 0.000120223)
  (COD1_COD2_COD3_atc2agc 0.00326885)
  (COD1_COD2_COD3_atc2agg 0.000423603)
  (COD1_COD2_COD3_atc2agt 0.000193283)
  (COD1_COD2_COD3_atc2ata 0.074171)
  (COD1_COD2_COD3_atc2atg 0.0302693)
  (COD1_COD2_COD3_atc2att 0.337042)
  (COD1_COD2_COD3_atc2caa 0.000181593)
  (COD1_COD2_COD3_atc2cac 0.00141302)
  (COD1_COD2_COD3_atc2cag 0.00219586)
  (COD1_COD2_COD3_atc2cat 4.98146e-06)
  (COD1_COD2_COD3_atc2cca 5.06151e-06)
  (COD1_COD2_COD3_atc2ccc 0.00281432)
  (COD1_COD2_COD3_atc2ccg 0.000265316)
  (COD1_COD2_COD3_atc2cct 1.33938e-06)
  (COD1_COD2_COD3_atc2cga 5.73411e-06)
  (COD1_COD2_COD3_atc2cgc 0.00348893)
  (COD1_COD2_COD3_atc2cgg 2.52388e-05)
  (COD1_COD2_COD3_atc2cgt 8.42395e-07)
  (COD1_COD2_COD3_atc2cta 0.00149463)
  (COD1_COD2_COD3_atc2ctc 0.0632847)
  (COD1_COD2_COD3_atc2ctg 0.0704137)
  (COD1_COD2_COD3_atc2ctt 0.00592077)
  (COD1_COD2_COD3_atc2gaa 0.000316173)
  (COD1_COD2_COD3_atc2gac 0.000821188)
  (COD1_COD2_COD3_atc2gag 0.000979274)
  (COD1_COD2_COD3_atc2gat 1.22567e-06)
  (COD1_COD2_COD3_atc2gca 9.78498e-06)
  (COD1_COD2_COD3_atc2gcc 0.0143879)
  (COD1_COD2_COD3_atc2gcg 8.44818e-05)
  (COD1_COD2_COD3_atc2gct 4.93335e-06)
  (COD1_COD2_COD3_atc2gga 1.06405e-06)
  (COD1_COD2_COD3_atc2ggc 0.00131202)
  (COD1_COD2_COD3_atc2ggg 2.92122e-06)
  (COD1_COD2_COD3_atc2ggt 3.1525e-07)
  (COD1_COD2_COD3_atc2gta 0.00342662)
  (COD1_COD2_COD3_atc2gtc 0.137929)
  (COD1_COD2_COD3_atc2gtg 0.0789455)
  (COD1_COD2_COD3_atc2gtt 0.010957)
  (COD1_COD2_COD3_atc2taa 0)
  (COD1_COD2_COD3_atc2tac 0.00427458)
  (COD1_COD2_COD3_atc2tag 0)
  (COD1_COD2_COD3_atc2tat 2.1623e-06)
  (COD1_COD2_COD3_atc2tca 6.0356e-07)
  (COD1_COD2_COD3_atc2tcc 0.00124901)
  (COD1_COD2_COD3_atc2tcg 3.36491e-06)
  (COD1_COD2_COD3_atc2tct 3.59128e-07)
  (COD1_COD2_COD3_atc2tga 0)
  (COD1_COD2_COD3_atc2tgc 0.00229705)
  (COD1_COD2_COD3_atc2tgg 0.00174446)
  (COD1_COD2_COD3_atc2tgt 1.68129e-06)
  (COD1_COD2_COD3_atc2tta 0.000682445)
  (COD1_COD2_COD3_atc2ttc 0.0214779)
  (COD1_COD2_COD3_atc2ttg 0.00751506)
  (COD1_COD2_COD3_atc2ttt 0.00119979)
  (COD1_COD2_COD3_atg2aaa 0.00856697)
  (COD1_COD2_COD3_atg2aac 0.00445611)
  (COD1_COD2_COD3_atg2aag 0.0132855)
  (COD1_COD2_COD3_atg2aat 0.00465501)
  (COD1_COD2_COD3_atg2aca 0.0112028)
  (COD1_COD2_COD3_atg2acc 0.00775854)
  (COD1_COD2_COD3_atg2acg 0.0147106)
  (COD1_COD2_COD3_atg2act 0.00531395)
  (COD1_COD2_COD3_atg2aga 0.00298098)
  (COD1_COD2_COD3_atg2agc 0.00417502)
  (COD1_COD2_COD3_atg2agg 0.00615084)
  (COD1_COD2_COD3_atg2agt 0.00315969)
  (COD1_COD2_COD3_atg2ata 0.0285968)
  (COD1_COD2_COD3_atg2atc 0.0339605)
  (COD1_COD2_COD3_atg2att 0.0286167)
  (COD1_COD2_COD3_atg2caa 0.00761996)
  (COD1_COD2_COD3_atg2cac 0.00180473)
  (COD1_COD2_COD3_atg2cag 0.017313)
  (COD1_COD2_COD3_atg2cat 0.00345524)
  (COD1_COD2_COD3_atg2cca 0.000469691)
  (COD1_COD2_COD3_atg2ccc 8.84977e-05)
  (COD1_COD2_COD3_atg2ccg 0.00236163)
  (COD1_COD2_COD3_atg2cct 0.000851326)
  (COD1_COD2_COD3_atg2cga 7.51952e-05)
  (COD1_COD2_COD3_atg2cgc 0.000435512)
  (COD1_COD2_COD3_atg2cgg 0.00333523)
  (COD1_COD2_COD3_atg2cgt 0.000244141)
  (COD1_COD2_COD3_atg2cta 0.0120402)
  (COD1_COD2_COD3_atg2ctc 0.0241138)
  (COD1_COD2_COD3_atg2ctg 0.063769)
  (COD1_COD2_COD3_atg2ctt 0.0185042)
  (COD1_COD2_COD3_atg2gaa 0.00117347)
  (COD1_COD2_COD3_atg2gac 0.000288328)
  (COD1_COD2_COD3_atg2gag 0.00803771)
  (COD1_COD2_COD3_atg2gat 0.00168855)
  (COD1_COD2_COD3_atg2gca 0.0085551)
  (COD1_COD2_COD3_atg2gcc 0.00454029)
  (COD1_COD2_COD3_atg2gcg 0.0153141)
  (COD1_COD2_COD3_atg2gct 0.00853141)
  (COD1_COD2_COD3_atg2gga 0.000618045)
  (COD1_COD2_COD3_atg2ggc 0.000439759)
  (COD1_COD2_COD3_atg2ggg 0.00419533)
  (COD1_COD2_COD3_atg2ggt 0.000463867)
  (COD1_COD2_COD3_atg2gta 0.00621513)
  (COD1_COD2_COD3_atg2gtc 0.00645664)
  (COD1_COD2_COD3_atg2gtg 0.0316862)
  (COD1_COD2_COD3_atg2gtt 0.00818388)
  (COD1_COD2_COD3_atg2taa 0)
  (COD1_COD2_COD3_atg2tac 0.00501686)
  (COD1_COD2_COD3_atg2tag 0)
  (COD1_COD2_COD3_atg2tat 0.00507632)
  (COD1_COD2_COD3_atg2tca 0.00255228)
  (COD1_COD2_COD3_atg2tcc 0.000455267)
  (COD1_COD2_COD3_atg2tcg 0.00459729)
  (COD1_COD2_COD3_atg2tct 0.00110327)
  (COD1_COD2_COD3_atg2tga 0)
  (COD1_COD2_COD3_atg2tgc 0.00240791)
  (COD1_COD2_COD3_atg2tgg 0.00410046)
  (COD1_COD2_COD3_atg2tgt 0.0019209)
  (COD1_COD2_COD3_atg2tta 0.0229876)
  (COD1_COD2_COD3_atg2ttc 0.0170983)
  (COD1_COD2_COD3_atg2ttg 0.0481607)
  (COD1_COD2_COD3_atg2ttt 0.0145484)
  (COD1_COD2_COD3_att2aaa 0.00689134)
  (COD1_COD2_COD3_att2aac 0.00051648)
  (COD1_COD2_COD3_att2aag 0.0011071)
  (COD1_COD2_COD3_att2aat 0.0053324)
  (COD1_COD2_COD3_att2aca 0.00388465)
  (COD1_COD2_COD3_att2acc 0.000618474)
  (COD1_COD2_COD3_att2acg 0.000969145)
  (COD1_COD2_COD3_att2act 0.0160309)
  (COD1_COD2_COD3_att2aga 0.00113115)
  (COD1_COD2_COD3_att2agc 0.000426845)
  (COD1_COD2_COD3_att2agg 0.000634981)
  (COD1_COD2_COD3_att2agt 0.00370323)
  (COD1_COD2_COD3_att2ata 0.21037)
  (COD1_COD2_COD3_att2atc 0.323536)
  (COD1_COD2_COD3_att2atg 0.0244673)
  (COD1_COD2_COD3_att2caa 0.00135546)
  (COD1_COD2_COD3_att2cac 1.65576e-07)
  (COD1_COD2_COD3_att2cag 8.04806e-05)
  (COD1_COD2_COD3_att2cat 0.00206449)
  (COD1_COD2_COD3_att2cca 2.94985e-05)
  (COD1_COD2_COD3_att2ccc 5.97317e-07)
  (COD1_COD2_COD3_att2ccg 3.26157e-06)
  (COD1_COD2_COD3_att2cct 0.00305418)
  (COD1_COD2_COD3_att2cga 2.04566e-06)
  (COD1_COD2_COD3_att2cgc 3.42814e-08)
  (COD1_COD2_COD3_att2cgg 1.66036e-07)
  (COD1_COD2_COD3_att2cgt 0.000649222)
  (COD1_COD2_COD3_att2cta 0.00977966)
  (COD1_COD2_COD3_att2ctc 0.00309252)
  (COD1_COD2_COD3_att2ctg 0.00254697)
  (COD1_COD2_COD3_att2ctt 0.0505201)
  (COD1_COD2_COD3_att2gaa 0.00133402)
  (COD1_COD2_COD3_att2gac 4.78801e-08)
  (COD1_COD2_COD3_att2gag 0.000101446)
  (COD1_COD2_COD3_att2gat 0.00191426)
  (COD1_COD2_COD3_att2gca 3.76498e-05)
  (COD1_COD2_COD3_att2gcc 5.44336e-07)
  (COD1_COD2_COD3_att2gcg 1.70358e-06)
  (COD1_COD2_COD3_att2gct 0.0101456)
  (COD1_COD2_COD3_att2gga 4.94705e-06)
  (COD1_COD2_COD3_att2ggc 6.88076e-06)
  (COD1_COD2_COD3_att2ggg 8.32527e-07)
  (COD1_COD2_COD3_att2ggt 0.00118245)
  (COD1_COD2_COD3_att2gta 0.0282031)
  (COD1_COD2_COD3_att2gtc 0.00844902)
  (COD1_COD2_COD3_att2gtg 0.0215824)
  (COD1_COD2_COD3_att2gtt 0.124568)
  (COD1_COD2_COD3_att2taa 0)
  (COD1_COD2_COD3_att2tac 2.70864e-06)
  (COD1_COD2_COD3_att2tag 0)
  (COD1_COD2_COD3_att2tat 0.00608301)
  (COD1_COD2_COD3_att2tca 3.78572e-06)
  (COD1_COD2_COD3_att2tcc 4.2619e-07)
  (COD1_COD2_COD3_att2tcg 2.45461e-07)
  (COD1_COD2_COD3_att2tct 0.00203466)
  (COD1_COD2_COD3_att2tga 0)
  (COD1_COD2_COD3_att2tgc 2.85713e-06)
  (COD1_COD2_COD3_att2tgg 0.000998465)
  (COD1_COD2_COD3_att2tgt 0.00277942)
  (COD1_COD2_COD3_att2tta 0.0345062)
  (COD1_COD2_COD3_att2ttc 0.00128057)
  (COD1_COD2_COD3_att2ttg 0.0203335)
  (COD1_COD2_COD3_att2ttt 0.0239804)
  (COD1_COD2_COD3_caa2aaa 0.099995)
  (COD1_COD2_COD3_caa2aac 0.00767297)
  (COD1_COD2_COD3_caa2aag 0.0123188)
  (COD1_COD2_COD3_caa2aat 0.0392001)
  (COD1_COD2_COD3_caa2aca 0.0217929)
  (COD1_COD2_COD3_caa2acc 0.00118985)
  (COD1_COD2_COD3_caa2acg 3.04331e-05)
  (COD1_COD2_COD3_caa2act 0.00473966)
  (COD1_COD2_COD3_caa2aga 0.0121071)
  (COD1_COD2_COD3_caa2agc 0.00318829)
  (COD1_COD2_COD3_caa2agg 2.55202e-05)
  (COD1_COD2_COD3_caa2agt 0.0121492)
  (COD1_COD2_COD3_caa2ata 0.0049995)
  (COD1_COD2_COD3_caa2atc 0.000251353)
  (COD1_COD2_COD3_caa2atg 0.0094008)
  (COD1_COD2_COD3_caa2att 0.00195448)
  (COD1_COD2_COD3_caa2cac 0.00830387)
  (COD1_COD2_COD3_caa2cag 0.371646)
  (COD1_COD2_COD3_caa2cat 0.0264058)
  (COD1_COD2_COD3_caa2cca 0.0117516)
  (COD1_COD2_COD3_caa2ccc 0.000779048)
  (COD1_COD2_COD3_caa2ccg 0.000773131)
  (COD1_COD2_COD3_caa2cct 0.00453714)
  (COD1_COD2_COD3_caa2cga 0.0227341)
  (COD1_COD2_COD3_caa2cgc 0.00217616)
  (COD1_COD2_COD3_caa2cgg 0.00244831)
  (COD1_COD2_COD3_caa2cgt 0.00988872)
  (COD1_COD2_COD3_caa2cta 0.00887523)
  (COD1_COD2_COD3_caa2ctc 0.00120646)
  (COD1_COD2_COD3_caa2ctg 0.000951176)
  (COD1_COD2_COD3_caa2ctt 0.00802657)
  (COD1_COD2_COD3_caa2gaa 0.106432)
  (COD1_COD2_COD3_caa2gac 0.00306903)
  (COD1_COD2_COD3_caa2gag 0.0105107)
  (COD1_COD2_COD3_caa2gat 0.020207)
  (COD1_COD2_COD3_caa2gca 0.0210505)
  (COD1_COD2_COD3_caa2gcc 0.00050515)
  (COD1_COD2_COD3_caa2gcg 2.18198e-05)
  (COD1_COD2_COD3_caa2gct 0.00516148)
  (COD1_COD2_COD3_caa2gga 0.00903663)
  (COD1_COD2_COD3_caa2ggc 0.000383629)
  (COD1_COD2_COD3_caa2ggg 3.11588e-05)
  (COD1_COD2_COD3_caa2ggt 0.000723359)
  (COD1_COD2_COD3_caa2gta 0.00662897)
  (COD1_COD2_COD3_caa2gtc 0.000124549)
  (COD1_COD2_COD3_caa2gtg 1.06042e-05)
  (COD1_COD2_COD3_caa2gtt 0.00286947)
  (COD1_COD2_COD3_caa2taa 0)
  (COD1_COD2_COD3_caa2tac 0.00132389)
  (COD1_COD2_COD3_caa2tag 0)
  (COD1_COD2_COD3_caa2tat 0.00838485)
  (COD1_COD2_COD3_caa2tca 0.0214333)
  (COD1_COD2_COD3_caa2tcc 0.000865607)
  (COD1_COD2_COD3_caa2tcg 1.90216e-05)
  (COD1_COD2_COD3_caa2tct 0.00599926)
  (COD1_COD2_COD3_caa2tga 0)
  (COD1_COD2_COD3_caa2tgc 0.000239116)
  (COD1_COD2_COD3_caa2tgg 0.000703091)
  (COD1_COD2_COD3_caa2tgt 0.00105174)
  (COD1_COD2_COD3_caa2tta 0.00934234)
  (COD1_COD2_COD3_caa2ttc 0.000354728)
  (COD1_COD2_COD3_caa2ttg 0.000117667)
  (COD1_COD2_COD3_caa2ttt 0.00270529)
  (COD1_COD2_COD3_cac2aaa 0.00446229)
  (COD1_COD2_COD3_cac2aac 0.0447801)
  (COD1_COD2_COD3_cac2aag 0.0152611)
  (COD1_COD2_COD3_cac2aat 0.00420115)
  (COD1_COD2_COD3_cac2aca 0.000287132)
  (COD1_COD2_COD3_cac2acc 0.017337)
  (COD1_COD2_COD3_cac2acg 0.00110174)
  (COD1_COD2_COD3_cac2act 3.84923e-06)
  (COD1_COD2_COD3_cac2aga 0.000248753)
  (COD1_COD2_COD3_cac2agc 0.017579)
  (COD1_COD2_COD3_cac2agg 0.00114773)
  (COD1_COD2_COD3_cac2agt 4.72911e-06)
  (COD1_COD2_COD3_cac2ata 3.55184e-06)
  (COD1_COD2_COD3_cac2atc 0.0030415)
  (COD1_COD2_COD3_cac2atg 0.00346243)
  (COD1_COD2_COD3_cac2att 3.71535e-07)
  (COD1_COD2_COD3_cac2caa 0.0129133)
  (COD1_COD2_COD3_cac2cag 0.0519044)
  (COD1_COD2_COD3_cac2cat 0.412939)
  (COD1_COD2_COD3_cac2cca 0.00100332)
  (COD1_COD2_COD3_cac2ccc 0.00760414)
  (COD1_COD2_COD3_cac2ccg 0.00444377)
  (COD1_COD2_COD3_cac2cct 0.000642507)
  (COD1_COD2_COD3_cac2cga 0.00253465)
  (COD1_COD2_COD3_cac2cgc 0.0361967)
  (COD1_COD2_COD3_cac2cgg 0.0180232)
  (COD1_COD2_COD3_cac2cgt 0.00235509)
  (COD1_COD2_COD3_cac2cta 0.00044098)
  (COD1_COD2_COD3_cac2ctc 0.0103804)
  (COD1_COD2_COD3_cac2ctg 0.00940732)
  (COD1_COD2_COD3_cac2ctt 0.000889326)
  (COD1_COD2_COD3_cac2gaa 0.00366493)
  (COD1_COD2_COD3_cac2gac 0.0246717)
  (COD1_COD2_COD3_cac2gag 0.017325)
  (COD1_COD2_COD3_cac2gat 0.00133773)
  (COD1_COD2_COD3_cac2gca 8.17172e-05)
  (COD1_COD2_COD3_cac2gcc 0.0186717)
  (COD1_COD2_COD3_cac2gcg 0.00347686)
  (COD1_COD2_COD3_cac2gct 2.08865e-06)
  (COD1_COD2_COD3_cac2gga 8.22286e-05)
  (COD1_COD2_COD3_cac2ggc 0.0134151)
  (COD1_COD2_COD3_cac2ggg 0.000549726)
  (COD1_COD2_COD3_cac2ggt 1.07592e-06)
  (COD1_COD2_COD3_cac2gta 3.09007e-05)
  (COD1_COD2_COD3_cac2gtc 0.00904916)
  (COD1_COD2_COD3_cac2gtg 0.000997774)
  (COD1_COD2_COD3_cac2gtt 1.65929e-06)
  (COD1_COD2_COD3_cac2taa 0)
  (COD1_COD2_COD3_cac2tac 0.0533636)
  (COD1_COD2_COD3_cac2tag 0)
  (COD1_COD2_COD3_cac2tat 0.00458139)
  (COD1_COD2_COD3_cac2tca 0.000150286)
  (COD1_COD2_COD3_cac2tcc 0.0123188)
  (COD1_COD2_COD3_cac2tcg 0.000999158)
  (COD1_COD2_COD3_cac2tct 6.57763e-06)
  (COD1_COD2_COD3_cac2tga 0)
  (COD1_COD2_COD3_cac2tgc 0.00358205)
  (COD1_COD2_COD3_cac2tgg 0.00470692)
  (COD1_COD2_COD3_cac2tgt 1.42165e-06)
  (COD1_COD2_COD3_cac2tta 1.64099e-06)
  (COD1_COD2_COD3_cac2ttc 0.016266)
  (COD1_COD2_COD3_cac2ttg 3.41679e-05)
  (COD1_COD2_COD3_cac2ttt 9.78498e-06)
  (COD1_COD2_COD3_cag2aaa 0.00887523)
  (COD1_COD2_COD3_cag2aac 0.0200257)
  (COD1_COD2_COD3_cag2aag 0.0610443)
  (COD1_COD2_COD3_cag2aat 0.00461006)
  (COD1_COD2_COD3_cag2aca 1.67904e-05)
  (COD1_COD2_COD3_cag2acc 0.0110485)
  (COD1_COD2_COD3_cag2acg 0.01888)
  (COD1_COD2_COD3_cag2act 0.000368512)
  (COD1_COD2_COD3_cag2aga 2.9764e-06)
  (COD1_COD2_COD3_cag2agc 0.0194377)
  (COD1_COD2_COD3_cag2agg 0.00779627)
  (COD1_COD2_COD3_cag2agt 0.000858437)
  (COD1_COD2_COD3_cag2ata 7.16763e-07)
  (COD1_COD2_COD3_cag2atc 0.00251714)
  (COD1_COD2_COD3_cag2atg 0.017689)
  (COD1_COD2_COD3_cag2att 9.61069e-05)
  (COD1_COD2_COD3_cag2caa 0.307786)
  (COD1_COD2_COD3_cag2cac 0.0276419)
  (COD1_COD2_COD3_cag2cat 0.0118087)
  (COD1_COD2_COD3_cag2cca 0.00205877)
  (COD1_COD2_COD3_cag2ccc 0.0055128)
  (COD1_COD2_COD3_cag2ccg 0.00873486)
  (COD1_COD2_COD3_cag2cct 0.00112022)
  (COD1_COD2_COD3_cag2cga 0.0029686)
  (COD1_COD2_COD3_cag2cgc 0.0296464)
  (COD1_COD2_COD3_cag2cgg 0.0384999)
  (COD1_COD2_COD3_cag2cgt 0.00289544)
  (COD1_COD2_COD3_cag2cta 0.000775815)
  (COD1_COD2_COD3_cag2ctc 0.00710475)
  (COD1_COD2_COD3_cag2ctg 0.0200674)
  (COD1_COD2_COD3_cag2ctt 0.00119979)
  (COD1_COD2_COD3_cag2gaa 0.0185299)
  (COD1_COD2_COD3_cag2gac 0.0234869)
  (COD1_COD2_COD3_cag2gag 0.0929759)
  (COD1_COD2_COD3_cag2gat 0.00374453)
  (COD1_COD2_COD3_cag2gca 3.64177e-05)
  (COD1_COD2_COD3_cag2gcc 0.0170274)
  (COD1_COD2_COD3_cag2gcg 0.0282619)
  (COD1_COD2_COD3_cag2gct 0.000774203)
  (COD1_COD2_COD3_cag2gga 4.9402e-06)
  (COD1_COD2_COD3_cag2ggc 0.00796009)
  (COD1_COD2_COD3_cag2ggg 0.00648355)
  (COD1_COD2_COD3_cag2ggt 5.70664e-05)
  (COD1_COD2_COD3_cag2gta 1.75877e-06)
  (COD1_COD2_COD3_cag2gtc 0.00127702)
  (COD1_COD2_COD3_cag2gtg 0.0107613)
  (COD1_COD2_COD3_cag2gtt 0.000182603)
  (COD1_COD2_COD3_cag2taa 0)
  (COD1_COD2_COD3_cag2tac 0.00555114)
  (COD1_COD2_COD3_cag2tag 0)
  (COD1_COD2_COD3_cag2tat 0.00158753)
  (COD1_COD2_COD3_cag2tca 2.22012e-05)
  (COD1_COD2_COD3_cag2tcc 0.00455605)
  (COD1_COD2_COD3_cag2tcg 0.0132579)
  (COD1_COD2_COD3_cag2tct 0.000581473)
  (COD1_COD2_COD3_cag2tga 0)
  (COD1_COD2_COD3_cag2tgc 0.000782837)
  (COD1_COD2_COD3_cag2tgg 0.00315532)
  (COD1_COD2_COD3_cag2tgt 0.00037288)
  (COD1_COD2_COD3_cag2tta 1.59383e-07)
  (COD1_COD2_COD3_cag2ttc 0.00228911)
  (COD1_COD2_COD3_cag2ttg 0.00394161)
  (COD1_COD2_COD3_cag2ttt 0.000209611)
  (COD1_COD2_COD3_cat2aaa 0.0279695)
  (COD1_COD2_COD3_cat2aac 0.00936828)
  (COD1_COD2_COD3_cat2aag 0.00866851)
  (COD1_COD2_COD3_cat2aat 0.059954)
  (COD1_COD2_COD3_cat2aca 0.00108131)
  (COD1_COD2_COD3_cat2acc 2.09455e-05)
  (COD1_COD2_COD3_cat2acg 0.000822327)
  (COD1_COD2_COD3_cat2act 0.0113828)
  (COD1_COD2_COD3_cat2aga 0.00400773)
  (COD1_COD2_COD3_cat2agc 6.99169e-05)
  (COD1_COD2_COD3_cat2agg 0.000957128)
  (COD1_COD2_COD3_cat2agt 0.0133409)
  (COD1_COD2_COD3_cat2ata 0.000513624)
  (COD1_COD2_COD3_cat2atc 1.10929e-05)
  (COD1_COD2_COD3_cat2atg 0.00685798)
  (COD1_COD2_COD3_cat2att 0.0047892)
  (COD1_COD2_COD3_cat2caa 0.0424821)
  (COD1_COD2_COD3_cat2cac 0.427205)
  (COD1_COD2_COD3_cat2cag 0.0229399)
  (COD1_COD2_COD3_cat2cca 0.00319715)
  (COD1_COD2_COD3_cat2ccc 0.000949858)
  (COD1_COD2_COD3_cat2ccg 0.000659655)
  (COD1_COD2_COD3_cat2cct 0.0080824)
  (COD1_COD2_COD3_cat2cga 0.00651509)
  (COD1_COD2_COD3_cat2cgc 0.00248593)
  (COD1_COD2_COD3_cat2cgg 0.00258253)
  (COD1_COD2_COD3_cat2cgt 0.0209051)
  (COD1_COD2_COD3_cat2cta 0.00322832)
  (COD1_COD2_COD3_cat2ctc 0.000963786)
  (COD1_COD2_COD3_cat2ctg 0.00127525)
  (COD1_COD2_COD3_cat2ctt 0.0100895)
  (COD1_COD2_COD3_cat2gaa 0.015625)
  (COD1_COD2_COD3_cat2gac 0.00255228)
  (COD1_COD2_COD3_cat2gag 0.00487291)
  (COD1_COD2_COD3_cat2gat 0.0213592)
  (COD1_COD2_COD3_cat2gca 0.000647874)
  (COD1_COD2_COD3_cat2gcc 1.46982e-05)
  (COD1_COD2_COD3_cat2gcg 0.0014659)
  (COD1_COD2_COD3_cat2gct 0.00795457)
  (COD1_COD2_COD3_cat2gga 0.00159968)
  (COD1_COD2_COD3_cat2ggc 2.096e-05)
  (COD1_COD2_COD3_cat2ggg 0.000859032)
  (COD1_COD2_COD3_cat2ggt 0.00655586)
  (COD1_COD2_COD3_cat2gta 0.000214461)
  (COD1_COD2_COD3_cat2gtc 7.67182e-06)
  (COD1_COD2_COD3_cat2gtg 0.000729907)
  (COD1_COD2_COD3_cat2gtt 0.0064343)
  (COD1_COD2_COD3_cat2taa 0)
  (COD1_COD2_COD3_cat2tac 0.00568746)
  (COD1_COD2_COD3_cat2tag 0)
  (COD1_COD2_COD3_cat2tat 0.0532528)
  (COD1_COD2_COD3_cat2tca 0.00132849)
  (COD1_COD2_COD3_cat2tcc 2.28571e-05)
  (COD1_COD2_COD3_cat2tcg 0.000642953)
  (COD1_COD2_COD3_cat2tct 0.0148027)
  (COD1_COD2_COD3_cat2tga 0)
  (COD1_COD2_COD3_cat2tgc 1.11392e-05)
  (COD1_COD2_COD3_cat2tgg 0.00248593)
  (COD1_COD2_COD3_cat2tgt 0.00408627)
  (COD1_COD2_COD3_cat2tta 0.000429814)
  (COD1_COD2_COD3_cat2ttc 2.81209e-05)
  (COD1_COD2_COD3_cat2ttg 0.000632784)
  (COD1_COD2_COD3_cat2ttt 0.013678)
  (COD1_COD2_COD3_cca2aaa 0.0324194)
  (COD1_COD2_COD3_cca2aac 0.00150711)
  (COD1_COD2_COD3_cca2aag 0.000320586)
  (COD1_COD2_COD3_cca2aat 0.00214174)
  (COD1_COD2_COD3_cca2aca 0.0133594)
  (COD1_COD2_COD3_cca2acc 0.000485581)
  (COD1_COD2_COD3_cca2acg 0.000892414)
  (COD1_COD2_COD3_cca2act 0.00437349)
  (COD1_COD2_COD3_cca2aga 0.00481249)
  (COD1_COD2_COD3_cca2agc 7.14355e-05)
  (COD1_COD2_COD3_cca2agg 6.27946e-05)
  (COD1_COD2_COD3_cca2agt 0.000216702)
  (COD1_COD2_COD3_cca2ata 0.00615938)
  (COD1_COD2_COD3_cca2atc 1.07746e-05)
  (COD1_COD2_COD3_cca2atg 0.00089056)
  (COD1_COD2_COD3_cca2att 6.54159e-05)
  (COD1_COD2_COD3_cca2caa 0.0180733)
  (COD1_COD2_COD3_cca2cac 0.000992256)
  (COD1_COD2_COD3_cca2cag 0.00382321)
  (COD1_COD2_COD3_cca2cat 0.00305418)
  (COD1_COD2_COD3_cca2ccc 0.160651)
  (COD1_COD2_COD3_cca2ccg 0.184539)
  (COD1_COD2_COD3_cca2cct 0.426909)
  (COD1_COD2_COD3_cca2cga 0.00301422)
  (COD1_COD2_COD3_cca2cgc 2.44298e-05)
  (COD1_COD2_COD3_cca2cgg 8.38982e-05)
  (COD1_COD2_COD3_cca2cgt 0.000538786)
  (COD1_COD2_COD3_cca2cta 0.0079051)
  (COD1_COD2_COD3_cca2ctc 0.000279474)
  (COD1_COD2_COD3_cca2ctg 0.00116617)
  (COD1_COD2_COD3_cca2ctt 0.000996391)
  (COD1_COD2_COD3_cca2gaa 0.0307771)
  (COD1_COD2_COD3_cca2gac 9.39335e-05)
  (COD1_COD2_COD3_cca2gag 5.58145e-05)
  (COD1_COD2_COD3_cca2gat 0.00422744)
  (COD1_COD2_COD3_cca2gca 0.0204324)
  (COD1_COD2_COD3_cca2gcc 0.000351789)
  (COD1_COD2_COD3_cca2gcg 0.000325738)
  (COD1_COD2_COD3_cca2gct 0.00656495)
  (COD1_COD2_COD3_cca2gga 0.00911842)
  (COD1_COD2_COD3_cca2ggc 1.48408e-06)
  (COD1_COD2_COD3_cca2ggg 1.27336e-05)
  (COD1_COD2_COD3_cca2ggt 4.28009e-05)
  (COD1_COD2_COD3_cca2gta 0.0079988)
  (COD1_COD2_COD3_cca2gtc 1.05675e-05)
  (COD1_COD2_COD3_cca2gtg 1.59068e-05)
  (COD1_COD2_COD3_cca2gtt 0.000176017)
  (COD1_COD2_COD3_cca2taa 0)
  (COD1_COD2_COD3_cca2tac 0.000432204)
  (COD1_COD2_COD3_cca2tag 0)
  (COD1_COD2_COD3_cca2tat 0.000553933)
  (COD1_COD2_COD3_cca2tca 0.0323745)
  (COD1_COD2_COD3_cca2tcc 0.00149359)
  (COD1_COD2_COD3_cca2tcg 0.00132206)
  (COD1_COD2_COD3_cca2tct 0.00575487)
  (COD1_COD2_COD3_cca2tga 0)
  (COD1_COD2_COD3_cca2tgc 7.62977e-05)
  (COD1_COD2_COD3_cca2tgg 0.000896754)
  (COD1_COD2_COD3_cca2tgt 1.75399e-05)
  (COD1_COD2_COD3_cca2tta 0.00423037)
  (COD1_COD2_COD3_cca2ttc 0.000140416)
  (COD1_COD2_COD3_cca2ttg 8.73399e-05)
  (COD1_COD2_COD3_cca2ttt 0.000540282)
  (COD1_COD2_COD3_ccc2aaa 0.000384428)
  (COD1_COD2_COD3_ccc2aac 0.0108137)
  (COD1_COD2_COD3_ccc2aag 0.00811046)
  (COD1_COD2_COD3_ccc2aat 3.41188e-06)
  (COD1_COD2_COD3_ccc2aca 0.000344072)
  (COD1_COD2_COD3_ccc2acc 0.0155817)
  (COD1_COD2_COD3_ccc2acg 0.00465178)
  (COD1_COD2_COD3_ccc2act 0.000681499)
  (COD1_COD2_COD3_ccc2aga 3.876e-06)
  (COD1_COD2_COD3_ccc2agc 0.0183383)
  (COD1_COD2_COD3_ccc2agg 0.000225904)
  (COD1_COD2_COD3_ccc2agt 3.4261e-06)
  (COD1_COD2_COD3_ccc2ata 3.25915e-07)
  (COD1_COD2_COD3_ccc2atc 0.00642539)
  (COD1_COD2_COD3_ccc2atg 0.000180089)
  (COD1_COD2_COD3_ccc2att 1.42067e-06)
  (COD1_COD2_COD3_ccc2caa 0.00128501)
  (COD1_COD2_COD3_ccc2cac 0.00806561)
  (COD1_COD2_COD3_ccc2cag 0.0109798)
  (COD1_COD2_COD3_ccc2cat 0.000973859)
  (COD1_COD2_COD3_ccc2cca 0.172301)
  (COD1_COD2_COD3_ccc2ccg 0.408102)
  (COD1_COD2_COD3_ccc2cct 0.245025)
  (COD1_COD2_COD3_ccc2cga 0.000741123)
  (COD1_COD2_COD3_ccc2cgc 0.00823509)
  (COD1_COD2_COD3_ccc2cgg 0.0073197)
  (COD1_COD2_COD3_ccc2cgt 0.0010554)
  (COD1_COD2_COD3_ccc2cta 0.000294795)
  (COD1_COD2_COD3_ccc2ctc 0.010904)
  (COD1_COD2_COD3_ccc2ctg 0.00426866)
  (COD1_COD2_COD3_ccc2ctt 0.000403261)
  (COD1_COD2_COD3_ccc2gaa 0.00362953)
  (COD1_COD2_COD3_ccc2gac 0.0246888)
  (COD1_COD2_COD3_ccc2gag 0.00901161)
  (COD1_COD2_COD3_ccc2gat 2.17443e-05)
  (COD1_COD2_COD3_ccc2gca 0.00115251)
  (COD1_COD2_COD3_ccc2gcc 0.0374212)
  (COD1_COD2_COD3_ccc2gcg 0.00764112)
  (COD1_COD2_COD3_ccc2gct 0.00138011)
  (COD1_COD2_COD3_ccc2gga 5.94868e-06)
  (COD1_COD2_COD3_ccc2ggc 0.0150718)
  (COD1_COD2_COD3_ccc2ggg 1.42963e-05)
  (COD1_COD2_COD3_ccc2ggt 4.44619e-06)
  (COD1_COD2_COD3_ccc2gta 1.58509e-06)
  (COD1_COD2_COD3_ccc2gtc 0.0140042)
  (COD1_COD2_COD3_ccc2gtg 3.4741e-05)
  (COD1_COD2_COD3_ccc2gtt 2.8139e-06)
  (COD1_COD2_COD3_ccc2taa 0)
  (COD1_COD2_COD3_ccc2tac 0.00518658)
  (COD1_COD2_COD3_ccc2tag 0)
  (COD1_COD2_COD3_ccc2tat 1.53004e-06)
  (COD1_COD2_COD3_ccc2tca 0.000750427)
  (COD1_COD2_COD3_ccc2tcc 0.0267744)
  (COD1_COD2_COD3_ccc2tcg 0.00377319)
  (COD1_COD2_COD3_ccc2tct 0.00119481)
  (COD1_COD2_COD3_ccc2tga 0)
  (COD1_COD2_COD3_ccc2tgc 0.00160079)
  (COD1_COD2_COD3_ccc2tgg 0.000415173)
  (COD1_COD2_COD3_ccc2tgt 3.12422e-07)
  (COD1_COD2_COD3_ccc2tta 1.47171e-07)
  (COD1_COD2_COD3_ccc2ttc 0.00609989)
  (COD1_COD2_COD3_ccc2ttg 5.87085e-06)
  (COD1_COD2_COD3_ccc2ttt 2.84725e-06)
  (COD1_COD2_COD3_ccg2aaa 1.19801e-05)
  (COD1_COD2_COD3_ccg2aac 0.00193963)
  (COD1_COD2_COD3_ccg2aag 0.0149989)
  (COD1_COD2_COD3_ccg2aat 6.03202e-05)
  (COD1_COD2_COD3_ccg2aca 0.000196934)
  (COD1_COD2_COD3_ccg2acc 0.00942691)
  (COD1_COD2_COD3_ccg2acg 0.0102801)
  (COD1_COD2_COD3_ccg2act 0.000145368)
  (COD1_COD2_COD3_ccg2aga 6.26548e-08)
  (COD1_COD2_COD3_ccg2agc 0.00143473)
  (COD1_COD2_COD3_ccg2agg 0.00185933)
  (COD1_COD2_COD3_ccg2agt 1.06116e-05)
  (COD1_COD2_COD3_ccg2ata 4.81487e-07)
  (COD1_COD2_COD3_ccg2atc 0.00051648)
  (COD1_COD2_COD3_ccg2atg 0.00410046)
  (COD1_COD2_COD3_ccg2att 6.61879e-06)
  (COD1_COD2_COD3_ccg2caa 0.00108808)
  (COD1_COD2_COD3_ccg2cac 0.00402164)
  (COD1_COD2_COD3_ccg2cag 0.0148438)
  (COD1_COD2_COD3_ccg2cat 0.000576657)
  (COD1_COD2_COD3_ccg2cca 0.168989)
  (COD1_COD2_COD3_ccg2ccc 0.348203)
  (COD1_COD2_COD3_ccg2cct 0.107321)
  (COD1_COD2_COD3_ccg2cga 0.000437934)
  (COD1_COD2_COD3_ccg2cgc 0.0116623)
  (COD1_COD2_COD3_ccg2cgg 0.00950564)
  (COD1_COD2_COD3_ccg2cgt 0.000379398)
  (COD1_COD2_COD3_ccg2cta 0.000121227)
  (COD1_COD2_COD3_ccg2ctc 0.00198589)
  (COD1_COD2_COD3_ccg2ctg 0.0141114)
  (COD1_COD2_COD3_ccg2ctt 0.000120891)
  (COD1_COD2_COD3_ccg2gaa 0.000187607)
  (COD1_COD2_COD3_ccg2gac 0.0162998)
  (COD1_COD2_COD3_ccg2gag 0.0303113)
  (COD1_COD2_COD3_ccg2gat 0.000901741)
  (COD1_COD2_COD3_ccg2gca 0.00144371)
  (COD1_COD2_COD3_ccg2gcc 0.0260423)
  (COD1_COD2_COD3_ccg2gcg 0.0356736)
  (COD1_COD2_COD3_ccg2gct 0.000255569)
  (COD1_COD2_COD3_ccg2gga 2.6918e-06)
  (COD1_COD2_COD3_ccg2ggc 0.00936828)
  (COD1_COD2_COD3_ccg2ggg 0.00718897)
  (COD1_COD2_COD3_ccg2ggt 2.15493e-05)
  (COD1_COD2_COD3_ccg2gta 1.0465e-06)
  (COD1_COD2_COD3_ccg2gtc 0.000582683)
  (COD1_COD2_COD3_ccg2gtg 0.0136307)
  (COD1_COD2_COD3_ccg2gtt 4.5239e-06)
  (COD1_COD2_COD3_ccg2taa 0)
  (COD1_COD2_COD3_ccg2tac 0.00062364)
  (COD1_COD2_COD3_ccg2tag 0)
  (COD1_COD2_COD3_ccg2tat 7.21321e-05)
  (COD1_COD2_COD3_ccg2tca 0.000673049)
  (COD1_COD2_COD3_ccg2tcc 0.00399386)
  (COD1_COD2_COD3_ccg2tcg 0.0180983)
  (COD1_COD2_COD3_ccg2tct 0.000253628)
  (COD1_COD2_COD3_ccg2tga 0)
  (COD1_COD2_COD3_ccg2tgc 3.40969e-05)
  (COD1_COD2_COD3_ccg2tgg 0.00275067)
  (COD1_COD2_COD3_ccg2tgt 8.11447e-07)
  (COD1_COD2_COD3_ccg2tta 3.38798e-08)
  (COD1_COD2_COD3_ccg2ttc 0.00169089)
  (COD1_COD2_COD3_ccg2ttg 0.00177619)
  (COD1_COD2_COD3_ccg2ttt 8.72189e-05)
  (COD1_COD2_COD3_cct2aaa 0.00768894)
  (COD1_COD2_COD3_cct2aac 1.87467e-05)
  (COD1_COD2_COD3_cct2aag 0.0058962)
  (COD1_COD2_COD3_cct2aat 0.0175425)
  (COD1_COD2_COD3_cct2aca 0.0048359)
  (COD1_COD2_COD3_cct2acc 0.000789376)
  (COD1_COD2_COD3_cct2acg 0.000851917)
  (COD1_COD2_COD3_cct2act 0.0132947)
  (COD1_COD2_COD3_cct2aga 0.000111784)
  (COD1_COD2_COD3_cct2agc 1.75035e-05)
  (COD1_COD2_COD3_cct2agg 0.000410594)
  (COD1_COD2_COD3_cct2agt 0.0106722)
  (COD1_COD2_COD3_cct2ata 2.61107e-05)
  (COD1_COD2_COD3_cct2atc 3.05372e-06)
  (COD1_COD2_COD3_cct2atg 0.00172882)
  (COD1_COD2_COD3_cct2att 0.00725404)
  (COD1_COD2_COD3_cct2caa 0.0074735)
  (COD1_COD2_COD3_cct2cac 0.000680084)
  (COD1_COD2_COD3_cct2cag 0.00222651)
  (COD1_COD2_COD3_cct2cat 0.00827514)
  (COD1_COD2_COD3_cct2cca 0.457233)
  (COD1_COD2_COD3_cct2ccc 0.244685)
  (COD1_COD2_COD3_cct2ccg 0.125608)
  (COD1_COD2_COD3_cct2cga 0.00169676)
  (COD1_COD2_COD3_cct2cgc 3.46689e-05)
  (COD1_COD2_COD3_cct2cgg 8.05922e-05)
  (COD1_COD2_COD3_cct2cgt 0.00425095)
  (COD1_COD2_COD3_cct2cta 0.00133495)
  (COD1_COD2_COD3_cct2ctc 0.000609536)
  (COD1_COD2_COD3_cct2ctg 0.000410025)
  (COD1_COD2_COD3_cct2ctt 0.0124821)
  (COD1_COD2_COD3_cct2gaa 0.00638101)
  (COD1_COD2_COD3_cct2gac 3.02214e-06)
  (COD1_COD2_COD3_cct2gag 0.00105466)
  (COD1_COD2_COD3_cct2gat 0.0189324)
  (COD1_COD2_COD3_cct2gca 0.00555114)
  (COD1_COD2_COD3_cct2gcc 0.000592869)
  (COD1_COD2_COD3_cct2gcg 0.000257168)
  (COD1_COD2_COD3_cct2gct 0.0240637)
  (COD1_COD2_COD3_cct2gga 5.7863e-05)
  (COD1_COD2_COD3_cct2ggc 1.38854e-06)
  (COD1_COD2_COD3_cct2ggg 1.92204e-05)
  (COD1_COD2_COD3_cct2ggt 0.00707527)
  (COD1_COD2_COD3_cct2gta 1.30282e-05)
  (COD1_COD2_COD3_cct2gtc 3.85724e-06)
  (COD1_COD2_COD3_cct2gtg 9.27642e-06)
  (COD1_COD2_COD3_cct2gtt 0.0134803)
  (COD1_COD2_COD3_cct2taa 0)
  (COD1_COD2_COD3_cct2tac 8.36038e-06)
  (COD1_COD2_COD3_cct2tag 0)
  (COD1_COD2_COD3_cct2tat 0.00582713)
  (COD1_COD2_COD3_cct2tca 0.00564426)
  (COD1_COD2_COD3_cct2tcc 0.00211372)
  (COD1_COD2_COD3_cct2tcg 0.000686715)
  (COD1_COD2_COD3_cct2tct 0.0335859)
  (COD1_COD2_COD3_cct2tga 0)
  (COD1_COD2_COD3_cct2tgc 2.10609e-06)
  (COD1_COD2_COD3_cct2tgg 0.000788282)
  (COD1_COD2_COD3_cct2tgt 0.00154519)
  (COD1_COD2_COD3_cct2tta 2.16241e-05)
  (COD1_COD2_COD3_cct2ttc 3.26836e-06)
  (COD1_COD2_COD3_cct2ttg 5.50843e-05)
  (COD1_COD2_COD3_cct2ttt 0.0069489)
  (COD1_COD2_COD3_cga2aaa 0.0953251)
  (COD1_COD2_COD3_cga2aac 0.000336525)
  (COD1_COD2_COD3_cga2aag 0.00092005)
  (COD1_COD2_COD3_cga2aat 0.000222794)
  (COD1_COD2_COD3_cga2aca 0.0047331)
  (COD1_COD2_COD3_cga2acc 3.58915e-05)
  (COD1_COD2_COD3_cga2acg 3.01601e-05)
  (COD1_COD2_COD3_cga2act 1.24112e-05)
  (COD1_COD2_COD3_cga2aga 0.311434)
  (COD1_COD2_COD3_cga2agc 0.00485942)
  (COD1_COD2_COD3_cga2agg 0.103809)
  (COD1_COD2_COD3_cga2agt 0.00547472)
  (COD1_COD2_COD3_cga2ata 0.00156242)
  (COD1_COD2_COD3_cga2atc 2.83165e-05)
  (COD1_COD2_COD3_cga2atg 0.000331202)
  (COD1_COD2_COD3_cga2att 1.0531e-05)
  (COD1_COD2_COD3_cga2caa 0.0811649)
  (COD1_COD2_COD3_cga2cac 0.00581905)
  (COD1_COD2_COD3_cga2cag 0.0127974)
  (COD1_COD2_COD3_cga2cat 0.0144479)
  (COD1_COD2_COD3_cga2cca 0.00699723)
  (COD1_COD2_COD3_cga2ccc 0.00160412)
  (COD1_COD2_COD3_cga2ccg 0.00111094)
  (COD1_COD2_COD3_cga2cct 0.0036802)
  (COD1_COD2_COD3_cga2cgc 0.18518)
  (COD1_COD2_COD3_cga2cgg 0.249481)
  (COD1_COD2_COD3_cga2cgt 0.336342)
  (COD1_COD2_COD3_cga2cta 0.00675888)
  (COD1_COD2_COD3_cga2ctc 0.0018879)
  (COD1_COD2_COD3_cga2ctg 0.00119647)
  (COD1_COD2_COD3_cga2ctt 0.00644323)
  (COD1_COD2_COD3_cga2gaa 0.0152083)
  (COD1_COD2_COD3_cga2gac 0.00025398)
  (COD1_COD2_COD3_cga2gag 3.13321e-05)
  (COD1_COD2_COD3_cga2gat 0.00020191)
  (COD1_COD2_COD3_cga2gca 0.0047331)
  (COD1_COD2_COD3_cga2gcc 1.21473e-05)
  (COD1_COD2_COD3_cga2gcg 9.96983e-06)
  (COD1_COD2_COD3_cga2gct 3.59662e-05)
  (COD1_COD2_COD3_cga2gga 0.00405242)
  (COD1_COD2_COD3_cga2ggc 0.000210338)
  (COD1_COD2_COD3_cga2ggg 0.0014659)
  (COD1_COD2_COD3_cga2ggt 0.000708473)
  (COD1_COD2_COD3_cga2gta 0.00278714)
  (COD1_COD2_COD3_cga2gtc 3.44055e-05)
  (COD1_COD2_COD3_cga2gtg 1.89821e-05)
  (COD1_COD2_COD3_cga2gtt 7.22321e-05)
  (COD1_COD2_COD3_cga2taa 0)
  (COD1_COD2_COD3_cga2tac 0.00086321)
  (COD1_COD2_COD3_cga2tag 0)
  (COD1_COD2_COD3_cga2tat 0.00075722)
  (COD1_COD2_COD3_cga2tca 0.00577485)
  (COD1_COD2_COD3_cga2tcc 4.37912e-05)
  (COD1_COD2_COD3_cga2tcg 2.76378e-05)
  (COD1_COD2_COD3_cga2tct 7.17829e-05)
  (COD1_COD2_COD3_cga2tga 0)
  (COD1_COD2_COD3_cga2tgc 0.00123267)
  (COD1_COD2_COD3_cga2tgg 0.00186708)
  (COD1_COD2_COD3_cga2tgt 0.00149981)
  (COD1_COD2_COD3_cga2tta 0.0022142)
  (COD1_COD2_COD3_cga2ttc 0.00034217)
  (COD1_COD2_COD3_cga2ttg 2.43959e-05)
  (COD1_COD2_COD3_cga2ttt 0.000645185)
  (COD1_COD2_COD3_cgc2aaa 0.00922011)
  (COD1_COD2_COD3_cgc2aac 0.0175425)
  (COD1_COD2_COD3_cgc2aag 0.072695)
  (COD1_COD2_COD3_cgc2aat 1.29915e-06)
  (COD1_COD2_COD3_cgc2aca 5.48855e-08)
  (COD1_COD2_COD3_cgc2acc 0.0264975)
  (COD1_COD2_COD3_cgc2acg 0.00015249)
  (COD1_COD2_COD3_cgc2act 5.9853e-08)
  (COD1_COD2_COD3_cgc2aga 0.0108587)
  (COD1_COD2_COD3_cgc2agc 0.0180983)
  (COD1_COD2_COD3_cgc2agg 0.0277571)
  (COD1_COD2_COD3_cgc2agt 0.000187087)
  (COD1_COD2_COD3_cgc2ata 3.25206e-09)
  (COD1_COD2_COD3_cgc2atc 0.00567564)
  (COD1_COD2_COD3_cgc2atg 0.000631469)
  (COD1_COD2_COD3_cgc2att 5.81357e-08)
  (COD1_COD2_COD3_cgc2caa 0.00255759)
  (COD1_COD2_COD3_cgc2cac 0.0273749)
  (COD1_COD2_COD3_cgc2cag 0.042101)
  (COD1_COD2_COD3_cgc2cat 0.00181603)
  (COD1_COD2_COD3_cgc2cca 1.86819e-05)
  (COD1_COD2_COD3_cgc2ccc 0.00587172)
  (COD1_COD2_COD3_cgc2ccg 0.00974582)
  (COD1_COD2_COD3_cgc2cct 2.47365e-05)
  (COD1_COD2_COD3_cgc2cga 0.061002)
  (COD1_COD2_COD3_cgc2cgg 0.290176)
  (COD1_COD2_COD3_cgc2cgt 0.231487)
  (COD1_COD2_COD3_cgc2cta 3.98793e-05)
  (COD1_COD2_COD3_cgc2ctc 0.00915642)
  (COD1_COD2_COD3_cgc2ctg 0.0160754)
  (COD1_COD2_COD3_cgc2ctt 0.000243296)
  (COD1_COD2_COD3_cgc2gaa 0.000885635)
  (COD1_COD2_COD3_cgc2gac 0.0231635)
  (COD1_COD2_COD3_cgc2gag 0.0161872)
  (COD1_COD2_COD3_cgc2gat 5.218e-07)
  (COD1_COD2_COD3_cgc2gca 6.33128e-07)
  (COD1_COD2_COD3_cgc2gcc 0.0431948)
  (COD1_COD2_COD3_cgc2gcg 0.00153239)
  (COD1_COD2_COD3_cgc2gct 2.31417e-07)
  (COD1_COD2_COD3_cgc2gga 1.24802e-05)
  (COD1_COD2_COD3_cgc2ggc 0.0159976)
  (COD1_COD2_COD3_cgc2ggg 0.000993633)
  (COD1_COD2_COD3_cgc2ggt 7.87151e-05)
  (COD1_COD2_COD3_cgc2gta 7.10298e-08)
  (COD1_COD2_COD3_cgc2gtc 0.0119487)
  (COD1_COD2_COD3_cgc2gtg 7.40059e-05)
  (COD1_COD2_COD3_cgc2gtt 2.13983e-07)
  (COD1_COD2_COD3_cgc2taa 0)
  (COD1_COD2_COD3_cgc2tac 0.0100965)
  (COD1_COD2_COD3_cgc2tag 0)
  (COD1_COD2_COD3_cgc2tat 2.17131e-06)
  (COD1_COD2_COD3_cgc2tca 9.5495e-08)
  (COD1_COD2_COD3_cgc2tcc 0.00738084)
  (COD1_COD2_COD3_cgc2tcg 0.000279667)
  (COD1_COD2_COD3_cgc2tct 2.13095e-07)
  (COD1_COD2_COD3_cgc2tga 0)
  (COD1_COD2_COD3_cgc2tgc 0.00383382)
  (COD1_COD2_COD3_cgc2tgg 0.00224978)
  (COD1_COD2_COD3_cgc2tgt 8.34921e-05)
  (COD1_COD2_COD3_cgc2tta 1.357e-08)
  (COD1_COD2_COD3_cgc2ttc 0.00428943)
  (COD1_COD2_COD3_cgc2ttg 1.31911e-06)
  (COD1_COD2_COD3_cgc2ttt 2.45802e-07)
  (COD1_COD2_COD3_cgg2aaa 0.000344789)
  (COD1_COD2_COD3_cgg2aac 0.00132481)
  (COD1_COD2_COD3_cgg2aag 0.10238)
  (COD1_COD2_COD3_cgg2aat 2.14747e-05)
  (COD1_COD2_COD3_cgg2aca 3.52469e-07)
  (COD1_COD2_COD3_cgg2acc 0.00031705)
  (COD1_COD2_COD3_cgg2acg 0.0319066)
  (COD1_COD2_COD3_cgg2act 3.14159e-07)
  (COD1_COD2_COD3_cgg2aga 0.0235848)
  (COD1_COD2_COD3_cgg2agc 0.0186459)
  (COD1_COD2_COD3_cgg2agg 0.125869)
  (COD1_COD2_COD3_cgg2agt 0.000257704)
  (COD1_COD2_COD3_cgg2ata 2.17864e-08)
  (COD1_COD2_COD3_cgg2atc 7.34947e-05)
  (COD1_COD2_COD3_cgg2atg 0.0086565)
  (COD1_COD2_COD3_cgg2att 5.04025e-07)
  (COD1_COD2_COD3_cgg2caa 0.00515076)
  (COD1_COD2_COD3_cgg2cac 0.0243827)
  (COD1_COD2_COD3_cgg2cag 0.0978693)
  (COD1_COD2_COD3_cgg2cat 0.0033771)
  (COD1_COD2_COD3_cgg2cca 0.000114767)
  (COD1_COD2_COD3_cgg2ccc 0.00934234)
  (COD1_COD2_COD3_cgg2ccg 0.0142095)
  (COD1_COD2_COD3_cgg2cct 0.000103005)
  (COD1_COD2_COD3_cgg2cga 0.147113)
  (COD1_COD2_COD3_cgg2cgc 0.51943)
  (COD1_COD2_COD3_cgg2cgt 0.109272)
  (COD1_COD2_COD3_cgg2cta 0.000300989)
  (COD1_COD2_COD3_cgg2ctc 0.0107613)
  (COD1_COD2_COD3_cgg2ctg 0.0226397)
  (COD1_COD2_COD3_cgg2ctt 0.00059865)
  (COD1_COD2_COD3_cgg2gaa 2.98688e-05)
  (COD1_COD2_COD3_cgg2gac 0.00380998)
  (COD1_COD2_COD3_cgg2gag 0.0665231)
  (COD1_COD2_COD3_cgg2gat 1.08797e-05)
  (COD1_COD2_COD3_cgg2gca 1.32278e-06)
  (COD1_COD2_COD3_cgg2gcc 0.000550107)
  (COD1_COD2_COD3_cgg2gcg 0.0623702)
  (COD1_COD2_COD3_cgg2gct 1.02141e-06)
  (COD1_COD2_COD3_cgg2gga 0.000170848)
  (COD1_COD2_COD3_cgg2ggc 0.0166539)
  (COD1_COD2_COD3_cgg2ggg 0.00604099)
  (COD1_COD2_COD3_cgg2ggt 0.000199545)
  (COD1_COD2_COD3_cgg2gta 2.98245e-07)
  (COD1_COD2_COD3_cgg2gtc 9.24479e-05)
  (COD1_COD2_COD3_cgg2gtg 0.0158761)
  (COD1_COD2_COD3_cgg2gtt 8.92899e-07)
  (COD1_COD2_COD3_cgg2taa 0)
  (COD1_COD2_COD3_cgg2tac 0.00136868)
  (COD1_COD2_COD3_cgg2tag 0)
  (COD1_COD2_COD3_cgg2tat 6.45153e-05)
  (COD1_COD2_COD3_cgg2tca 3.48823e-07)
  (COD1_COD2_COD3_cgg2tcc 5.18247e-05)
  (COD1_COD2_COD3_cgg2tcg 0.0166077)
  (COD1_COD2_COD3_cgg2tct 7.59735e-07)
  (COD1_COD2_COD3_cgg2tga 0)
  (COD1_COD2_COD3_cgg2tgc 0.00135828)
  (COD1_COD2_COD3_cgg2tgg 0.0144779)
  (COD1_COD2_COD3_cgg2tgt 0.000154404)
  (COD1_COD2_COD3_cgg2tta 2.60345e-08)
  (COD1_COD2_COD3_cgg2ttc 0.000584301)
  (COD1_COD2_COD3_cgg2ttg 0.00200803)
  (COD1_COD2_COD3_cgg2ttt 1.96651e-05)
  (COD1_COD2_COD3_cgt2aaa 0.0550543)
  (COD1_COD2_COD3_cgt2aac 6.41585e-05)
  (COD1_COD2_COD3_cgt2aag 0.0149781)
  (COD1_COD2_COD3_cgt2aat 0.0151977)
  (COD1_COD2_COD3_cgt2aca 7.73054e-06)
  (COD1_COD2_COD3_cgt2acc 1.89558e-05)
  (COD1_COD2_COD3_cgt2acg 4.88595e-05)
  (COD1_COD2_COD3_cgt2act 0.00436138)
  (COD1_COD2_COD3_cgt2aga 0.170164)
  (COD1_COD2_COD3_cgt2agc 0.00531763)
  (COD1_COD2_COD3_cgt2agg 0.0461668)
  (COD1_COD2_COD3_cgt2agt 0.00796561)
  (COD1_COD2_COD3_cgt2ata 5.70607e-07)
  (COD1_COD2_COD3_cgt2atc 2.35637e-06)
  (COD1_COD2_COD3_cgt2atg 0.000608692)
  (COD1_COD2_COD3_cgt2att 0.00189314)
  (COD1_COD2_COD3_cgt2caa 0.019998)
  (COD1_COD2_COD3_cgt2cac 0.00306266)
  (COD1_COD2_COD3_cgt2cag 0.00706546)
  (COD1_COD2_COD3_cgt2cat 0.026278)
  (COD1_COD2_COD3_cgt2cca 0.000708473)
  (COD1_COD2_COD3_cgt2ccc 0.00129395)
  (COD1_COD2_COD3_cgt2ccg 0.000544795)
  (COD1_COD2_COD3_cgt2cct 0.00521904)
  (COD1_COD2_COD3_cgt2cga 0.190518)
  (COD1_COD2_COD3_cgt2cgc 0.398044)
  (COD1_COD2_COD3_cgt2cgg 0.105039)
  (COD1_COD2_COD3_cgt2cta 0.000989509)
  (COD1_COD2_COD3_cgt2ctc 0.00115491)
  (COD1_COD2_COD3_cgt2ctg 0.000428327)
  (COD1_COD2_COD3_cgt2ctt 0.0100755)
  (COD1_COD2_COD3_cgt2gaa 0.0012073)
  (COD1_COD2_COD3_cgt2gac 5.96106e-06)
  (COD1_COD2_COD3_cgt2gag 0.000242791)
  (COD1_COD2_COD3_cgt2gat 0.00464856)
  (COD1_COD2_COD3_cgt2gca 1.0095e-05)
  (COD1_COD2_COD3_cgt2gcc 6.71584e-06)
  (COD1_COD2_COD3_cgt2gcg 2.74849e-05)
  (COD1_COD2_COD3_cgt2gct 0.00613381)
  (COD1_COD2_COD3_cgt2gga 0.000251527)
  (COD1_COD2_COD3_cgt2ggc 0.000507255)
  (COD1_COD2_COD3_cgt2ggg 0.000441898)
  (COD1_COD2_COD3_cgt2ggt 0.00407213)
  (COD1_COD2_COD3_cgt2gta 4.4002e-06)
  (COD1_COD2_COD3_cgt2gtc 1.13496e-05)
  (COD1_COD2_COD3_cgt2gtg 3.27533e-05)
  (COD1_COD2_COD3_cgt2gtt 0.0064076)
  (COD1_COD2_COD3_cgt2taa 0)
  (COD1_COD2_COD3_cgt2tac 2.32083e-05)
  (COD1_COD2_COD3_cgt2tag 0)
  (COD1_COD2_COD3_cgt2tat 0.00921372)
  (COD1_COD2_COD3_cgt2tca 5.84648e-06)
  (COD1_COD2_COD3_cgt2tcc 1.31553e-05)
  (COD1_COD2_COD3_cgt2tcg 1.48931e-05)
  (COD1_COD2_COD3_cgt2tct 0.00779627)
  (COD1_COD2_COD3_cgt2tga 0)
  (COD1_COD2_COD3_cgt2tgc 0.000577056)
  (COD1_COD2_COD3_cgt2tgg 0.00025083)
  (COD1_COD2_COD3_cgt2tgt 0.00468414)
  (COD1_COD2_COD3_cgt2tta 1.19718e-05)
  (COD1_COD2_COD3_cgt2ttc 2.24322e-06)
  (COD1_COD2_COD3_cgt2ttg 4.79204e-05)
  (COD1_COD2_COD3_cgt2ttt 0.00308182)
  (COD1_COD2_COD3_cta2aaa 0.0134803)
  (COD1_COD2_COD3_cta2aac 0.000295819)
  (COD1_COD2_COD3_cta2aag 1.88641e-05)
  (COD1_COD2_COD3_cta2aat 0.000437327)
  (COD1_COD2_COD3_cta2aca 0.0123958)
  (COD1_COD2_COD3_cta2acc 1.16687e-05)
  (COD1_COD2_COD3_cta2acg 4.17729e-06)
  (COD1_COD2_COD3_cta2act 5.56214e-05)
  (COD1_COD2_COD3_cta2aga 0.00399109)
  (COD1_COD2_COD3_cta2agc 3.78591e-05)
  (COD1_COD2_COD3_cta2agg 6.80959e-06)
  (COD1_COD2_COD3_cta2agt 9.79225e-05)
  (COD1_COD2_COD3_cta2ata 0.0564063)
  (COD1_COD2_COD3_cta2atc 0.00482585)
  (COD1_COD2_COD3_cta2atg 0.03465)
  (COD1_COD2_COD3_cta2att 0.0329175)
  (COD1_COD2_COD3_cta2caa 0.0207032)
  (COD1_COD2_COD3_cta2cac 0.000661487)
  (COD1_COD2_COD3_cta2cag 0.00218674)
  (COD1_COD2_COD3_cta2cat 0.00468414)
  (COD1_COD2_COD3_cta2cca 0.0119902)
  (COD1_COD2_COD3_cta2ccc 0.000416903)
  (COD1_COD2_COD3_cta2ccg 0.000200933)
  (COD1_COD2_COD3_cta2cct 0.00189183)
  (COD1_COD2_COD3_cta2cga 0.00441614)
  (COD1_COD2_COD3_cta2cgc 7.9098e-05)
  (COD1_COD2_COD3_cta2cgg 0.000333737)
  (COD1_COD2_COD3_cta2cgt 0.00114218)
  (COD1_COD2_COD3_cta2ctc 0.148034)
  (COD1_COD2_COD3_cta2ctg 0.237171)
  (COD1_COD2_COD3_cta2ctt 0.325335)
  (COD1_COD2_COD3_cta2gaa 0.0108287)
  (COD1_COD2_COD3_cta2gac 3.36975e-05)
  (COD1_COD2_COD3_cta2gag 2.95789e-06)
  (COD1_COD2_COD3_cta2gat 0.000352278)
  (COD1_COD2_COD3_cta2gca 0.0122421)
  (COD1_COD2_COD3_cta2gcc 2.85515e-06)
  (COD1_COD2_COD3_cta2gcg 6.7904e-07)
  (COD1_COD2_COD3_cta2gct 7.71486e-05)
  (COD1_COD2_COD3_cta2gga 0.00271845)
  (COD1_COD2_COD3_cta2ggc 4.5583e-07)
  (COD1_COD2_COD3_cta2ggg 6.83289e-07)
  (COD1_COD2_COD3_cta2ggt 5.0545e-06)
  (COD1_COD2_COD3_cta2gta 0.022359)
  (COD1_COD2_COD3_cta2gtc 0.000805962)
  (COD1_COD2_COD3_cta2gtg 0.00391981)
  (COD1_COD2_COD3_cta2gtt 0.0105618)
  (COD1_COD2_COD3_cta2taa 0)
  (COD1_COD2_COD3_cta2tac 0.000199683)
  (COD1_COD2_COD3_cta2tag 0)
  (COD1_COD2_COD3_cta2tat 0.00028753)
  (COD1_COD2_COD3_cta2tca 0.00514362)
  (COD1_COD2_COD3_cta2tcc 2.20468e-06)
  (COD1_COD2_COD3_cta2tcg 7.02499e-07)
  (COD1_COD2_COD3_cta2tct 8.10925e-06)
  (COD1_COD2_COD3_cta2tga 0)
  (COD1_COD2_COD3_cta2tgc 0.000160404)
  (COD1_COD2_COD3_cta2tgg 0.000217756)
  (COD1_COD2_COD3_cta2tgt 0.000133303)
  (COD1_COD2_COD3_cta2tta 0.441046)
  (COD1_COD2_COD3_cta2ttc 0.00423918)
  (COD1_COD2_COD3_cta2ttg 0.260436)
  (COD1_COD2_COD3_cta2ttt 0.0222046)
  (COD1_COD2_COD3_ctc2aaa 6.69758e-05)
  (COD1_COD2_COD3_ctc2aac 0.00382056)
  (COD1_COD2_COD3_ctc2aag 0.000598235)
  (COD1_COD2_COD3_ctc2aat 2.69539e-07)
  (COD1_COD2_COD3_ctc2aca 1.57196e-06)
  (COD1_COD2_COD3_ctc2acc 0.0160531)
  (COD1_COD2_COD3_ctc2acg 6.07788e-06)
  (COD1_COD2_COD3_ctc2act 1.06701e-06)
  (COD1_COD2_COD3_ctc2aga 6.93308e-07)
  (COD1_COD2_COD3_ctc2agc 0.00411754)
  (COD1_COD2_COD3_ctc2agg 6.49608e-06)
  (COD1_COD2_COD3_ctc2agt 3.56647e-07)
  (COD1_COD2_COD3_ctc2ata 0.00174567)
  (COD1_COD2_COD3_ctc2atc 0.089498)
  (COD1_COD2_COD3_ctc2atg 0.0303955)
  (COD1_COD2_COD3_ctc2att 0.00455921)
  (COD1_COD2_COD3_ctc2caa 0.00123352)
  (COD1_COD2_COD3_ctc2cac 0.00682006)
  (COD1_COD2_COD3_ctc2cag 0.00876518)
  (COD1_COD2_COD3_ctc2cat 0.000612077)
  (COD1_COD2_COD3_ctc2cca 0.000185795)
  (COD1_COD2_COD3_ctc2ccc 0.0067542)
  (COD1_COD2_COD3_ctc2ccg 0.00144271)
  (COD1_COD2_COD3_ctc2cct 0.000378348)
  (COD1_COD2_COD3_ctc2cga 0.000540657)
  (COD1_COD2_COD3_ctc2cgc 0.00796009)
  (COD1_COD2_COD3_ctc2cgg 0.00522628)
  (COD1_COD2_COD3_ctc2cgt 0.000583896)
  (COD1_COD2_COD3_ctc2cta 0.0648387)
  (COD1_COD2_COD3_ctc2ctg 0.441351)
  (COD1_COD2_COD3_ctc2ctt 0.240815)
  (COD1_COD2_COD3_ctc2gaa 9.28975e-05)
  (COD1_COD2_COD3_ctc2gac 0.00342187)
  (COD1_COD2_COD3_ctc2gag 0.000308381)
  (COD1_COD2_COD3_ctc2gat 6.38859e-07)
  (COD1_COD2_COD3_ctc2gca 5.76999e-06)
  (COD1_COD2_COD3_ctc2gcc 0.031973)
  (COD1_COD2_COD3_ctc2gcg 1.6241e-05)
  (COD1_COD2_COD3_ctc2gct 8.29688e-06)
  (COD1_COD2_COD3_ctc2gga 6.02724e-07)
  (COD1_COD2_COD3_ctc2ggc 0.0056914)
  (COD1_COD2_COD3_ctc2ggg 1.29466e-06)
  (COD1_COD2_COD3_ctc2ggt 6.28319e-07)
  (COD1_COD2_COD3_ctc2gta 0.000346946)
  (COD1_COD2_COD3_ctc2gtc 0.0501363)
  (COD1_COD2_COD3_ctc2gtg 0.0108062)
  (COD1_COD2_COD3_ctc2gtt 0.00138682)
  (COD1_COD2_COD3_ctc2taa 0)
  (COD1_COD2_COD3_ctc2tac 0.0103876)
  (COD1_COD2_COD3_ctc2tag 0)
  (COD1_COD2_COD3_ctc2tat 1.90471e-06)
  (COD1_COD2_COD3_ctc2tca 8.56526e-07)
  (COD1_COD2_COD3_ctc2tcc 0.00369042)
  (COD1_COD2_COD3_ctc2tcg 2.89903e-06)
  (COD1_COD2_COD3_ctc2tct 1.29466e-06)
  (COD1_COD2_COD3_ctc2tga 0)
  (COD1_COD2_COD3_ctc2tgc 0.0049995)
  (COD1_COD2_COD3_ctc2tgg 0.00144973)
  (COD1_COD2_COD3_ctc2tgt 3.03473e-06)
  (COD1_COD2_COD3_ctc2tta 0.0222508)
  (COD1_COD2_COD3_ctc2ttc 0.0492409)
  (COD1_COD2_COD3_ctc2ttg 0.0888798)
  (COD1_COD2_COD3_ctc2ttt 0.00179227)
  (COD1_COD2_COD3_ctg2aaa 9.23746e-07)
  (COD1_COD2_COD3_ctg2aac 0.000709456)
  (COD1_COD2_COD3_ctg2aag 0.00355238)
  (COD1_COD2_COD3_ctg2aat 3.13957e-06)
  (COD1_COD2_COD3_ctg2aca 8.40062e-07)
  (COD1_COD2_COD3_ctg2acc 0.00131748)
  (COD1_COD2_COD3_ctg2acg 0.00783419)
  (COD1_COD2_COD3_ctg2act 9.59642e-07)
  (COD1_COD2_COD3_ctg2aga 2.04263e-08)
  (COD1_COD2_COD3_ctg2agc 0.0002373)
  (COD1_COD2_COD3_ctg2agg 0.000273912)
  (COD1_COD2_COD3_ctg2agt 9.12293e-07)
  (COD1_COD2_COD3_ctg2ata 0.000626673)
  (COD1_COD2_COD3_ctg2atc 0.0586795)
  (COD1_COD2_COD3_ctg2atg 0.047399)
  (COD1_COD2_COD3_ctg2att 0.00221266)
  (COD1_COD2_COD3_ctg2caa 0.00057307)
  (COD1_COD2_COD3_ctg2cac 0.00364466)
  (COD1_COD2_COD3_ctg2cag 0.0145989)
  (COD1_COD2_COD3_ctg2cat 0.00047757)
  (COD1_COD2_COD3_ctg2cca 0.000457164)
  (COD1_COD2_COD3_ctg2ccc 0.00155918)
  (COD1_COD2_COD3_ctg2ccg 0.00604099)
  (COD1_COD2_COD3_ctg2cct 0.000150078)
  (COD1_COD2_COD3_ctg2cga 0.00020191)
  (COD1_COD2_COD3_ctg2cgc 0.0082408)
  (COD1_COD2_COD3_ctg2cgg 0.00648355)
  (COD1_COD2_COD3_ctg2cgt 0.000127607)
  (COD1_COD2_COD3_ctg2cta 0.0612562)
  (COD1_COD2_COD3_ctg2ctc 0.260255)
  (COD1_COD2_COD3_ctg2ctt 0.0728463)
  (COD1_COD2_COD3_ctg2gaa 5.15713e-06)
  (COD1_COD2_COD3_ctg2gac 0.000397158)
  (COD1_COD2_COD3_ctg2gag 0.0053956)
  (COD1_COD2_COD3_ctg2gat 1.43062e-05)
  (COD1_COD2_COD3_ctg2gca 5.69845e-06)
  (COD1_COD2_COD3_ctg2gcc 0.00278328)
  (COD1_COD2_COD3_ctg2gcg 0.0278728)
  (COD1_COD2_COD3_ctg2gct 1.48415e-05)
  (COD1_COD2_COD3_ctg2gga 8.66678e-07)
  (COD1_COD2_COD3_ctg2ggc 0.00111867)
  (COD1_COD2_COD3_ctg2ggg 0.00189183)
  (COD1_COD2_COD3_ctg2ggt 2.67506e-06)
  (COD1_COD2_COD3_ctg2gta 0.000224344)
  (COD1_COD2_COD3_ctg2gtc 0.019998)
  (COD1_COD2_COD3_ctg2gtg 0.0512253)
  (COD1_COD2_COD3_ctg2gtt 0.000220184)
  (COD1_COD2_COD3_ctg2taa 0)
  (COD1_COD2_COD3_ctg2tac 0.0051579)
  (COD1_COD2_COD3_ctg2tag 0)
  (COD1_COD2_COD3_ctg2tat 8.29154e-05)
  (COD1_COD2_COD3_ctg2tca 2.56774e-07)
  (COD1_COD2_COD3_ctg2tcc 1.08346e-05)
  (COD1_COD2_COD3_ctg2tcg 0.00232912)
  (COD1_COD2_COD3_ctg2tct 1.87449e-07)
  (COD1_COD2_COD3_ctg2tga 0)
  (COD1_COD2_COD3_ctg2tgc 0.00197217)
  (COD1_COD2_COD3_ctg2tgg 0.0064343)
  (COD1_COD2_COD3_ctg2tgt 1.67208e-05)
  (COD1_COD2_COD3_ctg2tta 0.016945)
  (COD1_COD2_COD3_ctg2ttc 0.0286365)
  (COD1_COD2_COD3_ctg2ttg 0.195874)
  (COD1_COD2_COD3_ctg2ttt 0.00157547)
  (COD1_COD2_COD3_ctt2aaa 0.00189577)
  (COD1_COD2_COD3_ctt2aac 6.81904e-06)
  (COD1_COD2_COD3_ctt2aag 0.00105174)
  (COD1_COD2_COD3_ctt2aat 0.00577886)
  (COD1_COD2_COD3_ctt2aca 1.13182e-05)
  (COD1_COD2_COD3_ctt2acc 7.603e-06)
  (COD1_COD2_COD3_ctt2acg 3.12654e-06)
  (COD1_COD2_COD3_ctt2act 0.0105253)
  (COD1_COD2_COD3_ctt2aga 4.61599e-05)
  (COD1_COD2_COD3_ctt2agc 4.83853e-06)
  (COD1_COD2_COD3_ctt2agg 6.4874e-05)
  (COD1_COD2_COD3_ctt2agt 0.0033283)
  (COD1_COD2_COD3_ctt2ata 0.0161536)
  (COD1_COD2_COD3_ctt2atc 0.00927781)
  (COD1_COD2_COD3_ctt2atg 0.0258445)
  (COD1_COD2_COD3_ctt2att 0.0824692)
  (COD1_COD2_COD3_ctt2caa 0.00908688)
  (COD1_COD2_COD3_ctt2cac 0.000647425)
  (COD1_COD2_COD3_ctt2cag 0.0016401)
  (COD1_COD2_COD3_ctt2cat 0.00709491)
  (COD1_COD2_COD3_ctt2cca 0.000733457)
  (COD1_COD2_COD3_ctt2ccc 0.000276775)
  (COD1_COD2_COD3_ctt2ccg 9.71787e-05)
  (COD1_COD2_COD3_ctt2cct 0.00857885)
  (COD1_COD2_COD3_ctt2cga 0.00204313)
  (COD1_COD2_COD3_ctt2cgc 0.000234195)
  (COD1_COD2_COD3_ctt2cgg 0.000321922)
  (COD1_COD2_COD3_ctt2cgt 0.00564035)
  (COD1_COD2_COD3_ctt2cta 0.157782)
  (COD1_COD2_COD3_ctt2ctc 0.266831)
  (COD1_COD2_COD3_ctt2ctg 0.136882)
  (COD1_COD2_COD3_ctt2gaa 0.00136489)
  (COD1_COD2_COD3_ctt2gac 9.58313e-07)
  (COD1_COD2_COD3_ctt2gag 0.000128673)
  (COD1_COD2_COD3_ctt2gat 0.00370323)
  (COD1_COD2_COD3_ctt2gca 3.86814e-05)
  (COD1_COD2_COD3_ctt2gcc 3.83857e-06)
  (COD1_COD2_COD3_ctt2gcg 1.63872e-06)
  (COD1_COD2_COD3_ctt2gct 0.0166423)
  (COD1_COD2_COD3_ctt2gga 5.85459e-06)
  (COD1_COD2_COD3_ctt2ggc 3.73342e-07)
  (COD1_COD2_COD3_ctt2ggg 8.41228e-07)
  (COD1_COD2_COD3_ctt2ggt 0.00286748)
  (COD1_COD2_COD3_ctt2gta 0.00556655)
  (COD1_COD2_COD3_ctt2gtc 0.00194098)
  (COD1_COD2_COD3_ctt2gtg 0.00207885)
  (COD1_COD2_COD3_ctt2gtt 0.0360964)
  (COD1_COD2_COD3_ctt2taa 0)
  (COD1_COD2_COD3_ctt2tac 5.23275e-06)
  (COD1_COD2_COD3_ctt2tag 0)
  (COD1_COD2_COD3_ctt2tat 0.00837323)
  (COD1_COD2_COD3_ctt2tca 3.2911e-06)
  (COD1_COD2_COD3_ctt2tcc 1.92862e-06)
  (COD1_COD2_COD3_ctt2tcg 4.83494e-07)
  (COD1_COD2_COD3_ctt2tct 0.0032802)
  (COD1_COD2_COD3_ctt2tga 0)
  (COD1_COD2_COD3_ctt2tgc 4.53332e-06)
  (COD1_COD2_COD3_ctt2tgg 0.000128852)
  (COD1_COD2_COD3_ctt2tgt 0.00350104)
  (COD1_COD2_COD3_ctt2tta 0.194926)
  (COD1_COD2_COD3_ctt2ttc 0.00443454)
  (COD1_COD2_COD3_ctt2ttg 0.175434)
  (COD1_COD2_COD3_ctt2ttt 0.0431948)
  (COD1_COD2_COD3_gaa2aaa 0.0702187)
  (COD1_COD2_COD3_gaa2aac 0.00393342)
  (COD1_COD2_COD3_gaa2aag 0.00824651)
  (COD1_COD2_COD3_gaa2aat 0.0217325)
  (COD1_COD2_COD3_gaa2aca 0.0152928)
  (COD1_COD2_COD3_gaa2acc 0.00196262)
  (COD1_COD2_COD3_gaa2acg 3.70286e-05)
  (COD1_COD2_COD3_gaa2act 0.00318388)
  (COD1_COD2_COD3_gaa2aga 0.00651509)
  (COD1_COD2_COD3_gaa2agc 0.00422744)
  (COD1_COD2_COD3_gaa2agg 3.00141e-05)
  (COD1_COD2_COD3_gaa2agt 0.0085848)
  (COD1_COD2_COD3_gaa2ata 0.00463248)
  (COD1_COD2_COD3_gaa2atc 0.000217605)
  (COD1_COD2_COD3_gaa2atg 0.000719858)
  (COD1_COD2_COD3_gaa2att 0.000956465)
  (COD1_COD2_COD3_gaa2caa 0.0529216)
  (COD1_COD2_COD3_gaa2cac 0.00117185)
  (COD1_COD2_COD3_gaa2cag 0.0111254)
  (COD1_COD2_COD3_gaa2cat 0.0048292)
  (COD1_COD2_COD3_gaa2cca 0.00995061)
  (COD1_COD2_COD3_gaa2ccc 0.00109413)
  (COD1_COD2_COD3_gaa2ccg 6.62831e-05)
  (COD1_COD2_COD3_gaa2cct 0.00192624)
  (COD1_COD2_COD3_gaa2cga 0.00211812)
  (COD1_COD2_COD3_gaa2cgc 0.000374694)
  (COD1_COD2_COD3_gaa2cgg 7.05462e-06)
  (COD1_COD2_COD3_gaa2cgt 0.000296846)
  (COD1_COD2_COD3_gaa2cta 0.00230663)
  (COD1_COD2_COD3_gaa2ctc 4.521e-05)
  (COD1_COD2_COD3_gaa2ctg 4.25621e-06)
  (COD1_COD2_COD3_gaa2ctt 0.000599896)
  (COD1_COD2_COD3_gaa2gac 0.022577)
  (COD1_COD2_COD3_gaa2gag 0.397493)
  (COD1_COD2_COD3_gaa2gat 0.0783458)
  (COD1_COD2_COD3_gaa2gca 0.0149574)
  (COD1_COD2_COD3_gaa2gcc 0.00377057)
  (COD1_COD2_COD3_gaa2gcg 0.00416346)
  (COD1_COD2_COD3_gaa2gct 0.00949906)
  (COD1_COD2_COD3_gaa2gga 0.0104743)
  (COD1_COD2_COD3_gaa2ggc 0.000723861)
  (COD1_COD2_COD3_gaa2ggg 0.00140035)
  (COD1_COD2_COD3_gaa2ggt 0.00150502)
  (COD1_COD2_COD3_gaa2gta 0.00512583)
  (COD1_COD2_COD3_gaa2gtc 0.000783923)
  (COD1_COD2_COD3_gaa2gtg 0.00124469)
  (COD1_COD2_COD3_gaa2gtt 0.00526263)
  (COD1_COD2_COD3_gaa2taa 0)
  (COD1_COD2_COD3_gaa2tac 0.000730413)
  (COD1_COD2_COD3_gaa2tag 0)
  (COD1_COD2_COD3_gaa2tat 0.00443454)
  (COD1_COD2_COD3_gaa2tca 0.0131208)
  (COD1_COD2_COD3_gaa2tcc 0.000921965)
  (COD1_COD2_COD3_gaa2tcg 1.89164e-05)
  (COD1_COD2_COD3_gaa2tct 0.00390084)
  (COD1_COD2_COD3_gaa2tga 0)
  (COD1_COD2_COD3_gaa2tgc 2.74088e-05)
  (COD1_COD2_COD3_gaa2tgg 0.000127254)
  (COD1_COD2_COD3_gaa2tgt 0.000186311)
  (COD1_COD2_COD3_gaa2tta 0.00465501)
  (COD1_COD2_COD3_gaa2ttc 0.000137718)
  (COD1_COD2_COD3_gaa2ttg 8.61374e-05)
  (COD1_COD2_COD3_gaa2ttt 0.00181981)
  (COD1_COD2_COD3_gac2aaa 0.0016538)
  (COD1_COD2_COD3_gac2aac 0.0554757)
  (COD1_COD2_COD3_gac2aag 0.0113986)
  (COD1_COD2_COD3_gac2aat 0.00372124)
  (COD1_COD2_COD3_gac2aca 4.40347e-05)
  (COD1_COD2_COD3_gac2acc 0.0181234)
  (COD1_COD2_COD3_gac2acg 0.00101733)
  (COD1_COD2_COD3_gac2act 2.38761e-06)
  (COD1_COD2_COD3_gac2aga 7.01075e-06)
  (COD1_COD2_COD3_gac2agc 0.027489)
  (COD1_COD2_COD3_gac2agg 6.11198e-05)
  (COD1_COD2_COD3_gac2agt 4.43695e-06)
  (COD1_COD2_COD3_gac2ata 4.51406e-08)
  (COD1_COD2_COD3_gac2atc 0.000771525)
  (COD1_COD2_COD3_gac2atg 0.000241615)
  (COD1_COD2_COD3_gac2att 4.68622e-08)
  (COD1_COD2_COD3_gac2caa 0.00208462)
  (COD1_COD2_COD3_gac2cac 0.0107763)
  (COD1_COD2_COD3_gac2cag 0.01925)
  (COD1_COD2_COD3_gac2cat 0.00107683)
  (COD1_COD2_COD3_gac2cca 4.14577e-05)
  (COD1_COD2_COD3_gac2ccc 0.0101667)
  (COD1_COD2_COD3_gac2ccg 0.00786139)
  (COD1_COD2_COD3_gac2cct 1.24623e-06)
  (COD1_COD2_COD3_gac2cga 4.82872e-05)
  (COD1_COD2_COD3_gac2cgc 0.0133779)
  (COD1_COD2_COD3_gac2cgg 0.0012284)
  (COD1_COD2_COD3_gac2cgt 2.00218e-06)
  (COD1_COD2_COD3_gac2cta 9.81215e-06)
  (COD1_COD2_COD3_gac2ctc 0.00227329)
  (COD1_COD2_COD3_gac2ctg 0.000447446)
  (COD1_COD2_COD3_gac2ctt 5.74974e-07)
  (COD1_COD2_COD3_gac2gaa 0.0308411)
  (COD1_COD2_COD3_gac2gag 0.0978693)
  (COD1_COD2_COD3_gac2gat 0.406408)
  (COD1_COD2_COD3_gac2gca 0.000460983)
  (COD1_COD2_COD3_gac2gcc 0.0215227)
  (COD1_COD2_COD3_gac2gcg 0.0128508)
  (COD1_COD2_COD3_gac2gct 0.000352278)
  (COD1_COD2_COD3_gac2gga 0.00110557)
  (COD1_COD2_COD3_gac2ggc 0.0290564)
  (COD1_COD2_COD3_gac2ggg 0.00412612)
  (COD1_COD2_COD3_gac2ggt 0.000592869)
  (COD1_COD2_COD3_gac2gta 7.3801e-05)
  (COD1_COD2_COD3_gac2gtc 0.00219282)
  (COD1_COD2_COD3_gac2gtg 0.000934835)
  (COD1_COD2_COD3_gac2gtt 7.83884e-05)
  (COD1_COD2_COD3_gac2taa 0)
  (COD1_COD2_COD3_gac2tac 0.0030861)
  (COD1_COD2_COD3_gac2tag 0)
  (COD1_COD2_COD3_gac2tat 0.000172873)
  (COD1_COD2_COD3_gac2tca 7.68284e-05)
  (COD1_COD2_COD3_gac2tcc 0.011018)
  (COD1_COD2_COD3_gac2tcg 0.00584736)
  (COD1_COD2_COD3_gac2tct 2.51328e-06)
  (COD1_COD2_COD3_gac2tga 0)
  (COD1_COD2_COD3_gac2tgc 0.000531368)
  (COD1_COD2_COD3_gac2tgg 0.000846032)
  (COD1_COD2_COD3_gac2tgt 4.43343e-08)
  (COD1_COD2_COD3_gac2tta 7.67144e-07)
  (COD1_COD2_COD3_gac2ttc 0.00118245)
  (COD1_COD2_COD3_gac2ttg 0.000106637)
  (COD1_COD2_COD3_gac2ttt 2.20151e-07)
  (COD1_COD2_COD3_gag2aaa 0.00495465)
  (COD1_COD2_COD3_gag2aac 0.00757783)
  (COD1_COD2_COD3_gag2aag 0.0417813)
  (COD1_COD2_COD3_gag2aat 0.00333291)
  (COD1_COD2_COD3_gag2aca 1.10316e-05)
  (COD1_COD2_COD3_gag2acc 0.00644323)
  (COD1_COD2_COD3_gag2acg 0.0154634)
  (COD1_COD2_COD3_gag2act 0.000425368)
  (COD1_COD2_COD3_gag2aga 4.40325e-06)
  (COD1_COD2_COD3_gag2agc 0.0112729)
  (COD1_COD2_COD3_gag2agg 0.00812171)
  (COD1_COD2_COD3_gag2agt 0.00133218)
  (COD1_COD2_COD3_gag2ata 1.41281e-06)
  (COD1_COD2_COD3_gag2atc 0.000760376)
  (COD1_COD2_COD3_gag2atg 0.0055627)
  (COD1_COD2_COD3_gag2att 8.20578e-05)
  (COD1_COD2_COD3_gag2caa 0.0058962)
  (COD1_COD2_COD3_gag2cac 0.00624969)
  (COD1_COD2_COD3_gag2cag 0.0629784)
  (COD1_COD2_COD3_gag2cat 0.00169912)
  (COD1_COD2_COD3_gag2cca 2.03586e-05)
  (COD1_COD2_COD3_gag2ccc 0.00306478)
  (COD1_COD2_COD3_gag2ccg 0.012082)
  (COD1_COD2_COD3_gag2cct 0.000359181)
  (COD1_COD2_COD3_gag2cga 4.92311e-06)
  (COD1_COD2_COD3_gag2cgc 0.00772634)
  (COD1_COD2_COD3_gag2cgg 0.0177259)
  (COD1_COD2_COD3_gag2cgt 6.73483e-05)
  (COD1_COD2_COD3_gag2cta 7.11319e-07)
  (COD1_COD2_COD3_gag2ctc 0.000169198)
  (COD1_COD2_COD3_gag2ctg 0.00502382)
  (COD1_COD2_COD3_gag2ctt 6.37595e-05)
  (COD1_COD2_COD3_gag2gaa 0.448444)
  (COD1_COD2_COD3_gag2gac 0.0808281)
  (COD1_COD2_COD3_gag2gat 0.0312067)
  (COD1_COD2_COD3_gag2gca 0.0026041)
  (COD1_COD2_COD3_gag2gcc 0.0251726)
  (COD1_COD2_COD3_gag2gcg 0.0228922)
  (COD1_COD2_COD3_gag2gct 0.00200943)
  (COD1_COD2_COD3_gag2gga 0.000880738)
  (COD1_COD2_COD3_gag2ggc 0.00567171)
  (COD1_COD2_COD3_gag2ggg 0.00990244)
  (COD1_COD2_COD3_gag2ggt 0.000510784)
  (COD1_COD2_COD3_gag2gta 0.000579461)
  (COD1_COD2_COD3_gag2gtc 0.00451518)
  (COD1_COD2_COD3_gag2gtg 0.00820091)
  (COD1_COD2_COD3_gag2gtt 0.000947885)
  (COD1_COD2_COD3_gag2taa 0)
  (COD1_COD2_COD3_gag2tac 0.00227803)
  (COD1_COD2_COD3_gag2tag 0)
  (COD1_COD2_COD3_gag2tat 0.000738047)
  (COD1_COD2_COD3_gag2tca 1.0678e-05)
  (COD1_COD2_COD3_gag2tcc 0.00304995)
  (COD1_COD2_COD3_gag2tcg 0.00898043)
  (COD1_COD2_COD3_gag2tct 0.000499925)
  (COD1_COD2_COD3_gag2tga 0)
  (COD1_COD2_COD3_gag2tgc 9.76514e-05)
  (COD1_COD2_COD3_gag2tgg 0.00175416)
  (COD1_COD2_COD3_gag2tgt 9.28975e-05)
  (COD1_COD2_COD3_gag2tta 1.36556e-07)
  (COD1_COD2_COD3_gag2ttc 0.000734984)
  (COD1_COD2_COD3_gag2ttg 0.00251365)
  (COD1_COD2_COD3_gag2ttt 0.000234683)
  (COD1_COD2_COD3_gat2aaa 0.0188146)
  (COD1_COD2_COD3_gat2aac 0.00755685)
  (COD1_COD2_COD3_gat2aag 0.00502382)
  (COD1_COD2_COD3_gat2aat 0.0786723)
  (COD1_COD2_COD3_gat2aca 0.00121992)
  (COD1_COD2_COD3_gat2acc 1.03644e-05)
  (COD1_COD2_COD3_gat2acg 0.000455267)
  (COD1_COD2_COD3_gat2act 0.0117597)
  (COD1_COD2_COD3_gat2aga 0.00029664)
  (COD1_COD2_COD3_gat2agc 0.000171441)
  (COD1_COD2_COD3_gat2agg 0.000106859)
  (COD1_COD2_COD3_gat2agt 0.0175669)
  (COD1_COD2_COD3_gat2ata 1.4688e-05)
  (COD1_COD2_COD3_gat2atc 1.01859e-06)
  (COD1_COD2_COD3_gat2atg 0.00125074)
  (COD1_COD2_COD3_gat2att 0.00165839)
  (COD1_COD2_COD3_gat2caa 0.0121407)
  (COD1_COD2_COD3_gat2cac 0.00051648)
  (COD1_COD2_COD3_gat2cag 0.00271468)
  (COD1_COD2_COD3_gat2cat 0.00797113)
  (COD1_COD2_COD3_gat2cca 0.00165151)
  (COD1_COD2_COD3_gat2ccc 7.91489e-06)
  (COD1_COD2_COD3_gat2ccg 0.000384694)
  (COD1_COD2_COD3_gat2cct 0.0069009)
  (COD1_COD2_COD3_gat2cga 3.39554e-05)
  (COD1_COD2_COD3_gat2cgc 2.66567e-07)
  (COD1_COD2_COD3_gat2cgg 3.1028e-06)
  (COD1_COD2_COD3_gat2cgt 0.00138011)
  (COD1_COD2_COD3_gat2cta 9.0671e-05)
  (COD1_COD2_COD3_gat2ctc 3.75418e-07)
  (COD1_COD2_COD3_gat2ctg 1.42567e-05)
  (COD1_COD2_COD3_gat2ctt 0.00196399)
  (COD1_COD2_COD3_gat2gaa 0.094601)
  (COD1_COD2_COD3_gat2gac 0.359484)
  (COD1_COD2_COD3_gat2gag 0.0334002)
  (COD1_COD2_COD3_gat2gca 0.00312484)
  (COD1_COD2_COD3_gat2gcc 0.000406629)
  (COD1_COD2_COD3_gat2gcg 0.000789923)
  (COD1_COD2_COD3_gat2gct 0.00749945)
  (COD1_COD2_COD3_gat2gga 0.00614658)
  (COD1_COD2_COD3_gat2ggc 0.000886249)
  (COD1_COD2_COD3_gat2ggg 0.00241126)
  (COD1_COD2_COD3_gat2ggt 0.0106279)
  (COD1_COD2_COD3_gat2gta 0.000856654)
  (COD1_COD2_COD3_gat2gtc 0.000131468)
  (COD1_COD2_COD3_gat2gtg 0.000259676)
  (COD1_COD2_COD3_gat2gtt 0.00291963)
  (COD1_COD2_COD3_gat2taa 0)
  (COD1_COD2_COD3_gat2tac 0.000258598)
  (COD1_COD2_COD3_gat2tag 0)
  (COD1_COD2_COD3_gat2tat 0.00385782)
  (COD1_COD2_COD3_gat2tca 0.00361447)
  (COD1_COD2_COD3_gat2tcc 3.999e-05)
  (COD1_COD2_COD3_gat2tcg 0.00102582)
  (COD1_COD2_COD3_gat2tct 0.0132212)
  (COD1_COD2_COD3_gat2tga 0)
  (COD1_COD2_COD3_gat2tgc 9.71017e-07)
  (COD1_COD2_COD3_gat2tgg 0.000360428)
  (COD1_COD2_COD3_gat2tgt 0.000644291)
  (COD1_COD2_COD3_gat2tta 0.000682918)
  (COD1_COD2_COD3_gat2ttc 4.5583e-07)
  (COD1_COD2_COD3_gat2ttg 0.000606166)
  (COD1_COD2_COD3_gat2ttt 0.00137915)
  (COD1_COD2_COD3_gca2aaa 0.0287359)
  (COD1_COD2_COD3_gca2aac 0.000842521)
  (COD1_COD2_COD3_gca2aag 0.000116693)
  (COD1_COD2_COD3_gca2aat 0.00188529)
  (COD1_COD2_COD3_gca2aca 0.0358968)
  (COD1_COD2_COD3_gca2acc 0.00138298)
  (COD1_COD2_COD3_gca2acg 0.00340295)
  (COD1_COD2_COD3_gca2act 0.00483255)
  (COD1_COD2_COD3_gca2aga 0.00450268)
  (COD1_COD2_COD3_gca2agc 0.00086801)
  (COD1_COD2_COD3_gca2agg 3.03699e-05)
  (COD1_COD2_COD3_gca2agt 0.00130295)
  (COD1_COD2_COD3_gca2ata 0.0126125)
  (COD1_COD2_COD3_gca2atc 1.40799e-05)
  (COD1_COD2_COD3_gca2atg 0.0109722)
  (COD1_COD2_COD3_gca2att 5.64761e-05)
  (COD1_COD2_COD3_gca2caa 0.0218837)
  (COD1_COD2_COD3_gca2cac 5.4628e-05)
  (COD1_COD2_COD3_gca2cag 4.57141e-05)
  (COD1_COD2_COD3_gca2cat 0.000418641)
  (COD1_COD2_COD3_gca2cca 0.0138114)
  (COD1_COD2_COD3_gca2ccc 0.000726374)
  (COD1_COD2_COD3_gca2ccg 0.00106643)
  (COD1_COD2_COD3_gca2cct 0.00350347)
  (COD1_COD2_COD3_gca2cga 0.0013782)
  (COD1_COD2_COD3_gca2cgc 5.59639e-07)
  (COD1_COD2_COD3_gca2cgg 6.53187e-07)
  (COD1_COD2_COD3_gca2cgt 5.1894e-06)
  (COD1_COD2_COD3_gca2cta 0.00545578)
  (COD1_COD2_COD3_gca2ctc 5.86678e-06)
  (COD1_COD2_COD3_gca2ctg 9.83257e-06)
  (COD1_COD2_COD3_gca2ctt 3.55202e-05)
  (COD1_COD2_COD3_gca2gaa 0.0312717)
  (COD1_COD2_COD3_gca2gac 0.000705532)
  (COD1_COD2_COD3_gca2gag 0.00482585)
  (COD1_COD2_COD3_gca2gat 0.00540683)
  (COD1_COD2_COD3_gca2gcc 0.180992)
  (COD1_COD2_COD3_gca2gcg 0.21154)
  (COD1_COD2_COD3_gca2gct 0.461371)
  (COD1_COD2_COD3_gca2gga 0.0277763)
  (COD1_COD2_COD3_gca2ggc 0.000661945)
  (COD1_COD2_COD3_gca2ggg 0.00687226)
  (COD1_COD2_COD3_gca2ggt 0.00988187)
  (COD1_COD2_COD3_gca2gta 0.0373176)
  (COD1_COD2_COD3_gca2gtc 0.00170265)
  (COD1_COD2_COD3_gca2gtg 0.00940732)
  (COD1_COD2_COD3_gca2gtt 0.013678)
  (COD1_COD2_COD3_gca2taa 0)
  (COD1_COD2_COD3_gca2tac 0.000580265)
  (COD1_COD2_COD3_gca2tag 0)
  (COD1_COD2_COD3_gca2tat 0.00304995)
  (COD1_COD2_COD3_gca2tca 0.0454366)
  (COD1_COD2_COD3_gca2tcc 0.00601175)
  (COD1_COD2_COD3_gca2tcg 0.00441002)
  (COD1_COD2_COD3_gca2tct 0.0227972)
  (COD1_COD2_COD3_gca2tga 0)
  (COD1_COD2_COD3_gca2tgc 0.00220654)
  (COD1_COD2_COD3_gca2tgg 0.000401031)
  (COD1_COD2_COD3_gca2tgt 0.00519738)
  (COD1_COD2_COD3_gca2tta 0.0168746)
  (COD1_COD2_COD3_gca2ttc 0.000352766)
  (COD1_COD2_COD3_gca2ttg 0.000304557)
  (COD1_COD2_COD3_gca2ttt 0.00235673)
  (COD1_COD2_COD3_gcc2aaa 0.000170848)
  (COD1_COD2_COD3_gcc2aac 0.0115018)
  (COD1_COD2_COD3_gcc2aag 0.00998515)
  (COD1_COD2_COD3_gcc2aat 1.18392e-06)
  (COD1_COD2_COD3_gcc2aca 0.000257882)
  (COD1_COD2_COD3_gcc2acc 0.0386336)
  (COD1_COD2_COD3_gcc2acg 0.0053287)
  (COD1_COD2_COD3_gcc2act 0.000487943)
  (COD1_COD2_COD3_gcc2aga 3.46894e-07)
  (COD1_COD2_COD3_gcc2agc 0.0303534)
  (COD1_COD2_COD3_gcc2agg 1.24026e-05)
  (COD1_COD2_COD3_gcc2agt 4.75541e-06)
  (COD1_COD2_COD3_gcc2ata 1.65347e-07)
  (COD1_COD2_COD3_gcc2atc 0.0127532)
  (COD1_COD2_COD3_gcc2atg 0.00358702)
  (COD1_COD2_COD3_gcc2att 5.02282e-07)
  (COD1_COD2_COD3_gcc2caa 0.000323264)
  (COD1_COD2_COD3_gcc2cac 0.00768894)
  (COD1_COD2_COD3_gcc2cag 0.0131664)
  (COD1_COD2_COD3_gcc2cat 5.85054e-06)
  (COD1_COD2_COD3_gcc2cca 0.000146481)
  (COD1_COD2_COD3_gcc2ccc 0.0145282)
  (COD1_COD2_COD3_gcc2ccg 0.0118497)
  (COD1_COD2_COD3_gcc2cct 0.000230491)
  (COD1_COD2_COD3_gcc2cga 2.17885e-06)
  (COD1_COD2_COD3_gcc2cgc 0.0235195)
  (COD1_COD2_COD3_gcc2cgg 0.000167332)
  (COD1_COD2_COD3_gcc2cgt 2.12663e-06)
  (COD1_COD2_COD3_gcc2cta 7.83806e-07)
  (COD1_COD2_COD3_gcc2ctc 0.0200257)
  (COD1_COD2_COD3_gcc2ctg 0.00295833)
  (COD1_COD2_COD3_gcc2ctt 2.17131e-06)
  (COD1_COD2_COD3_gcc2gaa 0.00485269)
  (COD1_COD2_COD3_gcc2gac 0.0203053)
  (COD1_COD2_COD3_gcc2gag 0.0287359)
  (COD1_COD2_COD3_gcc2gat 0.000433404)
  (COD1_COD2_COD3_gcc2gca 0.111491)
  (COD1_COD2_COD3_gcc2gcg 0.354535)
  (COD1_COD2_COD3_gcc2gct 0.181873)
  (COD1_COD2_COD3_gcc2gga 0.000589999)
  (COD1_COD2_COD3_gcc2ggc 0.0425411)
  (COD1_COD2_COD3_gcc2ggg 0.00507632)
  (COD1_COD2_COD3_gcc2ggt 0.00146793)
  (COD1_COD2_COD3_gcc2gta 0.000312036)
  (COD1_COD2_COD3_gcc2gtc 0.0482944)
  (COD1_COD2_COD3_gcc2gtg 0.0154527)
  (COD1_COD2_COD3_gcc2gtt 0.000886864)
  (COD1_COD2_COD3_gcc2taa 0)
  (COD1_COD2_COD3_gcc2tac 0.00635893)
  (COD1_COD2_COD3_gcc2tag 0)
  (COD1_COD2_COD3_gcc2tat 1.21974e-06)
  (COD1_COD2_COD3_gcc2tca 0.000873442)
  (COD1_COD2_COD3_gcc2tcc 0.0371627)
  (COD1_COD2_COD3_gcc2tcg 0.0172651)
  (COD1_COD2_COD3_gcc2tct 0.00132114)
  (COD1_COD2_COD3_gcc2tga 0)
  (COD1_COD2_COD3_gcc2tgc 0.0112339)
  (COD1_COD2_COD3_gcc2tgg 0.000563225)
  (COD1_COD2_COD3_gcc2tgt 3.15266e-06)
  (COD1_COD2_COD3_gcc2tta 2.64909e-07)
  (COD1_COD2_COD3_gcc2ttc 0.0109418)
  (COD1_COD2_COD3_gcc2ttg 2.6165e-05)
  (COD1_COD2_COD3_gcc2ttt 1.65929e-06)
  (COD1_COD2_COD3_gcg2aaa 1.47288e-05)
  (COD1_COD2_COD3_gcg2aac 0.0013926)
  (COD1_COD2_COD3_gcg2aag 0.0202772)
  (COD1_COD2_COD3_gcg2aat 8.38401e-05)
  (COD1_COD2_COD3_gcg2aca 0.000653738)
  (COD1_COD2_COD3_gcg2acc 0.0130392)
  (COD1_COD2_COD3_gcg2acg 0.0306281)
  (COD1_COD2_COD3_gcg2act 0.000302244)
  (COD1_COD2_COD3_gcg2aga 8.70247e-08)
  (COD1_COD2_COD3_gcg2agc 0.00274496)
  (COD1_COD2_COD3_gcg2agg 0.00127525)
  (COD1_COD2_COD3_gcg2agt 6.56885e-05)
  (COD1_COD2_COD3_gcg2ata 1.00805e-06)
  (COD1_COD2_COD3_gcg2atc 0.000109711)
  (COD1_COD2_COD3_gcg2atg 0.0177382)
  (COD1_COD2_COD3_gcg2att 2.30628e-06)
  (COD1_COD2_COD3_gcg2caa 2.0486e-05)
  (COD1_COD2_COD3_gcg2cac 0.00209912)
  (COD1_COD2_COD3_gcg2cag 0.0320396)
  (COD1_COD2_COD3_gcg2cat 0.000855467)
  (COD1_COD2_COD3_gcg2cca 0.000198854)
  (COD1_COD2_COD3_gcg2ccc 0.00434629)
  (COD1_COD2_COD3_gcg2ccg 0.0237983)
  (COD1_COD2_COD3_gcg2cct 0.000146582)
  (COD1_COD2_COD3_gcg2cga 2.62182e-06)
  (COD1_COD2_COD3_gcg2cgc 0.00122331)
  (COD1_COD2_COD3_gcg2cgg 0.0278149)
  (COD1_COD2_COD3_gcg2cgt 1.27601e-05)
  (COD1_COD2_COD3_gcg2cta 2.73302e-07)
  (COD1_COD2_COD3_gcg2ctc 1.49137e-05)
  (COD1_COD2_COD3_gcg2ctg 0.0434349)
  (COD1_COD2_COD3_gcg2ctt 1.35902e-06)
  (COD1_COD2_COD3_gcg2gaa 0.00786139)
  (COD1_COD2_COD3_gcg2gac 0.0177751)
  (COD1_COD2_COD3_gcg2gag 0.0383136)
  (COD1_COD2_COD3_gcg2gat 0.00123523)
  (COD1_COD2_COD3_gcg2gca 0.191047)
  (COD1_COD2_COD3_gcg2gcc 0.51979)
  (COD1_COD2_COD3_gcg2gct 0.0971933)
  (COD1_COD2_COD3_gcg2gga 0.000763545)
  (COD1_COD2_COD3_gcg2ggc 0.0330776)
  (COD1_COD2_COD3_gcg2ggg 0.0156034)
  (COD1_COD2_COD3_gcg2ggt 0.00141106)
  (COD1_COD2_COD3_gcg2gta 0.000894891)
  (COD1_COD2_COD3_gcg2gtc 0.0178368)
  (COD1_COD2_COD3_gcg2gtg 0.0500669)
  (COD1_COD2_COD3_gcg2gtt 0.00102867)
  (COD1_COD2_COD3_gcg2taa 0)
  (COD1_COD2_COD3_gcg2tac 0.00177373)
  (COD1_COD2_COD3_gcg2tag 0)
  (COD1_COD2_COD3_gcg2tat 0.000466124)
  (COD1_COD2_COD3_gcg2tca 0.00156134)
  (COD1_COD2_COD3_gcg2tcc 0.0135929)
  (COD1_COD2_COD3_gcg2tcg 0.0406388)
  (COD1_COD2_COD3_gcg2tct 0.00121992)
  (COD1_COD2_COD3_gcg2tga 0)
  (COD1_COD2_COD3_gcg2tgc 0.00434629)
  (COD1_COD2_COD3_gcg2tgg 0.00568746)
  (COD1_COD2_COD3_gcg2tgt 0.000184384)
  (COD1_COD2_COD3_gcg2tta 1.62506e-07)
  (COD1_COD2_COD3_gcg2ttc 0.00159857)
  (COD1_COD2_COD3_gcg2ttg 0.00347204)
  (COD1_COD2_COD3_gcg2ttt 0.000125589)
  (COD1_COD2_COD3_gct2aaa 0.00672617)
  (COD1_COD2_COD3_gct2aac 2.66593e-05)
  (COD1_COD2_COD3_gct2aag 0.00717403)
  (COD1_COD2_COD3_gct2aat 0.0143282)
  (COD1_COD2_COD3_gct2aca 0.00425389)
  (COD1_COD2_COD3_gct2acc 0.00214918)
  (COD1_COD2_COD3_gct2acg 0.00115973)
  (COD1_COD2_COD3_gct2act 0.0298733)
  (COD1_COD2_COD3_gct2aga 0.000134697)
  (COD1_COD2_COD3_gct2agc 7.29365e-05)
  (COD1_COD2_COD3_gct2agg 0.000308595)
  (COD1_COD2_COD3_gct2agt 0.0239638)
  (COD1_COD2_COD3_gct2ata 2.29682e-05)
  (COD1_COD2_COD3_gct2atc 6.20129e-06)
  (COD1_COD2_COD3_gct2atg 0.0095585)
  (COD1_COD2_COD3_gct2att 0.0132855)
  (COD1_COD2_COD3_gct2caa 0.00468739)
  (COD1_COD2_COD3_gct2cac 1.21889e-06)
  (COD1_COD2_COD3_gct2cag 0.000848969)
  (COD1_COD2_COD3_gct2cat 0.00449021)
  (COD1_COD2_COD3_gct2cca 0.00387658)
  (COD1_COD2_COD3_gct2ccc 0.000759849)
  (COD1_COD2_COD3_gct2ccg 0.000164914)
  (COD1_COD2_COD3_gct2cct 0.0132671)
  (COD1_COD2_COD3_gct2cga 9.14871e-06)
  (COD1_COD2_COD3_gct2cgc 1.78695e-07)
  (COD1_COD2_COD3_gct2cgg 4.40608e-07)
  (COD1_COD2_COD3_gct2cgt 0.00275449)
  (COD1_COD2_COD3_gct2cta 3.00349e-05)
  (COD1_COD2_COD3_gct2ctc 7.37462e-06)
  (COD1_COD2_COD3_gct2ctg 2.23712e-05)
  (COD1_COD2_COD3_gct2ctt 0.0133502)
  (COD1_COD2_COD3_gct2gaa 0.017349)
  (COD1_COD2_COD3_gct2gac 0.000470995)
  (COD1_COD2_COD3_gct2gag 0.00325303)
  (COD1_COD2_COD3_gct2gat 0.0113434)
  (COD1_COD2_COD3_gct2gca 0.403041)
  (COD1_COD2_COD3_gct2gcc 0.257921)
  (COD1_COD2_COD3_gct2gcg 0.0940127)
  (COD1_COD2_COD3_gct2gga 0.0107912)
  (COD1_COD2_COD3_gct2ggc 0.000526237)
  (COD1_COD2_COD3_gct2ggg 0.00244322)
  (COD1_COD2_COD3_gct2ggt 0.0265711)
  (COD1_COD2_COD3_gct2gta 0.00723897)
  (COD1_COD2_COD3_gct2gtc 0.00243139)
  (COD1_COD2_COD3_gct2gtg 0.00344567)
  (COD1_COD2_COD3_gct2gtt 0.0494804)
  (COD1_COD2_COD3_gct2taa 0)
  (COD1_COD2_COD3_gct2tac 7.22285e-06)
  (COD1_COD2_COD3_gct2tag 0)
  (COD1_COD2_COD3_gct2tat 0.00638543)
  (COD1_COD2_COD3_gct2tca 0.0189324)
  (COD1_COD2_COD3_gct2tcc 0.00830963)
  (COD1_COD2_COD3_gct2tcg 0.00218523)
  (COD1_COD2_COD3_gct2tct 0.0471369)
  (COD1_COD2_COD3_gct2tga 0)
  (COD1_COD2_COD3_gct2tgc 8.7461e-05)
  (COD1_COD2_COD3_gct2tgg 0.000456847)
  (COD1_COD2_COD3_gct2tgt 0.0122421)
  (COD1_COD2_COD3_gct2tta 0.000144064)
  (COD1_COD2_COD3_gct2ttc 4.75541e-06)
  (COD1_COD2_COD3_gct2ttg 0.000560111)
  (COD1_COD2_COD3_gct2ttt 0.0103588)
  (COD1_COD2_COD3_gga2aaa 0.0222046)
  (COD1_COD2_COD3_gga2aac 0.0056993)
  (COD1_COD2_COD3_gga2aag 4.74906e-05)
  (COD1_COD2_COD3_gga2aat 0.0202772)
  (COD1_COD2_COD3_gga2aca 0.00612532)
  (COD1_COD2_COD3_gga2acc 1.53748e-06)
  (COD1_COD2_COD3_gga2acg 1.94743e-06)
  (COD1_COD2_COD3_gga2act 3.13538e-05)
  (COD1_COD2_COD3_gga2aga 0.00915008)
  (COD1_COD2_COD3_gga2agc 0.00409478)
  (COD1_COD2_COD3_gga2agg 0.00112489)
  (COD1_COD2_COD3_gga2agt 0.0121576)
  (COD1_COD2_COD3_gga2ata 0.0024911)
  (COD1_COD2_COD3_gga2atc 1.54282e-06)
  (COD1_COD2_COD3_gga2atg 0.000798732)
  (COD1_COD2_COD3_gga2att 7.47238e-06)
  (COD1_COD2_COD3_gga2caa 0.00947276)
  (COD1_COD2_COD3_gga2cac 5.5429e-05)
  (COD1_COD2_COD3_gga2cag 6.24875e-06)
  (COD1_COD2_COD3_gga2cat 0.00104159)
  (COD1_COD2_COD3_gga2cca 0.00621082)
  (COD1_COD2_COD3_gga2ccc 3.77786e-06)
  (COD1_COD2_COD3_gga2ccg 2.00495e-06)
  (COD1_COD2_COD3_gga2cct 3.68239e-05)
  (COD1_COD2_COD3_gga2cga 0.00118985)
  (COD1_COD2_COD3_gga2cgc 1.11237e-05)
  (COD1_COD2_COD3_gga2cgg 8.50694e-05)
  (COD1_COD2_COD3_gga2cgt 0.000130379)
  (COD1_COD2_COD3_gga2cta 0.00122076)
  (COD1_COD2_COD3_gga2ctc 6.17953e-07)
  (COD1_COD2_COD3_gga2ctg 1.50688e-06)
  (COD1_COD2_COD3_gga2ctt 5.41728e-06)
  (COD1_COD2_COD3_gga2gaa 0.0220818)
  (COD1_COD2_COD3_gga2gac 0.0017062)
  (COD1_COD2_COD3_gga2gag 0.00164579)
  (COD1_COD2_COD3_gga2gat 0.0107241)
  (COD1_COD2_COD3_gga2gca 0.0279889)
  (COD1_COD2_COD3_gga2gcc 0.000965792)
  (COD1_COD2_COD3_gga2gcg 0.000852507)
  (COD1_COD2_COD3_gga2gct 0.0124561)
  (COD1_COD2_COD3_gga2ggc 0.12345)
  (COD1_COD2_COD3_gga2ggg 0.262247)
  (COD1_COD2_COD3_gga2ggt 0.357496)
  (COD1_COD2_COD3_gga2gta 0.00354745)
  (COD1_COD2_COD3_gga2gtc 0.000179715)
  (COD1_COD2_COD3_gga2gtg 0.000380979)
  (COD1_COD2_COD3_gga2gtt 0.00152921)
  (COD1_COD2_COD3_gga2taa 0)
  (COD1_COD2_COD3_gga2tac 0.00049785)
  (COD1_COD2_COD3_gga2tag 0)
  (COD1_COD2_COD3_gga2tat 0.0014199)
  (COD1_COD2_COD3_gga2tca 0.0178492)
  (COD1_COD2_COD3_gga2tcc 2.07864e-05)
  (COD1_COD2_COD3_gga2tcg 1.54076e-05)
  (COD1_COD2_COD3_gga2tct 0.000143367)
  (COD1_COD2_COD3_gga2tga 0)
  (COD1_COD2_COD3_gga2tgc 0.000456215)
  (COD1_COD2_COD3_gga2tgg 0.000939382)
  (COD1_COD2_COD3_gga2tgt 0.00127349)
  (COD1_COD2_COD3_gga2tta 0.00325754)
  (COD1_COD2_COD3_gga2ttc 0.000391419)
  (COD1_COD2_COD3_gga2ttg 1.70366e-05)
  (COD1_COD2_COD3_gga2ttt 0.00244153)
  (COD1_COD2_COD3_ggc2aaa 8.94847e-05)
  (COD1_COD2_COD3_ggc2aac 0.022577)
  (COD1_COD2_COD3_ggc2aag 0.00470692)
  (COD1_COD2_COD3_ggc2aat 2.48568e-05)
  (COD1_COD2_COD3_ggc2aca 3.24338e-07)
  (COD1_COD2_COD3_ggc2acc 0.00928424)
  (COD1_COD2_COD3_ggc2acg 6.8573e-05)
  (COD1_COD2_COD3_ggc2act 3.58879e-07)
  (COD1_COD2_COD3_ggc2aga 7.80592e-06)
  (COD1_COD2_COD3_ggc2agc 0.0327809)
  (COD1_COD2_COD3_ggc2agg 8.31456e-05)
  (COD1_COD2_COD3_ggc2agt 0.0005574)
  (COD1_COD2_COD3_ggc2ata 9.52258e-09)
  (COD1_COD2_COD3_ggc2atc 0.00131748)
  (COD1_COD2_COD3_ggc2atg 0.000393868)
  (COD1_COD2_COD3_ggc2att 7.20285e-06)
  (COD1_COD2_COD3_ggc2caa 0.000278314)
  (COD1_COD2_COD3_ggc2cac 0.00625836)
  (COD1_COD2_COD3_ggc2cag 0.00697302)
  (COD1_COD2_COD3_ggc2cat 9.45167e-06)
  (COD1_COD2_COD3_ggc2cca 7.00554e-07)
  (COD1_COD2_COD3_ggc2ccc 0.00663356)
  (COD1_COD2_COD3_ggc2ccg 0.0048292)
  (COD1_COD2_COD3_ggc2cct 6.11985e-07)
  (COD1_COD2_COD3_ggc2cga 4.27712e-05)
  (COD1_COD2_COD3_ggc2cgc 0.00987502)
  (COD1_COD2_COD3_ggc2cgg 0.00574292)
  (COD1_COD2_COD3_ggc2cgt 0.000182098)
  (COD1_COD2_COD3_ggc2cta 1.41765e-07)
  (COD1_COD2_COD3_ggc2ctc 0.0040412)
  (COD1_COD2_COD3_ggc2ctg 0.00134797)
  (COD1_COD2_COD3_ggc2ctt 2.39412e-07)
  (COD1_COD2_COD3_ggc2gaa 0.00105613)
  (COD1_COD2_COD3_ggc2gac 0.0310557)
  (COD1_COD2_COD3_ggc2gag 0.00734002)
  (COD1_COD2_COD3_ggc2gat 0.00107087)
  (COD1_COD2_COD3_ggc2gca 0.000462263)
  (COD1_COD2_COD3_ggc2gcc 0.0482275)
  (COD1_COD2_COD3_ggc2gcg 0.0255772)
  (COD1_COD2_COD3_ggc2gct 0.000420677)
  (COD1_COD2_COD3_ggc2gga 0.085496)
  (COD1_COD2_COD3_ggc2ggg 0.191047)
  (COD1_COD2_COD3_ggc2ggt 0.23537)
  (COD1_COD2_COD3_ggc2gta 1.68604e-05)
  (COD1_COD2_COD3_ggc2gtc 0.00371351)
  (COD1_COD2_COD3_ggc2gtg 0.00115571)
  (COD1_COD2_COD3_ggc2gtt 3.94668e-05)
  (COD1_COD2_COD3_ggc2taa 0)
  (COD1_COD2_COD3_ggc2tac 0.00212842)
  (COD1_COD2_COD3_ggc2tag 0)
  (COD1_COD2_COD3_ggc2tat 5.06477e-07)
  (COD1_COD2_COD3_ggc2tca 2.75026e-06)
  (COD1_COD2_COD3_ggc2tcc 0.00947276)
  (COD1_COD2_COD3_ggc2tcg 0.00310756)
  (COD1_COD2_COD3_ggc2tct 9.00981e-07)
  (COD1_COD2_COD3_ggc2tga 0)
  (COD1_COD2_COD3_ggc2tgc 0.00229387)
  (COD1_COD2_COD3_ggc2tgg 0.00134237)
  (COD1_COD2_COD3_ggc2tgt 4.87243e-05)
  (COD1_COD2_COD3_ggc2tta 3.09175e-08)
  (COD1_COD2_COD3_ggc2ttc 0.00369298)
  (COD1_COD2_COD3_ggc2ttg 3.4261e-06)
  (COD1_COD2_COD3_ggc2ttt 2.02862e-07)
  (COD1_COD2_COD3_ggg2aaa 3.63673e-05)
  (COD1_COD2_COD3_ggg2aac 0.00597436)
  (COD1_COD2_COD3_ggg2aag 0.0198736)
  (COD1_COD2_COD3_ggg2aat 0.00434328)
  (COD1_COD2_COD3_ggg2aca 7.96442e-06)
  (COD1_COD2_COD3_ggg2acc 7.93136e-06)
  (COD1_COD2_COD3_ggg2acg 0.00765703)
  (COD1_COD2_COD3_ggg2act 7.13329e-06)
  (COD1_COD2_COD3_ggg2aga 0.000677731)
  (COD1_COD2_COD3_ggg2agc 0.0157555)
  (COD1_COD2_COD3_ggg2agg 0.0105838)
  (COD1_COD2_COD3_ggg2agt 0.00706546)
  (COD1_COD2_COD3_ggg2ata 7.81636e-07)
  (COD1_COD2_COD3_ggg2atc 6.79074e-06)
  (COD1_COD2_COD3_ggg2atg 0.00869258)
  (COD1_COD2_COD3_ggg2att 2.0161e-06)
  (COD1_COD2_COD3_ggg2caa 5.23301e-05)
  (COD1_COD2_COD3_ggg2cac 0.000593691)
  (COD1_COD2_COD3_ggg2cag 0.0131481)
  (COD1_COD2_COD3_ggg2cat 0.000896754)
  (COD1_COD2_COD3_ggg2cca 1.39054e-05)
  (COD1_COD2_COD3_ggg2ccc 1.45563e-05)
  (COD1_COD2_COD3_ggg2ccg 0.00857885)
  (COD1_COD2_COD3_ggg2cct 1.96107e-05)
  (COD1_COD2_COD3_ggg2cga 0.000689577)
  (COD1_COD2_COD3_ggg2cgc 0.0014199)
  (COD1_COD2_COD3_ggg2cgg 0.00482251)
  (COD1_COD2_COD3_ggg2cgt 0.000366983)
  (COD1_COD2_COD3_ggg2cta 4.91945e-07)
  (COD1_COD2_COD3_ggg2ctc 2.12663e-06)
  (COD1_COD2_COD3_ggg2ctg 0.00527359)
  (COD1_COD2_COD3_ggg2ctt 1.24796e-06)
  (COD1_COD2_COD3_ggg2gaa 0.00472982)
  (COD1_COD2_COD3_ggg2gac 0.0102091)
  (COD1_COD2_COD3_ggg2gag 0.0296464)
  (COD1_COD2_COD3_ggg2gat 0.00674484)
  (COD1_COD2_COD3_ggg2gca 0.0111023)
  (COD1_COD2_COD3_ggg2gcc 0.0133224)
  (COD1_COD2_COD3_ggg2gcg 0.0279114)
  (COD1_COD2_COD3_ggg2gct 0.00451831)
  (COD1_COD2_COD3_ggg2gga 0.420157)
  (COD1_COD2_COD3_ggg2ggc 0.441964)
  (COD1_COD2_COD3_ggg2ggt 0.236187)
  (COD1_COD2_COD3_ggg2gta 0.000813821)
  (COD1_COD2_COD3_ggg2gtc 0.00118409)
  (COD1_COD2_COD3_ggg2gtg 0.00625402)
  (COD1_COD2_COD3_ggg2gtt 0.000612077)
  (COD1_COD2_COD3_ggg2taa 0)
  (COD1_COD2_COD3_ggg2tac 0.000239282)
  (COD1_COD2_COD3_ggg2tag 0)
  (COD1_COD2_COD3_ggg2tat 0.000136577)
  (COD1_COD2_COD3_ggg2tca 3.45968e-05)
  (COD1_COD2_COD3_ggg2tcc 3.19461e-05)
  (COD1_COD2_COD3_ggg2tcg 0.0188669)
  (COD1_COD2_COD3_ggg2tct 3.34183e-05)
  (COD1_COD2_COD3_ggg2tga 0)
  (COD1_COD2_COD3_ggg2tgc 0.00136394)
  (COD1_COD2_COD3_ggg2tgg 0.00486616)
  (COD1_COD2_COD3_ggg2tgt 0.000682445)
  (COD1_COD2_COD3_ggg2tta 3.14377e-07)
  (COD1_COD2_COD3_ggg2ttc 0.00068434)
  (COD1_COD2_COD3_ggg2ttg 0.00276597)
  (COD1_COD2_COD3_ggg2ttt 0.000401865)
  (COD1_COD2_COD3_ggt2aaa 0.00151549)
  (COD1_COD2_COD3_ggt2aac 0.00011989)
  (COD1_COD2_COD3_ggt2aag 0.000578258)
  (COD1_COD2_COD3_ggt2aat 0.029462)
  (COD1_COD2_COD3_ggt2aca 7.14814e-06)
  (COD1_COD2_COD3_ggt2acc 2.33362e-06)
  (COD1_COD2_COD3_ggt2acg 4.23854e-06)
  (COD1_COD2_COD3_ggt2act 0.00481583)
  (COD1_COD2_COD3_ggt2aga 0.000367492)
  (COD1_COD2_COD3_ggt2agc 0.00446848)
  (COD1_COD2_COD3_ggt2agg 0.000321922)
  (COD1_COD2_COD3_ggt2agt 0.0245693)
  (COD1_COD2_COD3_ggt2ata 1.58729e-06)
  (COD1_COD2_COD3_ggt2atc 4.43673e-07)
  (COD1_COD2_COD3_ggt2atg 0.000581876)
  (COD1_COD2_COD3_ggt2att 0.00173362)
  (COD1_COD2_COD3_ggt2caa 0.000735493)
  (COD1_COD2_COD3_ggt2cac 7.03474e-07)
  (COD1_COD2_COD3_ggt2cag 7.00624e-05)
  (COD1_COD2_COD3_ggt2cat 0.00414331)
  (COD1_COD2_COD3_ggt2cca 2.82968e-05)
  (COD1_COD2_COD3_ggt2ccc 2.74075e-06)
  (COD1_COD2_COD3_ggt2ccg 1.55686e-05)
  (COD1_COD2_COD3_ggt2cct 0.00436743)
  (COD1_COD2_COD3_ggt2cga 0.00020177)
  (COD1_COD2_COD3_ggt2cgc 6.80521e-05)
  (COD1_COD2_COD3_ggt2cgg 9.64406e-05)
  (COD1_COD2_COD3_ggt2cgt 0.00204881)
  (COD1_COD2_COD3_ggt2cta 2.20315e-06)
  (COD1_COD2_COD3_ggt2ctc 6.25278e-07)
  (COD1_COD2_COD3_ggt2ctg 4.51764e-06)
  (COD1_COD2_COD3_ggt2ctt 0.00257538)
  (COD1_COD2_COD3_ggt2gaa 0.00307969)
  (COD1_COD2_COD3_ggt2gac 0.000888094)
  (COD1_COD2_COD3_ggt2gag 0.000925808)
  (COD1_COD2_COD3_ggt2gat 0.0179982)
  (COD1_COD2_COD3_ggt2gca 0.0096651)
  (COD1_COD2_COD3_ggt2gcc 0.00233235)
  (COD1_COD2_COD3_ggt2gcg 0.00152921)
  (COD1_COD2_COD3_ggt2gct 0.0297493)
  (COD1_COD2_COD3_ggt2gga 0.346998)
  (COD1_COD2_COD3_ggt2ggc 0.329877)
  (COD1_COD2_COD3_ggt2ggg 0.142991)
  (COD1_COD2_COD3_ggt2gta 0.000792116)
  (COD1_COD2_COD3_ggt2gtc 0.000308808)
  (COD1_COD2_COD3_ggt2gtg 0.000563616)
  (COD1_COD2_COD3_ggt2gtt 0.00312268)
  (COD1_COD2_COD3_ggt2taa 0)
  (COD1_COD2_COD3_ggt2tac 1.63758e-06)
  (COD1_COD2_COD3_ggt2tag 0)
  (COD1_COD2_COD3_ggt2tat 0.00225759)
  (COD1_COD2_COD3_ggt2tca 0.000121479)
  (COD1_COD2_COD3_ggt2tcc 2.09455e-05)
  (COD1_COD2_COD3_ggt2tcg 6.21882e-05)
  (COD1_COD2_COD3_ggt2tct 0.0115177)
  (COD1_COD2_COD3_ggt2tga 0)
  (COD1_COD2_COD3_ggt2tgc 0.000333969)
  (COD1_COD2_COD3_ggt2tgg 0.000500966)
  (COD1_COD2_COD3_ggt2tgt 0.00271092)
  (COD1_COD2_COD3_ggt2tta 1.18644e-05)
  (COD1_COD2_COD3_ggt2ttc 2.91515e-06)
  (COD1_COD2_COD3_ggt2ttg 6.09084e-05)
  (COD1_COD2_COD3_ggt2ttt 0.00352784)
  (COD1_COD2_COD3_gta2aaa 0.0220206)
  (COD1_COD2_COD3_gta2aac 4.80868e-05)
  (COD1_COD2_COD3_gta2aag 1.4336e-05)
  (COD1_COD2_COD3_gta2aat 0.000196934)
  (COD1_COD2_COD3_gta2aca 0.0509067)
  (COD1_COD2_COD3_gta2acc 3.52993e-05)
  (COD1_COD2_COD3_gta2acg 2.69193e-05)
  (COD1_COD2_COD3_gta2act 0.000114054)
  (COD1_COD2_COD3_gta2aga 0.00546713)
  (COD1_COD2_COD3_gta2agc 8.58947e-06)
  (COD1_COD2_COD3_gta2agg 1.42666e-05)
  (COD1_COD2_COD3_gta2agt 6.47841e-05)
  (COD1_COD2_COD3_gta2ata 0.144486)
  (COD1_COD2_COD3_gta2atc 0.00745798)
  (COD1_COD2_COD3_gta2atg 0.0120569)
  (COD1_COD2_COD3_gta2att 0.0639461)
  (COD1_COD2_COD3_gta2caa 0.0104236)
  (COD1_COD2_COD3_gta2cac 3.12237e-05)
  (COD1_COD2_COD3_gta2cag 3.33704e-06)
  (COD1_COD2_COD3_gta2cat 0.000209611)
  (COD1_COD2_COD3_gta2cca 0.00817821)
  (COD1_COD2_COD3_gta2ccc 1.51107e-06)
  (COD1_COD2_COD3_gta2ccg 1.16924e-06)
  (COD1_COD2_COD3_gta2cct 1.2437e-05)
  (COD1_COD2_COD3_gta2cga 0.00122755)
  (COD1_COD2_COD3_gta2cgc 9.50327e-08)
  (COD1_COD2_COD3_gta2cgg 2.22761e-07)
  (COD1_COD2_COD3_gta2cgt 3.42136e-06)
  (COD1_COD2_COD3_gta2cta 0.0150718)
  (COD1_COD2_COD3_gta2ctc 0.000533583)
  (COD1_COD2_COD3_gta2ctg 0.000585518)
  (COD1_COD2_COD3_gta2ctt 0.00773169)
  (COD1_COD2_COD3_gta2gaa 0.0162097)
  (COD1_COD2_COD3_gta2gac 0.000170848)
  (COD1_COD2_COD3_gta2gag 0.00162426)
  (COD1_COD2_COD3_gta2gat 0.002242)
  (COD1_COD2_COD3_gta2gca 0.0564454)
  (COD1_COD2_COD3_gta2gcc 0.000766195)
  (COD1_COD2_COD3_gta2gcg 0.00149878)
  (COD1_COD2_COD3_gta2gct 0.0125341)
  (COD1_COD2_COD3_gta2gga 0.00532132)
  (COD1_COD2_COD3_gta2ggc 3.65441e-05)
  (COD1_COD2_COD3_gta2ggg 0.000761958)
  (COD1_COD2_COD3_gta2ggt 0.001225)
  (COD1_COD2_COD3_gta2gtc 0.103881)
  (COD1_COD2_COD3_gta2gtg 0.315563)
  (COD1_COD2_COD3_gta2gtt 0.481631)
  (COD1_COD2_COD3_gta2taa 0)
  (COD1_COD2_COD3_gta2tac 0.0010096)
  (COD1_COD2_COD3_gta2tag 0)
  (COD1_COD2_COD3_gta2tat 0.00407213)
  (COD1_COD2_COD3_gta2tca 0.00843732)
  (COD1_COD2_COD3_gta2tcc 2.85318e-06)
  (COD1_COD2_COD3_gta2tcg 9.05363e-07)
  (COD1_COD2_COD3_gta2tct 1.56335e-05)
  (COD1_COD2_COD3_gta2tga 0)
  (COD1_COD2_COD3_gta2tgc 0.000644738)
  (COD1_COD2_COD3_gta2tgg 0.000746795)
  (COD1_COD2_COD3_gta2tgt 0.00485942)
  (COD1_COD2_COD3_gta2tta 0.0372659)
  (COD1_COD2_COD3_gta2ttc 0.0010124)
  (COD1_COD2_COD3_gta2ttg 0.0119902)
  (COD1_COD2_COD3_gta2ttt 0.0121155)
  (COD1_COD2_COD3_gtc2aaa 6.01949e-05)
  (COD1_COD2_COD3_gtc2aac 0.00437955)
  (COD1_COD2_COD3_gtc2aag 0.00149981)
  (COD1_COD2_COD3_gtc2aat 4.81153e-07)
  (COD1_COD2_COD3_gtc2aca 3.31186e-05)
  (COD1_COD2_COD3_gtc2acc 0.0511898)
  (COD1_COD2_COD3_gtc2acg 0.000109711)
  (COD1_COD2_COD3_gtc2act 1.34038e-05)
  (COD1_COD2_COD3_gtc2aga 2.13401e-06)
  (COD1_COD2_COD3_gtc2agc 0.00522628)
  (COD1_COD2_COD3_gtc2agg 1.39536e-05)
  (COD1_COD2_COD3_gtc2agt 5.69421e-07)
  (COD1_COD2_COD3_gtc2ata 0.00197628)
  (COD1_COD2_COD3_gtc2atc 0.178006)
  (COD1_COD2_COD3_gtc2atg 0.00742702)
  (COD1_COD2_COD3_gtc2att 0.0113592)
  (COD1_COD2_COD3_gtc2caa 0.000116047)
  (COD1_COD2_COD3_gtc2cac 0.00542185)
  (COD1_COD2_COD3_gtc2cag 0.00143772)
  (COD1_COD2_COD3_gtc2cat 4.44619e-06)
  (COD1_COD2_COD3_gtc2cca 6.40664e-06)
  (COD1_COD2_COD3_gtc2ccc 0.00791607)
  (COD1_COD2_COD3_gtc2ccg 0.000385762)
  (COD1_COD2_COD3_gtc2cct 2.18339e-06)
  (COD1_COD2_COD3_gtc2cga 8.97909e-06)
  (COD1_COD2_COD3_gtc2cgc 0.00947276)
  (COD1_COD2_COD3_gtc2cgg 4.09437e-05)
  (COD1_COD2_COD3_gtc2cgt 5.23275e-06)
  (COD1_COD2_COD3_gtc2cta 0.000321922)
  (COD1_COD2_COD3_gtc2ctc 0.045721)
  (COD1_COD2_COD3_gtc2ctg 0.0309268)
  (COD1_COD2_COD3_gtc2ctt 0.00159857)
  (COD1_COD2_COD3_gtc2gaa 0.00146997)
  (COD1_COD2_COD3_gtc2gac 0.00301004)
  (COD1_COD2_COD3_gtc2gag 0.00749945)
  (COD1_COD2_COD3_gtc2gat 0.00020402)
  (COD1_COD2_COD3_gtc2gca 0.00152709)
  (COD1_COD2_COD3_gtc2gcc 0.0703162)
  (COD1_COD2_COD3_gtc2gcg 0.0177136)
  (COD1_COD2_COD3_gtc2gct 0.00249629)
  (COD1_COD2_COD3_gtc2gga 0.000159849)
  (COD1_COD2_COD3_gtc2ggc 0.00476932)
  (COD1_COD2_COD3_gtc2ggg 0.000656918)
  (COD1_COD2_COD3_gtc2ggt 0.000283179)
  (COD1_COD2_COD3_gtc2gta 0.0615968)
  (COD1_COD2_COD3_gtc2gtg 0.424254)
  (COD1_COD2_COD3_gtc2gtt 0.208483)
  (COD1_COD2_COD3_gtc2taa 0)
  (COD1_COD2_COD3_gtc2tac 0.00973232)
  (COD1_COD2_COD3_gtc2tag 0)
  (COD1_COD2_COD3_gtc2tat 2.0147e-06)
  (COD1_COD2_COD3_gtc2tca 1.31273e-06)
  (COD1_COD2_COD3_gtc2tcc 0.00427162)
  (COD1_COD2_COD3_gtc2tcg 1.07896e-05)
  (COD1_COD2_COD3_gtc2tct 5.32395e-07)
  (COD1_COD2_COD3_gtc2tga 0)
  (COD1_COD2_COD3_gtc2tgc 0.00944653)
  (COD1_COD2_COD3_gtc2tgg 0.00117347)
  (COD1_COD2_COD3_gtc2tgt 7.32876e-06)
  (COD1_COD2_COD3_gtc2tta 9.69769e-05)
  (COD1_COD2_COD3_gtc2ttc 0.020207)
  (COD1_COD2_COD3_gtc2ttg 0.0022142)
  (COD1_COD2_COD3_gtc2ttt 0.000564007)
  (COD1_COD2_COD3_gtg2aaa 6.1244e-06)
  (COD1_COD2_COD3_gtg2aac 0.000275817)
  (COD1_COD2_COD3_gtg2aag 0.00832693)
  (COD1_COD2_COD3_gtg2aat 1.44157e-05)
  (COD1_COD2_COD3_gtg2aca 0.000150182)
  (COD1_COD2_COD3_gtg2acc 0.00160301)
  (COD1_COD2_COD3_gtg2acg 0.0330547)
  (COD1_COD2_COD3_gtg2act 4.6096e-05)
  (COD1_COD2_COD3_gtg2aga 5.84214e-07)
  (COD1_COD2_COD3_gtg2agc 5.71059e-05)
  (COD1_COD2_COD3_gtg2agg 0.00213433)
  (COD1_COD2_COD3_gtg2agt 1.38476e-05)
  (COD1_COD2_COD3_gtg2ata 0.0116785)
  (COD1_COD2_COD3_gtg2atc 0.0828704)
  (COD1_COD2_COD3_gtg2atg 0.0296464)
  (COD1_COD2_COD3_gtg2att 0.0236175)
  (COD1_COD2_COD3_gtg2caa 8.04766e-06)
  (COD1_COD2_COD3_gtg2cac 0.000486929)
  (COD1_COD2_COD3_gtg2cag 0.00986134)
  (COD1_COD2_COD3_gtg2cat 0.000344311)
  (COD1_COD2_COD3_gtg2cca 7.84932e-06)
  (COD1_COD2_COD3_gtg2ccc 1.59731e-05)
  (COD1_COD2_COD3_gtg2ccg 0.0073502)
  (COD1_COD2_COD3_gtg2cct 4.27395e-06)
  (COD1_COD2_COD3_gtg2cga 4.035e-06)
  (COD1_COD2_COD3_gtg2cgc 4.77877e-05)
  (COD1_COD2_COD3_gtg2cgg 0.00572305)
  (COD1_COD2_COD3_gtg2cgt 1.22913e-05)
  (COD1_COD2_COD3_gtg2cta 0.00127525)
  (COD1_COD2_COD3_gtg2ctc 0.00802101)
  (COD1_COD2_COD3_gtg2ctg 0.0644802)
  (COD1_COD2_COD3_gtg2ctt 0.00139357)
  (COD1_COD2_COD3_gtg2gaa 0.00189972)
  (COD1_COD2_COD3_gtg2gac 0.00104448)
  (COD1_COD2_COD3_gtg2gag 0.0110946)
  (COD1_COD2_COD3_gtg2gat 0.000328004)
  (COD1_COD2_COD3_gtg2gca 0.0068675)
  (COD1_COD2_COD3_gtg2gcc 0.0183128)
  (COD1_COD2_COD3_gtg2gcg 0.0404701)
  (COD1_COD2_COD3_gtg2gct 0.00287943)
  (COD1_COD2_COD3_gtg2gga 0.000276008)
  (COD1_COD2_COD3_gtg2ggc 0.00120814)
  (COD1_COD2_COD3_gtg2ggg 0.00282604)
  (COD1_COD2_COD3_gtg2ggt 0.000420677)
  (COD1_COD2_COD3_gtg2gta 0.152301)
  (COD1_COD2_COD3_gtg2gtc 0.345319)
  (COD1_COD2_COD3_gtg2gtt 0.168054)
  (COD1_COD2_COD3_gtg2taa 0)
  (COD1_COD2_COD3_gtg2tac 0.00194232)
  (COD1_COD2_COD3_gtg2tag 0)
  (COD1_COD2_COD3_gtg2tat 0.000166176)
  (COD1_COD2_COD3_gtg2tca 7.86566e-06)
  (COD1_COD2_COD3_gtg2tcc 1.4526e-05)
  (COD1_COD2_COD3_gtg2tcg 0.00394161)
  (COD1_COD2_COD3_gtg2tct 2.70676e-06)
  (COD1_COD2_COD3_gtg2tga 0)
  (COD1_COD2_COD3_gtg2tgc 0.00393342)
  (COD1_COD2_COD3_gtg2tgg 0.00353273)
  (COD1_COD2_COD3_gtg2tgt 0.000817213)
  (COD1_COD2_COD3_gtg2tta 0.000400475)
  (COD1_COD2_COD3_gtg2ttc 0.00792705)
  (COD1_COD2_COD3_gtg2ttg 0.0174334)
  (COD1_COD2_COD3_gtg2ttt 0.00147712)
  (COD1_COD2_COD3_gtt2aaa 0.00626704)
  (COD1_COD2_COD3_gtt2aac 6.09476e-06)
  (COD1_COD2_COD3_gtt2aag 0.00192624)
  (COD1_COD2_COD3_gtt2aat 0.00709983)
  (COD1_COD2_COD3_gtt2aca 0.000738559)
  (COD1_COD2_COD3_gtt2acc 5.4063e-05)
  (COD1_COD2_COD3_gtt2acg 5.16096e-05)
  (COD1_COD2_COD3_gtt2act 0.0367019)
  (COD1_COD2_COD3_gtt2aga 0.000313336)
  (COD1_COD2_COD3_gtt2agc 3.38362e-06)
  (COD1_COD2_COD3_gtt2agg 0.000351546)
  (COD1_COD2_COD3_gtt2agt 0.00517222)
  (COD1_COD2_COD3_gtt2ata 0.0330318)
  (COD1_COD2_COD3_gtt2atc 0.0136875)
  (COD1_COD2_COD3_gtt2atg 0.0091121)
  (COD1_COD2_COD3_gtt2att 0.162105)
  (COD1_COD2_COD3_gtt2caa 0.0025897)
  (COD1_COD2_COD3_gtt2cac 9.62974e-07)
  (COD1_COD2_COD3_gtt2cag 0.00019913)
  (COD1_COD2_COD3_gtt2cat 0.00360946)
  (COD1_COD2_COD3_gtt2cca 0.000103291)
  (COD1_COD2_COD3_gtt2ccc 1.53961e-06)
  (COD1_COD2_COD3_gtt2ccg 2.90305e-06)
  (COD1_COD2_COD3_gtt2cct 0.00738595)
  (COD1_COD2_COD3_gtt2cga 1.82594e-05)
  (COD1_COD2_COD3_gtt2cgc 1.64319e-07)
  (COD1_COD2_COD3_gtt2cgg 3.8304e-07)
  (COD1_COD2_COD3_gtt2cgt 0.00286153)
  (COD1_COD2_COD3_gtt2cta 0.00408627)
  (COD1_COD2_COD3_gtt2ctc 0.00122415)
  (COD1_COD2_COD3_gtt2ctg 0.000330056)
  (COD1_COD2_COD3_gtt2ctt 0.0287758)
  (COD1_COD2_COD3_gtt2gaa 0.00955188)
  (COD1_COD2_COD3_gtt2gac 0.000104154)
  (COD1_COD2_COD3_gtt2gag 0.00152603)
  (COD1_COD2_COD3_gtt2gat 0.00438867)
  (COD1_COD2_COD3_gtt2gca 0.0118744)
  (COD1_COD2_COD3_gtt2gcc 0.00124988)
  (COD1_COD2_COD3_gtt2gcg 0.000988823)
  (COD1_COD2_COD3_gtt2gct 0.0491727)
  (COD1_COD2_COD3_gtt2gga 0.00131748)
  (COD1_COD2_COD3_gtt2ggc 4.90972e-05)
  (COD1_COD2_COD3_gtt2ggg 0.000328914)
  (COD1_COD2_COD3_gtt2ggt 0.00277173)
  (COD1_COD2_COD3_gtt2gta 0.276624)
  (COD1_COD2_COD3_gtt2gtc 0.20194)
  (COD1_COD2_COD3_gtt2gtg 0.199851)
  (COD1_COD2_COD3_gtt2taa 0)
  (COD1_COD2_COD3_gtt2tac 6.32282e-06)
  (COD1_COD2_COD3_gtt2tag 0)
  (COD1_COD2_COD3_gtt2tat 0.00985451)
  (COD1_COD2_COD3_gtt2tca 2.3144e-05)
  (COD1_COD2_COD3_gtt2tcc 2.20927e-06)
  (COD1_COD2_COD3_gtt2tcg 1.16762e-06)
  (COD1_COD2_COD3_gtt2tct 0.00665659)
  (COD1_COD2_COD3_gtt2tga 0)
  (COD1_COD2_COD3_gtt2tgc 3.17474e-05)
  (COD1_COD2_COD3_gtt2tgg 0.000494069)
  (COD1_COD2_COD3_gtt2tgt 0.0103015)
  (COD1_COD2_COD3_gtt2tta 0.0121239)
  (COD1_COD2_COD3_gtt2ttc 0.00140813)
  (COD1_COD2_COD3_gtt2ttg 0.00832693)
  (COD1_COD2_COD3_gtt2ttt 0.0185042)
  (COD1_COD2_COD3_taa2aaa 0)
  (COD1_COD2_COD3_taa2aac 0)
  (COD1_COD2_COD3_taa2aag 0)
  (COD1_COD2_COD3_taa2aat 0)
  (COD1_COD2_COD3_taa2aca 0)
  (COD1_COD2_COD3_taa2acc 0)
  (COD1_COD2_COD3_taa2acg 0)
  (COD1_COD2_COD3_taa2act 0)
  (COD1_COD2_COD3_taa2aga 0)
  (COD1_COD2_COD3_taa2agc 0)
  (COD1_COD2_COD3_taa2agg 0)
  (COD1_COD2_COD3_taa2agt 0)
  (COD1_COD2_COD3_taa2ata 0)
  (COD1_COD2_COD3_taa2atc 0)
  (COD1_COD2_COD3_taa2atg 0)
  (COD1_COD2_COD3_taa2att 0)
  (COD1_COD2_COD3_taa2caa 0)
  (COD1_COD2_COD3_taa2cac 0)
  (COD1_COD2_COD3_taa2cag 0)
  (COD1_COD2_COD3_taa2cat 0)
  (COD1_COD2_COD3_taa2cca 0)
  (COD1_COD2_COD3_taa2ccc 0)
  (COD1_COD2_COD3_taa2ccg 0)
  (COD1_COD2_COD3_taa2cct 0)
  (COD1_COD2_COD3_taa2cga 0)
  (COD1_COD2_COD3_taa2cgc 0)
  (COD1_COD2_COD3_taa2cgg 0)
  (COD1_COD2_COD3_taa2cgt 0)
  (COD1_COD2_COD3_taa2cta 0)
  (COD1_COD2_COD3_taa2ctc 0)
  (COD1_COD2_COD3_taa2ctg 0)
  (COD1_COD2_COD3_taa2ctt 0)
  (COD1_COD2_COD3_taa2gaa 0)
  (COD1_COD2_COD3_taa2gac 0)
  (COD1_COD2_COD3_taa2gag 0)
  (COD1_COD2_COD3_taa2gat 0)
  (COD1_COD2_COD3_taa2gca 0)
  (COD1_COD2_COD3_taa2gcc 0)
  (COD1_COD2_COD3_taa2gcg 0)
  (COD1_COD2_COD3_taa2gct 0)
  (COD1_COD2_COD3_taa2gga 0)
  (COD1_COD2_COD3_taa2ggc 0)
  (COD1_COD2_COD3_taa2ggg 0)
  (COD1_COD2_COD3_taa2ggt 0)
  (COD1_COD2_COD3_taa2gta 0)
  (COD1_COD2_COD3_taa2gtc 0)
  (COD1_COD2_COD3_taa2gtg 0)
  (COD1_COD2_COD3_taa2gtt 0)
  (COD1_COD2_COD3_taa2tac 0)
  (COD1_COD2_COD3_taa2tag 0)
  (COD1_COD2_COD3_taa2tat 0)
  (COD1_COD2_COD3_taa2tca 0)
  (COD1_COD2_COD3_taa2tcc 0)
  (COD1_COD2_COD3_taa2tcg 0)
  (COD1_COD2_COD3_taa2tct 0)
  (COD1_COD2_COD3_taa2tga 0)
  (COD1_COD2_COD3_taa2tgc 0)
  (COD1_COD2_COD3_taa2tgg 0)
  (COD1_COD2_COD3_taa2tgt 0)
  (COD1_COD2_COD3_taa2tta 0)
  (COD1_COD2_COD3_taa2ttc 0)
  (COD1_COD2_COD3_taa2ttg 0)
  (COD1_COD2_COD3_taa2ttt 0)
  (COD1_COD2_COD3_tac2aaa 0.00133495)
  (COD1_COD2_COD3_tac2aac 0.00991618)
  (COD1_COD2_COD3_tac2aag 0.00376274)
  (COD1_COD2_COD3_tac2aat 0.000952496)
  (COD1_COD2_COD3_tac2aca 0.000700659)
  (COD1_COD2_COD3_tac2acc 0.00704102)
  (COD1_COD2_COD3_tac2acg 0.00189972)
  (COD1_COD2_COD3_tac2act 7.59773e-06)
  (COD1_COD2_COD3_tac2aga 0.000593691)
  (COD1_COD2_COD3_tac2agc 0.00475282)
  (COD1_COD2_COD3_tac2agg 0.00182992)
  (COD1_COD2_COD3_tac2agt 1.5906e-06)
  (COD1_COD2_COD3_tac2ata 0.00032619)
  (COD1_COD2_COD3_tac2atc 0.00608301)
  (COD1_COD2_COD3_tac2atg 0.00636334)
  (COD1_COD2_COD3_tac2att 4.01825e-06)
  (COD1_COD2_COD3_tac2caa 0.00136111)
  (COD1_COD2_COD3_tac2cac 0.0352801)
  (COD1_COD2_COD3_tac2cag 0.00689134)
  (COD1_COD2_COD3_tac2cat 0.00363457)
  (COD1_COD2_COD3_tac2cca 0.000288928)
  (COD1_COD2_COD3_tac2ccc 0.0032328)
  (COD1_COD2_COD3_tac2ccg 0.000455898)
  (COD1_COD2_COD3_tac2cct 5.21826e-06)
  (COD1_COD2_COD3_tac2cga 0.00024858)
  (COD1_COD2_COD3_tac2cgc 0.00883227)
  (COD1_COD2_COD3_tac2cgg 0.0006684)
  (COD1_COD2_COD3_tac2cgt 1.17988e-05)
  (COD1_COD2_COD3_tac2cta 8.80084e-05)
  (COD1_COD2_COD3_tac2ctc 0.0104526)
  (COD1_COD2_COD3_tac2ctg 0.00880171)
  (COD1_COD2_COD3_tac2ctt 4.75211e-06)
  (COD1_COD2_COD3_tac2gaa 0.00151025)
  (COD1_COD2_COD3_tac2gac 0.00467117)
  (COD1_COD2_COD3_tac2gag 0.00417502)
  (COD1_COD2_COD3_tac2gat 0.000442511)
  (COD1_COD2_COD3_tac2gca 0.000574263)
  (COD1_COD2_COD3_tac2gcc 0.0102162)
  (COD1_COD2_COD3_tac2gcg 0.00194232)
  (COD1_COD2_COD3_tac2gct 8.17698e-06)
  (COD1_COD2_COD3_tac2gga 0.00048862)
  (COD1_COD2_COD3_tac2ggc 0.00301631)
  (COD1_COD2_COD3_tac2ggg 0.000146582)
  (COD1_COD2_COD3_tac2ggt 1.65585e-06)
  (COD1_COD2_COD3_tac2gta 0.000660113)
  (COD1_COD2_COD3_tac2gtc 0.010739)
  (COD1_COD2_COD3_tac2gtg 0.00263314)
  (COD1_COD2_COD3_tac2gtt 7.20285e-06)
  (COD1_COD2_COD3_tac2taa 0)
  (COD1_COD2_COD3_tac2tag 0)
  (COD1_COD2_COD3_tac2tat 0.416966)
  (COD1_COD2_COD3_tac2tca 0.000733966)
  (COD1_COD2_COD3_tac2tcc 0.00564035)
  (COD1_COD2_COD3_tac2tcg 0.00186579)
  (COD1_COD2_COD3_tac2tct 0.000821757)
  (COD1_COD2_COD3_tac2tga 0)
  (COD1_COD2_COD3_tac2tgc 0.00466147)
  (COD1_COD2_COD3_tac2tgg 0.0181234)
  (COD1_COD2_COD3_tac2tgt 0.000353501)
  (COD1_COD2_COD3_tac2tta 0.00054291)
  (COD1_COD2_COD3_tac2ttc 0.0957224)
  (COD1_COD2_COD3_tac2ttg 0.00279488)
  (COD1_COD2_COD3_tac2ttt 0.0103015)
  (COD1_COD2_COD3_tag2aaa 0)
  (COD1_COD2_COD3_tag2aac 0)
  (COD1_COD2_COD3_tag2aag 0)
  (COD1_COD2_COD3_tag2aat 0)
  (COD1_COD2_COD3_tag2aca 0)
  (COD1_COD2_COD3_tag2acc 0)
  (COD1_COD2_COD3_tag2acg 0)
  (COD1_COD2_COD3_tag2act 0)
  (COD1_COD2_COD3_tag2aga 0)
  (COD1_COD2_COD3_tag2agc 0)
  (COD1_COD2_COD3_tag2agg 0)
  (COD1_COD2_COD3_tag2agt 0)
  (COD1_COD2_COD3_tag2ata 0)
  (COD1_COD2_COD3_tag2atc 0)
  (COD1_COD2_COD3_tag2atg 0)
  (COD1_COD2_COD3_tag2att 0)
  (COD1_COD2_COD3_tag2caa 0)
  (COD1_COD2_COD3_tag2cac 0)
  (COD1_COD2_COD3_tag2cag 0)
  (COD1_COD2_COD3_tag2cat 0)
  (COD1_COD2_COD3_tag2cca 0)
  (COD1_COD2_COD3_tag2ccc 0)
  (COD1_COD2_COD3_tag2ccg 0)
  (COD1_COD2_COD3_tag2cct 0)
  (COD1_COD2_COD3_tag2cga 0)
  (COD1_COD2_COD3_tag2cgc 0)
  (COD1_COD2_COD3_tag2cgg 0)
  (COD1_COD2_COD3_tag2cgt 0)
  (COD1_COD2_COD3_tag2cta 0)
  (COD1_COD2_COD3_tag2ctc 0)
  (COD1_COD2_COD3_tag2ctg 0)
  (COD1_COD2_COD3_tag2ctt 0)
  (COD1_COD2_COD3_tag2gaa 0)
  (COD1_COD2_COD3_tag2gac 0)
  (COD1_COD2_COD3_tag2gag 0)
  (COD1_COD2_COD3_tag2gat 0)
  (COD1_COD2_COD3_tag2gca 0)
  (COD1_COD2_COD3_tag2gcc 0)
  (COD1_COD2_COD3_tag2gcg 0)
  (COD1_COD2_COD3_tag2gct 0)
  (COD1_COD2_COD3_tag2gga 0)
  (COD1_COD2_COD3_tag2ggc 0)
  (COD1_COD2_COD3_tag2ggg 0)
  (COD1_COD2_COD3_tag2ggt 0)
  (COD1_COD2_COD3_tag2gta 0)
  (COD1_COD2_COD3_tag2gtc 0)
  (COD1_COD2_COD3_tag2gtg 0)
  (COD1_COD2_COD3_tag2gtt 0)
  (COD1_COD2_COD3_tag2taa 0)
  (COD1_COD2_COD3_tag2tac 0)
  (COD1_COD2_COD3_tag2tat 0)
  (COD1_COD2_COD3_tag2tca 0)
  (COD1_COD2_COD3_tag2tcc 0)
  (COD1_COD2_COD3_tag2tcg 0)
  (COD1_COD2_COD3_tag2tct 0)
  (COD1_COD2_COD3_tag2tga 0)
  (COD1_COD2_COD3_tag2tgc 0)
  (COD1_COD2_COD3_tag2tgg 0)
  (COD1_COD2_COD3_tag2tgt 0)
  (COD1_COD2_COD3_tag2tta 0)
  (COD1_COD2_COD3_tag2ttc 0)
  (COD1_COD2_COD3_tag2ttg 0)
  (COD1_COD2_COD3_tag2ttt 0)
  (COD1_COD2_COD3_tat2aaa 0.0145282)
  (COD1_COD2_COD3_tat2aac 0.000947228)
  (COD1_COD2_COD3_tat2aag 0.00194637)
  (COD1_COD2_COD3_tat2aat 0.0166654)
  (COD1_COD2_COD3_tat2aca 0.00272978)
  (COD1_COD2_COD3_tat2acc 1.65241e-06)
  (COD1_COD2_COD3_tat2acg 0.000450246)
  (COD1_COD2_COD3_tat2act 0.0075936)
  (COD1_COD2_COD3_tat2aga 0.00413757)
  (COD1_COD2_COD3_tat2agc 1.88109e-06)
  (COD1_COD2_COD3_tat2agg 0.00136111)
  (COD1_COD2_COD3_tat2agt 0.00522266)
  (COD1_COD2_COD3_tat2ata 0.00734511)
  (COD1_COD2_COD3_tat2atc 2.98053e-06)
  (COD1_COD2_COD3_tat2atg 0.00623671)
  (COD1_COD2_COD3_tat2att 0.00873486)
  (COD1_COD2_COD3_tat2caa 0.00835005)
  (COD1_COD2_COD3_tat2cac 0.00293383)
  (COD1_COD2_COD3_tat2cag 0.00190896)
  (COD1_COD2_COD3_tat2cat 0.0329632)
  (COD1_COD2_COD3_tat2cca 0.000358684)
  (COD1_COD2_COD3_tat2ccc 9.23746e-07)
  (COD1_COD2_COD3_tat2ccg 5.10404e-05)
  (COD1_COD2_COD3_tat2cct 0.00352295)
  (COD1_COD2_COD3_tat2cga 0.000211215)
  (COD1_COD2_COD3_tat2cgc 1.83983e-06)
  (COD1_COD2_COD3_tat2cgg 3.05176e-05)
  (COD1_COD2_COD3_tat2cgt 0.00454029)
  (COD1_COD2_COD3_tat2cta 0.000122749)
  (COD1_COD2_COD3_tat2ctc 1.85648e-06)
  (COD1_COD2_COD3_tat2ctg 0.000137051)
  (COD1_COD2_COD3_tat2ctt 0.0073655)
  (COD1_COD2_COD3_tat2gaa 0.00888138)
  (COD1_COD2_COD3_tat2gac 0.000253628)
  (COD1_COD2_COD3_tat2gag 0.00131111)
  (COD1_COD2_COD3_tat2gat 0.00639872)
  (COD1_COD2_COD3_tat2gca 0.00292165)
  (COD1_COD2_COD3_tat2gcc 1.8968e-06)
  (COD1_COD2_COD3_tat2gcg 0.000494754)
  (COD1_COD2_COD3_tat2gct 0.00700208)
  (COD1_COD2_COD3_tat2gga 0.00134984)
  (COD1_COD2_COD3_tat2ggc 6.95233e-07)
  (COD1_COD2_COD3_tat2ggg 8.09842e-05)
  (COD1_COD2_COD3_tat2ggt 0.00221113)
  (COD1_COD2_COD3_tat2gta 0.00258074)
  (COD1_COD2_COD3_tat2gtc 2.15333e-06)
  (COD1_COD2_COD3_tat2gtg 0.000218209)
  (COD1_COD2_COD3_tat2gtt 0.0108738)
  (COD1_COD2_COD3_tat2taa 0)
  (COD1_COD2_COD3_tat2tac 0.40388)
  (COD1_COD2_COD3_tat2tag 0)
  (COD1_COD2_COD3_tat2tca 0.003545)
  (COD1_COD2_COD3_tat2tcc 0.000507255)
  (COD1_COD2_COD3_tat2tcg 0.000507607)
  (COD1_COD2_COD3_tat2tct 0.00812171)
  (COD1_COD2_COD3_tat2tga 0)
  (COD1_COD2_COD3_tat2tgc 0.000506904)
  (COD1_COD2_COD3_tat2tgg 0.0145383)
  (COD1_COD2_COD3_tat2tgt 0.00530291)
  (COD1_COD2_COD3_tat2tta 0.0129402)
  (COD1_COD2_COD3_tat2ttc 0.0107539)
  (COD1_COD2_COD3_tat2ttg 0.00530291)
  (COD1_COD2_COD3_tat2ttt 0.0925258)
  (COD1_COD2_COD3_tca2aaa 0.0465524)
  (COD1_COD2_COD3_tca2aac 0.00381791)
  (COD1_COD2_COD3_tca2aag 0.000118815)
  (COD1_COD2_COD3_tca2aat 0.0194377)
  (COD1_COD2_COD3_tca2aca 0.0749462)
  (COD1_COD2_COD3_tca2acc 0.00268474)
  (COD1_COD2_COD3_tca2acg 0.00552044)
  (COD1_COD2_COD3_tca2act 0.0290967)
  (COD1_COD2_COD3_tca2aga 0.0106722)
  (COD1_COD2_COD3_tca2agc 0.0143879)
  (COD1_COD2_COD3_tca2agg 6.29253e-05)
  (COD1_COD2_COD3_tca2agt 0.0615115)
  (COD1_COD2_COD3_tca2ata 0.00517222)
  (COD1_COD2_COD3_tca2atc 1.38278e-06)
  (COD1_COD2_COD3_tca2atg 0.00521181)
  (COD1_COD2_COD3_tca2att 9.04781e-06)
  (COD1_COD2_COD3_tca2caa 0.0355009)
  (COD1_COD2_COD3_tca2cac 0.000160071)
  (COD1_COD2_COD3_tca2cag 4.44025e-05)
  (COD1_COD2_COD3_tca2cat 0.00136773)
  (COD1_COD2_COD3_tca2cca 0.0348669)
  (COD1_COD2_COD3_tca2ccc 0.000753555)
  (COD1_COD2_COD3_tca2ccg 0.000792116)
  (COD1_COD2_COD3_tca2cct 0.00567958)
  (COD1_COD2_COD3_tca2cga 0.00267917)
  (COD1_COD2_COD3_tca2cgc 1.3449e-07)
  (COD1_COD2_COD3_tca2cgg 2.74441e-07)
  (COD1_COD2_COD3_tca2cgt 4.78848e-06)
  (COD1_COD2_COD3_tca2cta 0.00365225)
  (COD1_COD2_COD3_tca2ctc 1.38854e-06)
  (COD1_COD2_COD3_tca2ctg 7.05916e-07)
  (COD1_COD2_COD3_tca2ctt 4.81845e-06)
  (COD1_COD2_COD3_tca2gaa 0.0437068)
  (COD1_COD2_COD3_tca2gac 0.000187347)
  (COD1_COD2_COD3_tca2gag 3.15281e-05)
  (COD1_COD2_COD3_tca2gat 0.00996441)
  (COD1_COD2_COD3_tca2gca 0.0723933)
  (COD1_COD2_COD3_tca2gcc 0.00225916)
  (COD1_COD2_COD3_tca2gcg 0.00275449)
  (COD1_COD2_COD3_tca2gct 0.0345301)
  (COD1_COD2_COD3_tca2gga 0.0282031)
  (COD1_COD2_COD3_tca2ggc 6.27915e-06)
  (COD1_COD2_COD3_tca2ggg 3.41205e-05)
  (COD1_COD2_COD3_tca2ggt 0.000197892)
  (COD1_COD2_COD3_tca2gta 0.00888754)
  (COD1_COD2_COD3_tca2gtc 2.33362e-06)
  (COD1_COD2_COD3_tca2gtg 1.7167e-05)
  (COD1_COD2_COD3_tca2gtt 4.24758e-05)
  (COD1_COD2_COD3_tca2taa 0)
  (COD1_COD2_COD3_tca2tac 0.00118163)
  (COD1_COD2_COD3_tca2tag 0)
  (COD1_COD2_COD3_tca2tat 0.00589211)
  (COD1_COD2_COD3_tca2tcc 0.172062)
  (COD1_COD2_COD3_tca2tcg 0.151985)
  (COD1_COD2_COD3_tca2tct 0.42396)
  (COD1_COD2_COD3_tca2tga 0)
  (COD1_COD2_COD3_tca2tgc 0.000904244)
  (COD1_COD2_COD3_tca2tgg 0.00249629)
  (COD1_COD2_COD3_tca2tgt 0.00484597)
  (COD1_COD2_COD3_tca2tta 0.0191967)
  (COD1_COD2_COD3_tca2ttc 0.00121065)
  (COD1_COD2_COD3_tca2ttg 0.00961832)
  (COD1_COD2_COD3_tca2ttt 0.00595782)
  (COD1_COD2_COD3_tcc2aaa 0.000650123)
  (COD1_COD2_COD3_tcc2aac 0.0227341)
  (COD1_COD2_COD3_tcc2aag 0.0067215)
  (COD1_COD2_COD3_tcc2aat 2.03869e-05)
  (COD1_COD2_COD3_tcc2aca 0.00537694)
  (COD1_COD2_COD3_tcc2acc 0.0684882)
  (COD1_COD2_COD3_tcc2acg 0.0167001)
  (COD1_COD2_COD3_tcc2act 0.00616792)
  (COD1_COD2_COD3_tcc2aga 1.96515e-05)
  (COD1_COD2_COD3_tcc2agc 0.14927)
  (COD1_COD2_COD3_tcc2agg 0.000194627)
  (COD1_COD2_COD3_tcc2agt 0.00086321)
  (COD1_COD2_COD3_tcc2ata 3.14159e-07)
  (COD1_COD2_COD3_tcc2atc 0.00275831)
  (COD1_COD2_COD3_tcc2atg 0.000896133)
  (COD1_COD2_COD3_tcc2att 9.81166e-07)
  (COD1_COD2_COD3_tcc2caa 0.00138107)
  (COD1_COD2_COD3_tcc2cac 0.0126388)
  (COD1_COD2_COD3_tcc2cag 0.00877734)
  (COD1_COD2_COD3_tcc2cat 2.2652e-05)
  (COD1_COD2_COD3_tcc2cca 0.00154948)
  (COD1_COD2_COD3_tcc2ccc 0.0258983)
  (COD1_COD2_COD3_tcc2ccg 0.00452772)
  (COD1_COD2_COD3_tcc2cct 0.00204739)
  (COD1_COD2_COD3_tcc2cga 1.957e-05)
  (COD1_COD2_COD3_tcc2cgc 0.0100198)
  (COD1_COD2_COD3_tcc2cgg 3.92758e-05)
  (COD1_COD2_COD3_tcc2cgt 1.03788e-05)
  (COD1_COD2_COD3_tcc2cta 1.50793e-06)
  (COD1_COD2_COD3_tcc2ctc 0.00575886)
  (COD1_COD2_COD3_tcc2ctg 2.86918e-05)
  (COD1_COD2_COD3_tcc2ctt 2.71993e-06)
  (COD1_COD2_COD3_tcc2gaa 0.00295833)
  (COD1_COD2_COD3_tcc2gac 0.0258803)
  (COD1_COD2_COD3_tcc2gag 0.00867452)
  (COD1_COD2_COD3_tcc2gat 0.000106268)
  (COD1_COD2_COD3_tcc2gca 0.00922651)
  (COD1_COD2_COD3_tcc2gcc 0.09259)
  (COD1_COD2_COD3_tcc2gcg 0.0230994)
  (COD1_COD2_COD3_tcc2gct 0.0145989)
  (COD1_COD2_COD3_tcc2gga 3.16376e-05)
  (COD1_COD2_COD3_tcc2ggc 0.0208328)
  (COD1_COD2_COD3_tcc2ggg 3.03488e-05)
  (COD1_COD2_COD3_tcc2ggt 3.28442e-05)
  (COD1_COD2_COD3_tcc2gta 2.89501e-06)
  (COD1_COD2_COD3_tcc2gtc 0.00731463)
  (COD1_COD2_COD3_tcc2gtg 3.05599e-05)
  (COD1_COD2_COD3_tcc2gtt 3.90567e-06)
  (COD1_COD2_COD3_tcc2taa 0)
  (COD1_COD2_COD3_tcc2tac 0.00875304)
  (COD1_COD2_COD3_tcc2tag 0)
  (COD1_COD2_COD3_tcc2tat 0.000812694)
  (COD1_COD2_COD3_tcc2tca 0.165741)
  (COD1_COD2_COD3_tcc2tcg 0.308427)
  (COD1_COD2_COD3_tcc2tct 0.279128)
  (COD1_COD2_COD3_tcc2tga 0)
  (COD1_COD2_COD3_tcc2tgc 0.00982042)
  (COD1_COD2_COD3_tcc2tgg 0.00103726)
  (COD1_COD2_COD3_tcc2tgt 0.00221266)
  (COD1_COD2_COD3_tcc2tta 0.000188128)
  (COD1_COD2_COD3_tcc2ttc 0.0140139)
  (COD1_COD2_COD3_tcc2ttg 0.00249802)
  (COD1_COD2_COD3_tcc2ttt 0.000644291)
  (COD1_COD2_COD3_tcg2aaa 1.4516e-05)
  (COD1_COD2_COD3_tcg2aac 0.0147514)
  (COD1_COD2_COD3_tcg2aag 0.0205317)
  (COD1_COD2_COD3_tcg2aat 0.000907383)
  (COD1_COD2_COD3_tcg2aca 0.00459092)
  (COD1_COD2_COD3_tcg2acc 0.0428667)
  (COD1_COD2_COD3_tcg2acg 0.052556)
  (COD1_COD2_COD3_tcg2act 0.00252938)
  (COD1_COD2_COD3_tcg2aga 8.76951e-07)
  (COD1_COD2_COD3_tcg2agc 0.13453)
  (COD1_COD2_COD3_tcg2agg 0.00349377)
  (COD1_COD2_COD3_tcg2agt 0.00703127)
  (COD1_COD2_COD3_tcg2ata 2.26184e-07)
  (COD1_COD2_COD3_tcg2atc 9.16776e-06)
  (COD1_COD2_COD3_tcg2atg 0.011164)
  (COD1_COD2_COD3_tcg2att 6.9668e-07)
  (COD1_COD2_COD3_tcg2caa 3.74416e-05)
  (COD1_COD2_COD3_tcg2cac 0.00126557)
  (COD1_COD2_COD3_tcg2cag 0.031511)
  (COD1_COD2_COD3_tcg2cat 0.00078719)
  (COD1_COD2_COD3_tcg2cca 0.00169206)
  (COD1_COD2_COD3_tcg2ccc 0.00450268)
  (COD1_COD2_COD3_tcg2ccg 0.0253302)
  (COD1_COD2_COD3_tcg2cct 0.000821188)
  (COD1_COD2_COD3_tcg2cga 1.52377e-05)
  (COD1_COD2_COD3_tcg2cgc 0.000468391)
  (COD1_COD2_COD3_tcg2cgg 0.0155386)
  (COD1_COD2_COD3_tcg2cgt 1.45059e-05)
  (COD1_COD2_COD3_tcg2cta 5.9278e-07)
  (COD1_COD2_COD3_tcg2ctc 5.58117e-06)
  (COD1_COD2_COD3_tcg2ctg 0.00760941)
  (COD1_COD2_COD3_tcg2ctt 8.40645e-07)
  (COD1_COD2_COD3_tcg2gaa 7.4935e-05)
  (COD1_COD2_COD3_tcg2gac 0.016945)
  (COD1_COD2_COD3_tcg2gag 0.031511)
  (COD1_COD2_COD3_tcg2gat 0.00336308)
  (COD1_COD2_COD3_tcg2gca 0.00835584)
  (COD1_COD2_COD3_tcg2gcc 0.0530685)
  (COD1_COD2_COD3_tcg2gcg 0.0852002)
  (COD1_COD2_COD3_tcg2gct 0.00473966)
  (COD1_COD2_COD3_tcg2gga 2.89315e-05)
  (COD1_COD2_COD3_tcg2ggc 0.00842563)
  (COD1_COD2_COD3_tcg2ggg 0.0221277)
  (COD1_COD2_COD3_tcg2ggt 0.00012039)
  (COD1_COD2_COD3_tcg2gta 1.13412e-06)
  (COD1_COD2_COD3_tcg2gtc 2.2778e-05)
  (COD1_COD2_COD3_tcg2gtg 0.0102304)
  (COD1_COD2_COD3_tcg2gtt 2.54659e-06)
  (COD1_COD2_COD3_tcg2taa 0)
  (COD1_COD2_COD3_tcg2tac 0.00357213)
  (COD1_COD2_COD3_tcg2tag 0)
  (COD1_COD2_COD3_tcg2tat 0.00100332)
  (COD1_COD2_COD3_tcg2tca 0.180742)
  (COD1_COD2_COD3_tcg2tcc 0.380508)
  (COD1_COD2_COD3_tcg2tct 0.135842)
  (COD1_COD2_COD3_tcg2tga 0)
  (COD1_COD2_COD3_tcg2tgc 0.00365731)
  (COD1_COD2_COD3_tcg2tgg 0.00522628)
  (COD1_COD2_COD3_tcg2tgt 0.000924525)
  (COD1_COD2_COD3_tcg2tta 0.000165143)
  (COD1_COD2_COD3_tcg2ttc 0.0042333)
  (COD1_COD2_COD3_tcg2ttg 0.0109116)
  (COD1_COD2_COD3_tcg2ttt 0.000731426)
  (COD1_COD2_COD3_tct2aaa 0.0106058)
  (COD1_COD2_COD3_tct2aac 6.86681e-05)
  (COD1_COD2_COD3_tct2aag 0.00417792)
  (COD1_COD2_COD3_tct2aat 0.0395001)
  (COD1_COD2_COD3_tct2aca 0.0264975)
  (COD1_COD2_COD3_tct2acc 0.00352295)
  (COD1_COD2_COD3_tct2acg 0.00385782)
  (COD1_COD2_COD3_tct2act 0.0594162)
  (COD1_COD2_COD3_tct2aga 0.000466447)
  (COD1_COD2_COD3_tct2agc 0.00129844)
  (COD1_COD2_COD3_tct2agg 0.000434607)
  (COD1_COD2_COD3_tct2agt 0.125521)
  (COD1_COD2_COD3_tct2ata 5.9281e-06)
  (COD1_COD2_COD3_tct2atc 7.23251e-07)
  (COD1_COD2_COD3_tct2atg 0.00198039)
  (COD1_COD2_COD3_tct2att 0.00426866)
  (COD1_COD2_COD3_tct2caa 0.00872881)
  (COD1_COD2_COD3_tct2cac 6.15419e-06)
  (COD1_COD2_COD3_tct2cag 0.00102157)
  (COD1_COD2_COD3_tct2cat 0.0133872)
  (COD1_COD2_COD3_tct2cca 0.00544444)
  (COD1_COD2_COD3_tct2ccc 0.00105393)
  (COD1_COD2_COD3_tct2ccg 0.000262208)
  (COD1_COD2_COD3_tct2cct 0.0296875)
  (COD1_COD2_COD3_tct2cga 2.92541e-05)
  (COD1_COD2_COD3_tct2cgc 2.6381e-07)
  (COD1_COD2_COD3_tct2cgg 5.25429e-07)
  (COD1_COD2_COD3_tct2cgt 0.00560916)
  (COD1_COD2_COD3_tct2cta 5.05801e-06)
  (COD1_COD2_COD3_tct2ctc 1.84238e-06)
  (COD1_COD2_COD3_tct2ctg 4.52682e-07)
  (COD1_COD2_COD3_tct2ctt 0.00421574)
  (COD1_COD2_COD3_tct2gaa 0.0114144)
  (COD1_COD2_COD3_tct2gac 5.38732e-06)
  (COD1_COD2_COD3_tct2gag 0.00129665)
  (COD1_COD2_COD3_tct2gat 0.0320396)
  (COD1_COD2_COD3_tct2gca 0.0319066)
  (COD1_COD2_COD3_tct2gcc 0.00300379)
  (COD1_COD2_COD3_tct2gcg 0.00189052)
  (COD1_COD2_COD3_tct2gct 0.0755722)
  (COD1_COD2_COD3_tct2gga 0.00019913)
  (COD1_COD2_COD3_tct2ggc 1.80697e-06)
  (COD1_COD2_COD3_tct2ggg 2.89515e-05)
  (COD1_COD2_COD3_tct2ggt 0.0164816)
  (COD1_COD2_COD3_tct2gta 1.44758e-05)
  (COD1_COD2_COD3_tct2gtc 8.31373e-07)
  (COD1_COD2_COD3_tct2gtg 5.1894e-06)
  (COD1_COD2_COD3_tct2gtt 0.0107315)
  (COD1_COD2_COD3_tct2taa 0)
  (COD1_COD2_COD3_tct2tac 0.00116295)
  (COD1_COD2_COD3_tct2tag 0)
  (COD1_COD2_COD3_tct2tat 0.0118662)
  (COD1_COD2_COD3_tct2tca 0.372419)
  (COD1_COD2_COD3_tct2tcc 0.254546)
  (COD1_COD2_COD3_tct2tcg 0.100412)
  (COD1_COD2_COD3_tct2tga 0)
  (COD1_COD2_COD3_tct2tgc 0.00102511)
  (COD1_COD2_COD3_tct2tgg 0.00088871)
  (COD1_COD2_COD3_tct2tgt 0.0116704)
  (COD1_COD2_COD3_tct2tta 0.00404681)
  (COD1_COD2_COD3_tct2ttc 0.00112411)
  (COD1_COD2_COD3_tct2ttg 0.00432226)
  (COD1_COD2_COD3_tct2ttt 0.0165732)
  (COD1_COD2_COD3_tga2aaa 0)
  (COD1_COD2_COD3_tga2aac 0)
  (COD1_COD2_COD3_tga2aag 0)
  (COD1_COD2_COD3_tga2aat 0)
  (COD1_COD2_COD3_tga2aca 0)
  (COD1_COD2_COD3_tga2acc 0)
  (COD1_COD2_COD3_tga2acg 0)
  (COD1_COD2_COD3_tga2act 0)
  (COD1_COD2_COD3_tga2aga 0)
  (COD1_COD2_COD3_tga2agc 0)
  (COD1_COD2_COD3_tga2agg 0)
  (COD1_COD2_COD3_tga2agt 0)
  (COD1_COD2_COD3_tga2ata 0)
  (COD1_COD2_COD3_tga2atc 0)
  (COD1_COD2_COD3_tga2atg 0)
  (COD1_COD2_COD3_tga2att 0)
  (COD1_COD2_COD3_tga2caa 0)
  (COD1_COD2_COD3_tga2cac 0)
  (COD1_COD2_COD3_tga2cag 0)
  (COD1_COD2_COD3_tga2cat 0)
  (COD1_COD2_COD3_tga2cca 0)
  (COD1_COD2_COD3_tga2ccc 0)
  (COD1_COD2_COD3_tga2ccg 0)
  (COD1_COD2_COD3_tga2cct 0)
  (COD1_COD2_COD3_tga2cga 0)
  (COD1_COD2_COD3_tga2cgc 0)
  (COD1_COD2_COD3_tga2cgg 0)
  (COD1_COD2_COD3_tga2cgt 0)
  (COD1_COD2_COD3_tga2cta 0)
  (COD1_COD2_COD3_tga2ctc 0)
  (COD1_COD2_COD3_tga2ctg 0)
  (COD1_COD2_COD3_tga2ctt 0)
  (COD1_COD2_COD3_tga2gaa 0)
  (COD1_COD2_COD3_tga2gac 0)
  (COD1_COD2_COD3_tga2gag 0)
  (COD1_COD2_COD3_tga2gat 0)
  (COD1_COD2_COD3_tga2gca 0)
  (COD1_COD2_COD3_tga2gcc 0)
  (COD1_COD2_COD3_tga2gcg 0)
  (COD1_COD2_COD3_tga2gct 0)
  (COD1_COD2_COD3_tga2gga 0)
  (COD1_COD2_COD3_tga2ggc 0)
  (COD1_COD2_COD3_tga2ggg 0)
  (COD1_COD2_COD3_tga2ggt 0)
  (COD1_COD2_COD3_tga2gta 0)
  (COD1_COD2_COD3_tga2gtc 0)
  (COD1_COD2_COD3_tga2gtg 0)
  (COD1_COD2_COD3_tga2gtt 0)
  (COD1_COD2_COD3_tga2taa 0)
  (COD1_COD2_COD3_tga2tac 0)
  (COD1_COD2_COD3_tga2tag 0)
  (COD1_COD2_COD3_tga2tat 0)
  (COD1_COD2_COD3_tga2tca 0)
  (COD1_COD2_COD3_tga2tcc 0)
  (COD1_COD2_COD3_tga2tcg 0)
  (COD1_COD2_COD3_tga2tct 0)
  (COD1_COD2_COD3_tga2tgc 0)
  (COD1_COD2_COD3_tga2tgg 0)
  (COD1_COD2_COD3_tga2tgt 0)
  (COD1_COD2_COD3_tga2tta 0)
  (COD1_COD2_COD3_tga2ttc 0)
  (COD1_COD2_COD3_tga2ttg 0)
  (COD1_COD2_COD3_tga2ttt 0)
  (COD1_COD2_COD3_tgc2aaa 0.000132108)
  (COD1_COD2_COD3_tgc2aac 0.0066152)
  (COD1_COD2_COD3_tgc2aag 0.000930311)
  (COD1_COD2_COD3_tgc2aat 4.00991e-06)
  (COD1_COD2_COD3_tgc2aca 0.000803731)
  (COD1_COD2_COD3_tgc2acc 0.0144179)
  (COD1_COD2_COD3_tgc2acg 0.00185162)
  (COD1_COD2_COD3_tgc2act 1.83609e-05)
  (COD1_COD2_COD3_tgc2aga 0.000300989)
  (COD1_COD2_COD3_tgc2agc 0.0188016)
  (COD1_COD2_COD3_tgc2agg 0.000478896)
  (COD1_COD2_COD3_tgc2agt 0.00187486)
  (COD1_COD2_COD3_tgc2ata 0.000155155)
  (COD1_COD2_COD3_tgc2atc 0.0073604)
  (COD1_COD2_COD3_tgc2atg 0.00687702)
  (COD1_COD2_COD3_tgc2att 9.53722e-06)
  (COD1_COD2_COD3_tgc2caa 0.00055355)
  (COD1_COD2_COD3_tgc2cac 0.00533609)
  (COD1_COD2_COD3_tgc2cag 0.00218978)
  (COD1_COD2_COD3_tgc2cat 1.60285e-05)
  (COD1_COD2_COD3_tgc2cca 0.000114847)
  (COD1_COD2_COD3_tgc2ccc 0.00224666)
  (COD1_COD2_COD3_tgc2ccg 5.6086e-05)
  (COD1_COD2_COD3_tgc2cct 2.96199e-06)
  (COD1_COD2_COD3_tgc2cga 0.000799286)
  (COD1_COD2_COD3_tgc2cgc 0.00755161)
  (COD1_COD2_COD3_tgc2cgg 0.00149463)
  (COD1_COD2_COD3_tgc2cgt 0.00066057)
  (COD1_COD2_COD3_tgc2cta 0.000159186)
  (COD1_COD2_COD3_tgc2ctc 0.0113277)
  (COD1_COD2_COD3_tgc2ctg 0.00757783)
  (COD1_COD2_COD3_tgc2ctt 9.27642e-06)
  (COD1_COD2_COD3_tgc2gaa 0.000127607)
  (COD1_COD2_COD3_tgc2gac 0.00181225)
  (COD1_COD2_COD3_tgc2gag 0.000402981)
  (COD1_COD2_COD3_tgc2gat 3.74397e-06)
  (COD1_COD2_COD3_tgc2gca 0.00491361)
  (COD1_COD2_COD3_tgc2gcc 0.0406106)
  (COD1_COD2_COD3_tgc2gcg 0.0107241)
  (COD1_COD2_COD3_tgc2gct 0.000222949)
  (COD1_COD2_COD3_tgc2gga 0.0010082)
  (COD1_COD2_COD3_tgc2ggc 0.0073197)
  (COD1_COD2_COD3_tgc2ggg 0.00188007)
  (COD1_COD2_COD3_tgc2ggt 0.000760376)
  (COD1_COD2_COD3_tgc2gta 0.000949858)
  (COD1_COD2_COD3_tgc2gtc 0.0234706)
  (COD1_COD2_COD3_tgc2gtg 0.0119985)
  (COD1_COD2_COD3_tgc2gtt 8.14345e-05)
  (COD1_COD2_COD3_tgc2taa 0)
  (COD1_COD2_COD3_tgc2tac 0.0104961)
  (COD1_COD2_COD3_tgc2tag 0)
  (COD1_COD2_COD3_tgc2tat 0.00117836)
  (COD1_COD2_COD3_tgc2tca 0.00126381)
  (COD1_COD2_COD3_tgc2tcc 0.014249)
  (COD1_COD2_COD3_tgc2tcg 0.00430133)
  (COD1_COD2_COD3_tgc2tct 0.00163103)
  (COD1_COD2_COD3_tgc2tga 0)
  (COD1_COD2_COD3_tgc2tgg 0.00267917)
  (COD1_COD2_COD3_tgc2tgt 0.366275)
  (COD1_COD2_COD3_tgc2tta 0.000648323)
  (COD1_COD2_COD3_tgc2ttc 0.0119902)
  (COD1_COD2_COD3_tgc2ttg 0.00378366)
  (COD1_COD2_COD3_tgc2ttt 0.000984719)
  (COD1_COD2_COD3_tgg2aaa 0.00158533)
  (COD1_COD2_COD3_tgg2aac 0.00109489)
  (COD1_COD2_COD3_tgg2aag 0.00281042)
  (COD1_COD2_COD3_tgg2aat 0.00166761)
  (COD1_COD2_COD3_tgg2aca 0.000793215)
  (COD1_COD2_COD3_tgg2acc 0.000676323)
  (COD1_COD2_COD3_tgg2acg 0.00387658)
  (COD1_COD2_COD3_tgg2act 0.000455267)
  (COD1_COD2_COD3_tgg2aga 0.00141596)
  (COD1_COD2_COD3_tgg2agc 0.00229069)
  (COD1_COD2_COD3_tgg2agg 0.00337242)
  (COD1_COD2_COD3_tgg2agt 0.00143374)
  (COD1_COD2_COD3_tgg2ata 0.00217918)
  (COD1_COD2_COD3_tgg2atc 0.00325303)
  (COD1_COD2_COD3_tgg2atg 0.00681533)
  (COD1_COD2_COD3_tgg2att 0.00193963)
  (COD1_COD2_COD3_tgg2caa 0.000947228)
  (COD1_COD2_COD3_tgg2cac 0.00407778)
  (COD1_COD2_COD3_tgg2cag 0.00513294)
  (COD1_COD2_COD3_tgg2cat 0.00208173)
  (COD1_COD2_COD3_tgg2cca 0.000785555)
  (COD1_COD2_COD3_tgg2ccc 0.0003391)
  (COD1_COD2_COD3_tgg2ccg 0.00263314)
  (COD1_COD2_COD3_tgg2cct 0.000644738)
  (COD1_COD2_COD3_tgg2cga 0.000704555)
  (COD1_COD2_COD3_tgg2cgc 0.00257716)
  (COD1_COD2_COD3_tgg2cgg 0.00927138)
  (COD1_COD2_COD3_tgg2cgt 0.0001671)
  (COD1_COD2_COD3_tgg2cta 0.000125763)
  (COD1_COD2_COD3_tgg2ctc 0.00191028)
  (COD1_COD2_COD3_tgg2ctg 0.0143879)
  (COD1_COD2_COD3_tgg2ctt 0.000153338)
  (COD1_COD2_COD3_tgg2gaa 0.000344789)
  (COD1_COD2_COD3_tgg2gac 0.00167921)
  (COD1_COD2_COD3_tgg2gag 0.00421281)
  (COD1_COD2_COD3_tgg2gat 0.00080876)
  (COD1_COD2_COD3_tgg2gca 0.000519712)
  (COD1_COD2_COD3_tgg2gcc 0.00118492)
  (COD1_COD2_COD3_tgg2gcg 0.00816122)
  (COD1_COD2_COD3_tgg2gct 0.000677731)
  (COD1_COD2_COD3_tgg2gga 0.00120814)
  (COD1_COD2_COD3_tgg2ggc 0.0024911)
  (COD1_COD2_COD3_tgg2ggg 0.00390354)
  (COD1_COD2_COD3_tgg2ggt 0.000663323)
  (COD1_COD2_COD3_tgg2gta 0.000639841)
  (COD1_COD2_COD3_tgg2gtc 0.00169676)
  (COD1_COD2_COD3_tgg2gtg 0.00627139)
  (COD1_COD2_COD3_tgg2gtt 0.000737535)
  (COD1_COD2_COD3_tgg2taa 0)
  (COD1_COD2_COD3_tgg2tac 0.0237488)
  (COD1_COD2_COD3_tgg2tag 0)
  (COD1_COD2_COD3_tgg2tat 0.0196545)
  (COD1_COD2_COD3_tgg2tca 0.00203043)
  (COD1_COD2_COD3_tgg2tcc 0.000875867)
  (COD1_COD2_COD3_tgg2tcg 0.00357708)
  (COD1_COD2_COD3_tgg2tct 0.000822897)
  (COD1_COD2_COD3_tgg2tga 0)
  (COD1_COD2_COD3_tgg2tgc 0.00155918)
  (COD1_COD2_COD3_tgg2tgt 0.00170501)
  (COD1_COD2_COD3_tgg2tta 0.00439171)
  (COD1_COD2_COD3_tgg2ttc 0.0252775)
  (COD1_COD2_COD3_tgg2ttg 0.00995061)
  (COD1_COD2_COD3_tgg2ttt 0.0168513)
  (COD1_COD2_COD3_tgt2aaa 0.00309467)
  (COD1_COD2_COD3_tgt2aac 1.77232e-05)
  (COD1_COD2_COD3_tgt2aag 0.000307527)
  (COD1_COD2_COD3_tgt2aat 0.00802101)
  (COD1_COD2_COD3_tgt2aca 0.00545578)
  (COD1_COD2_COD3_tgt2acc 1.31279e-05)
  (COD1_COD2_COD3_tgt2acg 0.000576257)
  (COD1_COD2_COD3_tgt2act 0.0137922)
  (COD1_COD2_COD3_tgt2aga 0.00140716)
  (COD1_COD2_COD3_tgt2agc 0.00340059)
  (COD1_COD2_COD3_tgt2agg 0.000830346)
  (COD1_COD2_COD3_tgt2agt 0.0207032)
  (COD1_COD2_COD3_tgt2ata 0.000892414)
  (COD1_COD2_COD3_tgt2atc 6.20129e-06)
  (COD1_COD2_COD3_tgt2atg 0.00631939)
  (COD1_COD2_COD3_tgt2att 0.010687)
  (COD1_COD2_COD3_tgt2caa 0.00280458)
  (COD1_COD2_COD3_tgt2cac 2.43609e-06)
  (COD1_COD2_COD3_tgt2cag 0.00119979)
  (COD1_COD2_COD3_tgt2cat 0.00676826)
  (COD1_COD2_COD3_tgt2cca 3.0412e-05)
  (COD1_COD2_COD3_tgt2ccc 5.05075e-07)
  (COD1_COD2_COD3_tgt2ccg 1.53748e-06)
  (COD1_COD2_COD3_tgt2cct 0.00249975)
  (COD1_COD2_COD3_tgt2cga 0.00112022)
  (COD1_COD2_COD3_tgt2cgc 0.000189305)
  (COD1_COD2_COD3_tgt2cgg 0.000195574)
  (COD1_COD2_COD3_tgt2cgt 0.00617648)
  (COD1_COD2_COD3_tgt2cta 0.000152279)
  (COD1_COD2_COD3_tgt2ctc 7.91489e-06)
  (COD1_COD2_COD3_tgt2ctg 7.40059e-05)
  (COD1_COD2_COD3_tgt2ctt 0.0082408)
  (COD1_COD2_COD3_tgt2gaa 0.000998465)
  (COD1_COD2_COD3_tgt2gac 1.74049e-07)
  (COD1_COD2_COD3_tgt2gag 0.000441591)
  (COD1_COD2_COD3_tgt2gat 0.00286153)
  (COD1_COD2_COD3_tgt2gca 0.0133224)
  (COD1_COD2_COD3_tgt2gcc 1.31279e-05)
  (COD1_COD2_COD3_tgt2gcg 0.00052369)
  (COD1_COD2_COD3_tgt2gct 0.0359466)
  (COD1_COD2_COD3_tgt2gga 0.00323953)
  (COD1_COD2_COD3_tgt2ggc 0.000178969)
  (COD1_COD2_COD3_tgt2ggg 0.00108281)
  (COD1_COD2_COD3_tgt2ggt 0.00710968)
  (COD1_COD2_COD3_tgt2gta 0.0082408)
  (COD1_COD2_COD3_tgt2gtc 2.096e-05)
  (COD1_COD2_COD3_tgt2gtg 0.00287146)
  (COD1_COD2_COD3_tgt2gtt 0.0304165)
  (COD1_COD2_COD3_tgt2taa 0)
  (COD1_COD2_COD3_tgt2tac 0.000916232)
  (COD1_COD2_COD3_tgt2tag 0)
  (COD1_COD2_COD3_tgt2tat 0.0141898)
  (COD1_COD2_COD3_tgt2tca 0.00779627)
  (COD1_COD2_COD3_tgt2tcc 0.0036981)
  (COD1_COD2_COD3_tgt2tcg 0.00125161)
  (COD1_COD2_COD3_tgt2tct 0.0213888)
  (COD1_COD2_COD3_tgt2tga 0)
  (COD1_COD2_COD3_tgt2tgc 0.421616)
  (COD1_COD2_COD3_tgt2tgg 0.00337242)
  (COD1_COD2_COD3_tgt2tta 0.0104598)
  (COD1_COD2_COD3_tgt2ttc 0.00164238)
  (COD1_COD2_COD3_tgt2ttg 0.00622375)
  (COD1_COD2_COD3_tgt2ttt 0.0157119)
  (COD1_COD2_COD3_tta2aaa 0.0190509)
  (COD1_COD2_COD3_tta2aac 5.73042e-05)
  (COD1_COD2_COD3_tta2aag 1.16439e-06)
  (COD1_COD2_COD3_tta2aat 0.00605356)
  (COD1_COD2_COD3_tta2aca 0.0114382)
  (COD1_COD2_COD3_tta2acc 8.08639e-07)
  (COD1_COD2_COD3_tta2acg 6.01055e-07)
  (COD1_COD2_COD3_tta2act 0.000158306)
  (COD1_COD2_COD3_tta2aga 0.00413184)
  (COD1_COD2_COD3_tta2agc 5.74604e-06)
  (COD1_COD2_COD3_tta2agg 1.5048e-06)
  (COD1_COD2_COD3_tta2agt 0.000112796)
  (COD1_COD2_COD3_tta2ata 0.0661553)
  (COD1_COD2_COD3_tta2atc 0.00100124)
  (COD1_COD2_COD3_tta2atg 0.0300602)
  (COD1_COD2_COD3_tta2att 0.0527751)
  (COD1_COD2_COD3_tta2caa 0.00990244)
  (COD1_COD2_COD3_tta2cac 1.11928e-06)
  (COD1_COD2_COD3_tta2cag 2.0399e-07)
  (COD1_COD2_COD3_tta2cat 0.000283375)
  (COD1_COD2_COD3_tta2cca 0.00291558)
  (COD1_COD2_COD3_tta2ccc 9.45728e-08)
  (COD1_COD2_COD3_tta2ccg 2.55341e-08)
  (COD1_COD2_COD3_tta2cct 1.3915e-05)
  (COD1_COD2_COD3_tta2cga 0.000657373)
  (COD1_COD2_COD3_tta2cgc 1.22385e-08)
  (COD1_COD2_COD3_tta2cgg 1.31078e-08)
  (COD1_COD2_COD3_tta2cgt 6.2748e-06)
  (COD1_COD2_COD3_tta2cta 0.200406)
  (COD1_COD2_COD3_tta2ctc 0.0230834)
  (COD1_COD2_COD3_tta2ctg 0.0298112)
  (COD1_COD2_COD3_tta2ctt 0.182631)
  (COD1_COD2_COD3_tta2gaa 0.00992305)
  (COD1_COD2_COD3_tta2gac 1.19795e-06)
  (COD1_COD2_COD3_tta2gag 2.58022e-07)
  (COD1_COD2_COD3_tta2gat 0.00120563)
  (COD1_COD2_COD3_tta2gca 0.0172053)
  (COD1_COD2_COD3_tta2gcc 4.3878e-07)
  (COD1_COD2_COD3_tta2gcg 1.83591e-07)
  (COD1_COD2_COD3_tta2gct 0.000168262)
  (COD1_COD2_COD3_tta2gga 0.00329616)
  (COD1_COD2_COD3_tta2ggc 4.51406e-08)
  (COD1_COD2_COD3_tta2ggg 1.98412e-07)
  (COD1_COD2_COD3_tta2ggt 1.23682e-05)
  (COD1_COD2_COD3_tta2gta 0.0251203)
  (COD1_COD2_COD3_tta2gtc 0.000110321)
  (COD1_COD2_COD3_tta2gtg 0.000559335)
  (COD1_COD2_COD3_tta2gtt 0.0142391)
  (COD1_COD2_COD3_tta2taa 0)
  (COD1_COD2_COD3_tta2tac 0.000559723)
  (COD1_COD2_COD3_tta2tag 0)
  (COD1_COD2_COD3_tta2tat 0.0137731)
  (COD1_COD2_COD3_tta2tca 0.0122846)
  (COD1_COD2_COD3_tta2tcc 0.000124981)
  (COD1_COD2_COD3_tta2tcg 8.89282e-05)
  (COD1_COD2_COD3_tta2tct 0.0029481)
  (COD1_COD2_COD3_tta2tga 0)
  (COD1_COD2_COD3_tta2tgc 0.000296846)
  (COD1_COD2_COD3_tta2tgg 0.00345524)
  (COD1_COD2_COD3_tta2tgt 0.00416058)
  (COD1_COD2_COD3_tta2ttc 0.00324402)
  (COD1_COD2_COD3_tta2ttg 0.291587)
  (COD1_COD2_COD3_tta2ttt 0.0513319)
  (COD1_COD2_COD3_ttc2aaa 0.000187087)
  (COD1_COD2_COD3_ttc2aac 0.00291963)
  (COD1_COD2_COD3_ttc2aag 0.00116537)
  (COD1_COD2_COD3_ttc2aat 2.53778e-06)
  (COD1_COD2_COD3_ttc2aca 0.000138196)
  (COD1_COD2_COD3_ttc2acc 0.00664737)
  (COD1_COD2_COD3_ttc2acg 0.000389794)
  (COD1_COD2_COD3_ttc2act 1.72138e-06)
  (COD1_COD2_COD3_ttc2aga 0.000122664)
  (COD1_COD2_COD3_ttc2agc 0.00228435)
  (COD1_COD2_COD3_ttc2agg 0.000385495)
  (COD1_COD2_COD3_ttc2agt 4.88208e-07)
  (COD1_COD2_COD3_ttc2ata 0.000832651)
  (COD1_COD2_COD3_ttc2atc 0.023914)
  (COD1_COD2_COD3_ttc2atg 0.0169685)
  (COD1_COD2_COD3_ttc2att 0.00148533)
  (COD1_COD2_COD3_ttc2caa 0.000285148)
  (COD1_COD2_COD3_ttc2cac 0.00841396)
  (COD1_COD2_COD3_ttc2cag 0.00222343)
  (COD1_COD2_COD3_ttc2cat 1.40604e-05)
  (COD1_COD2_COD3_ttc2cca 7.34438e-05)
  (COD1_COD2_COD3_ttc2ccc 0.00297478)
  (COD1_COD2_COD3_ttc2ccg 0.000966462)
  (COD1_COD2_COD3_ttc2cct 1.59612e-06)
  (COD1_COD2_COD3_ttc2cga 7.70952e-05)
  (COD1_COD2_COD3_ttc2cgc 0.00293383)
  (COD1_COD2_COD3_ttc2cgg 0.000223258)
  (COD1_COD2_COD3_ttc2cgt 8.91662e-07)
  (COD1_COD2_COD3_ttc2cta 0.00146184)
  (COD1_COD2_COD3_ttc2ctc 0.0387409)
  (COD1_COD2_COD3_ttc2ctg 0.038234)
  (COD1_COD2_COD3_ttc2ctt 0.00315094)
  (COD1_COD2_COD3_ttc2gaa 0.00022264)
  (COD1_COD2_COD3_ttc2gac 0.00140035)
  (COD1_COD2_COD3_ttc2gag 0.00105393)
  (COD1_COD2_COD3_ttc2gat 6.10291e-07)
  (COD1_COD2_COD3_ttc2gca 0.000272964)
  (COD1_COD2_COD3_ttc2gcc 0.0137445)
  (COD1_COD2_COD3_ttc2gcg 0.00136963)
  (COD1_COD2_COD3_ttc2gct 4.21218e-06)
  (COD1_COD2_COD3_ttc2gga 0.000300364)
  (COD1_COD2_COD3_ttc2ggc 0.00409194)
  (COD1_COD2_COD3_ttc2ggg 0.000327776)
  (COD1_COD2_COD3_ttc2ggt 2.30628e-06)
  (COD1_COD2_COD3_ttc2gta 0.000517914)
  (COD1_COD2_COD3_ttc2gtc 0.0174334)
  (COD1_COD2_COD3_ttc2gtg 0.00839648)
  (COD1_COD2_COD3_ttc2gtt 0.00125421)
  (COD1_COD2_COD3_ttc2taa 0)
  (COD1_COD2_COD3_ttc2tac 0.0748943)
  (COD1_COD2_COD3_ttc2tag 0)
  (COD1_COD2_COD3_ttc2tat 0.00868054)
  (COD1_COD2_COD3_ttc2tca 0.000587958)
  (COD1_COD2_COD3_ttc2tcc 0.00706057)
  (COD1_COD2_COD3_ttc2tcg 0.00173001)
  (COD1_COD2_COD3_ttc2tct 0.000621051)
  (COD1_COD2_COD3_ttc2tga 0)
  (COD1_COD2_COD3_ttc2tgc 0.00416635)
  (COD1_COD2_COD3_ttc2tgg 0.0150823)
  (COD1_COD2_COD3_ttc2tgt 0.000495441)
  (COD1_COD2_COD3_ttc2tta 0.00246021)
  (COD1_COD2_COD3_ttc2ttg 0.0106205)
  (COD1_COD2_COD3_ttc2ttt 0.342933)
  (COD1_COD2_COD3_ttg2aaa 0.000155155)
  (COD1_COD2_COD3_ttg2aac 0.00123438)
  (COD1_COD2_COD3_ttg2aag 0.00891222)
  (COD1_COD2_COD3_ttg2aat 0.00173482)
  (COD1_COD2_COD3_ttg2aca 0.000146075)
  (COD1_COD2_COD3_ttg2acc 5.29504e-05)
  (COD1_COD2_COD3_ttg2acg 0.00627139)
  (COD1_COD2_COD3_ttg2act 0.000197892)
  (COD1_COD2_COD3_ttg2aga 1.65478e-05)
  (COD1_COD2_COD3_ttg2agc 7.91528e-05)
  (COD1_COD2_COD3_ttg2agg 0.0034409)
  (COD1_COD2_COD3_ttg2agt 9.55755e-05)
  (COD1_COD2_COD3_ttg2ata 0.013659)
  (COD1_COD2_COD3_ttg2atc 0.0113592)
  (COD1_COD2_COD3_ttg2atg 0.0648837)
  (COD1_COD2_COD3_ttg2att 0.0320174)
  (COD1_COD2_COD3_ttg2caa 0.000128584)
  (COD1_COD2_COD3_ttg2cac 2.39934e-05)
  (COD1_COD2_COD3_ttg2cag 0.00520098)
  (COD1_COD2_COD3_ttg2cat 0.000429516)
  (COD1_COD2_COD3_ttg2cca 6.2059e-05)
  (COD1_COD2_COD3_ttg2ccc 3.88676e-06)
  (COD1_COD2_COD3_ttg2ccg 0.00137915)
  (COD1_COD2_COD3_ttg2cct 3.65441e-05)
  (COD1_COD2_COD3_ttg2cga 7.46721e-06)
  (COD1_COD2_COD3_ttg2cgc 1.22567e-06)
  (COD1_COD2_COD3_ttg2cgg 0.00104231)
  (COD1_COD2_COD3_ttg2cgt 2.58944e-05)
  (COD1_COD2_COD3_ttg2cta 0.121919)
  (COD1_COD2_COD3_ttg2ctc 0.0949953)
  (COD1_COD2_COD3_ttg2ctg 0.355027)
  (COD1_COD2_COD3_ttg2ctt 0.169341)
  (COD1_COD2_COD3_ttg2gaa 0.000189174)
  (COD1_COD2_COD3_ttg2gac 0.00017156)
  (COD1_COD2_COD3_ttg2gag 0.00489661)
  (COD1_COD2_COD3_ttg2gat 0.0011025)
  (COD1_COD2_COD3_ttg2gca 0.000320142)
  (COD1_COD2_COD3_ttg2gcc 4.46185e-05)
  (COD1_COD2_COD3_ttg2gcg 0.0040412)
  (COD1_COD2_COD3_ttg2gct 0.000673983)
  (COD1_COD2_COD3_ttg2gga 1.77601e-05)
  (COD1_COD2_COD3_ttg2ggc 5.15713e-06)
  (COD1_COD2_COD3_ttg2ggg 0.00179849)
  (COD1_COD2_COD3_ttg2ggt 6.54159e-05)
  (COD1_COD2_COD3_ttg2gta 0.00832693)
  (COD1_COD2_COD3_ttg2gtc 0.00259509)
  (COD1_COD2_COD3_ttg2gtg 0.0251029)
  (COD1_COD2_COD3_ttg2gtt 0.0100755)
  (COD1_COD2_COD3_ttg2taa 0)
  (COD1_COD2_COD3_ttg2tac 0.00297066)
  (COD1_COD2_COD3_ttg2tag 0)
  (COD1_COD2_COD3_ttg2tat 0.00581502)
  (COD1_COD2_COD3_ttg2tca 0.00634572)
  (COD1_COD2_COD3_ttg2tcc 0.00170975)
  (COD1_COD2_COD3_ttg2tcg 0.00605356)
  (COD1_COD2_COD3_ttg2tct 0.00324627)
  (COD1_COD2_COD3_ttg2tga 0)
  (COD1_COD2_COD3_ttg2tgc 0.00178483)
  (COD1_COD2_COD3_ttg2tgg 0.00807121)
  (COD1_COD2_COD3_ttg2tgt 0.00255051)
  (COD1_COD2_COD3_ttg2tta 0.300409)
  (COD1_COD2_COD3_ttg2ttc 0.0144278)
  (COD1_COD2_COD3_ttg2ttt 0.0257908)
  (COD1_COD2_COD3_ttt2aaa 0.00461006)
  (COD1_COD2_COD3_ttt2aac 1.60388e-06)
  (COD1_COD2_COD3_ttt2aag 0.000349844)
  (COD1_COD2_COD3_ttt2aat 0.00587172)
  (COD1_COD2_COD3_ttt2aca 0.00104956)
  (COD1_COD2_COD3_ttt2acc 1.38662e-06)
  (COD1_COD2_COD3_ttt2acg 8.60181e-05)
  (COD1_COD2_COD3_ttt2act 0.00629752)
  (COD1_COD2_COD3_ttt2aga 0.00136963)
  (COD1_COD2_COD3_ttt2agc 1.42264e-06)
  (COD1_COD2_COD3_ttt2agg 0.000565965)
  (COD1_COD2_COD3_ttt2agt 0.00232589)
  (COD1_COD2_COD3_ttt2ata 0.0137827)
  (COD1_COD2_COD3_ttt2atc 0.00133218)
  (COD1_COD2_COD3_ttt2atg 0.0143979)
  (COD1_COD2_COD3_ttt2att 0.0277186)
  (COD1_COD2_COD3_ttt2caa 0.00217013)
  (COD1_COD2_COD3_ttt2cac 5.044e-06)
  (COD1_COD2_COD3_ttt2cag 0.000202892)
  (COD1_COD2_COD3_ttt2cat 0.00682006)
  (COD1_COD2_COD3_ttt2cca 0.000281808)
  (COD1_COD2_COD3_ttt2ccc 1.3847e-06)
  (COD1_COD2_COD3_ttt2ccg 4.96791e-05)
  (COD1_COD2_COD3_ttt2cct 0.00338178)
  (COD1_COD2_COD3_ttt2cga 0.000144865)
  (COD1_COD2_COD3_ttt2cgc 1.67655e-07)
  (COD1_COD2_COD3_ttt2cgg 7.48794e-06)
  (COD1_COD2_COD3_ttt2cgt 0.00122161)
  (COD1_COD2_COD3_ttt2cta 0.00763054)
  (COD1_COD2_COD3_ttt2ctc 0.00140618)
  (COD1_COD2_COD3_ttt2ctg 0.00209621)
  (COD1_COD2_COD3_ttt2ctt 0.0305857)
  (COD1_COD2_COD3_ttt2gaa 0.00293383)
  (COD1_COD2_COD3_ttt2gac 2.59997e-07)
  (COD1_COD2_COD3_ttt2gag 0.00033536)
  (COD1_COD2_COD3_ttt2gat 0.00184138)
  (COD1_COD2_COD3_ttt2gca 0.00181728)
  (COD1_COD2_COD3_ttt2gcc 2.07854e-06)
  (COD1_COD2_COD3_ttt2gcg 0.000107305)
  (COD1_COD2_COD3_ttt2gct 0.00914374)
  (COD1_COD2_COD3_ttt2gga 0.00186838)
  (COD1_COD2_COD3_ttt2ggc 2.24155e-07)
  (COD1_COD2_COD3_ttt2ggg 0.000191815)
  (COD1_COD2_COD3_ttt2ggt 0.00278135)
  (COD1_COD2_COD3_ttt2gta 0.00618076)
  (COD1_COD2_COD3_ttt2gtc 0.000485245)
  (COD1_COD2_COD3_ttt2gtg 0.00156026)
  (COD1_COD2_COD3_ttt2gtt 0.016436)
  (COD1_COD2_COD3_ttt2taa 0)
  (COD1_COD2_COD3_ttt2tac 0.00803214)
  (COD1_COD2_COD3_ttt2tag 0)
  (COD1_COD2_COD3_ttt2tat 0.0744802)
  (COD1_COD2_COD3_ttt2tca 0.00288343)
  (COD1_COD2_COD3_ttt2tcc 0.000323712)
  (COD1_COD2_COD3_ttt2tcg 0.000297876)
  (COD1_COD2_COD3_ttt2tct 0.0091374)
  (COD1_COD2_COD3_ttt2tga 0)
  (COD1_COD2_COD3_ttt2tgc 0.000340986)
  (COD1_COD2_COD3_ttt2tgg 0.0100337)
  (COD1_COD2_COD3_ttt2tgt 0.00472654)
  (COD1_COD2_COD3_ttt2tta 0.0388215)
  (COD1_COD2_COD3_ttt2ttc 0.341984)
  (COD1_COD2_COD3_ttt2ttg 0.0189324)

  (COD1_COD2_COD3_aaa2gapgapgap 1)
  (COD1_COD2_COD3_aac2gapgapgap 1)
  (COD1_COD2_COD3_aag2gapgapgap 1)
  (COD1_COD2_COD3_aat2gapgapgap 1)
  (COD1_COD2_COD3_aca2gapgapgap 1)
  (COD1_COD2_COD3_acc2gapgapgap 1)
  (COD1_COD2_COD3_acg2gapgapgap 1)
  (COD1_COD2_COD3_act2gapgapgap 1)
  (COD1_COD2_COD3_aga2gapgapgap 1)
  (COD1_COD2_COD3_agc2gapgapgap 1)
  (COD1_COD2_COD3_agg2gapgapgap 1)
  (COD1_COD2_COD3_agt2gapgapgap 1)
  (COD1_COD2_COD3_ata2gapgapgap 1)
  (COD1_COD2_COD3_atc2gapgapgap 1)
  (COD1_COD2_COD3_atg2gapgapgap 1)
  (COD1_COD2_COD3_att2gapgapgap 1)
  (COD1_COD2_COD3_caa2gapgapgap 1)
  (COD1_COD2_COD3_cac2gapgapgap 1)
  (COD1_COD2_COD3_cag2gapgapgap 1)
  (COD1_COD2_COD3_cat2gapgapgap 1)
  (COD1_COD2_COD3_cca2gapgapgap 1)
  (COD1_COD2_COD3_ccc2gapgapgap 1)
  (COD1_COD2_COD3_ccg2gapgapgap 1)
  (COD1_COD2_COD3_cct2gapgapgap 1)
  (COD1_COD2_COD3_cga2gapgapgap 1)
  (COD1_COD2_COD3_cgc2gapgapgap 1)
  (COD1_COD2_COD3_cgg2gapgapgap 1)
  (COD1_COD2_COD3_cgt2gapgapgap 1)
  (COD1_COD2_COD3_cta2gapgapgap 1)
  (COD1_COD2_COD3_ctc2gapgapgap 1)
  (COD1_COD2_COD3_ctg2gapgapgap 1)
  (COD1_COD2_COD3_ctt2gapgapgap 1)
  (COD1_COD2_COD3_gaa2gapgapgap 1)
  (COD1_COD2_COD3_gac2gapgapgap 1)
  (COD1_COD2_COD3_gag2gapgapgap 1)
  (COD1_COD2_COD3_gat2gapgapgap 1)
  (COD1_COD2_COD3_gca2gapgapgap 1)
  (COD1_COD2_COD3_gcc2gapgapgap 1)
  (COD1_COD2_COD3_gcg2gapgapgap 1)
  (COD1_COD2_COD3_gct2gapgapgap 1)
  (COD1_COD2_COD3_gga2gapgapgap 1)
  (COD1_COD2_COD3_ggc2gapgapgap 1)
  (COD1_COD2_COD3_ggg2gapgapgap 1)
  (COD1_COD2_COD3_ggt2gapgapgap 1)
  (COD1_COD2_COD3_gta2gapgapgap 1)
  (COD1_COD2_COD3_gtc2gapgapgap 1)
  (COD1_COD2_COD3_gtg2gapgapgap 1)
  (COD1_COD2_COD3_gtt2gapgapgap 1)
  (COD1_COD2_COD3_taa2gapgapgap 1)
  (COD1_COD2_COD3_tac2gapgapgap 1)
  (COD1_COD2_COD3_tag2gapgapgap 1)
  (COD1_COD2_COD3_tat2gapgapgap 1)
  (COD1_COD2_COD3_tca2gapgapgap 1)
  (COD1_COD2_COD3_tcc2gapgapgap 1)
  (COD1_COD2_COD3_tcg2gapgapgap 1)
  (COD1_COD2_COD3_tct2gapgapgap 1)
  (COD1_COD2_COD3_tga2gapgapgap 1)
  (COD1_COD2_COD3_tgc2gapgapgap 1)
  (COD1_COD2_COD3_tgg2gapgapgap 1)
  (COD1_COD2_COD3_tgt2gapgapgap 1)
  (COD1_COD2_COD3_tta2gapgapgap 1)
  (COD1_COD2_COD3_ttc2gapgapgap 1)
  (COD1_COD2_COD3_ttg2gapgapgap 1)
  (COD1_COD2_COD3_ttt2gapgapgap 1)

  )  ;; end const-rate


;;;;;;;;;;;;;;;;
;; PSEUDOGENE ;;
;;;;;;;;;;;;;;;;

 ;; constant probability parameters
 (const-pgroup
  ((PG1_PG2_PG3_aaa 0.0310987)
   (PG1_PG2_PG3_aac 0.0203335)
   (PG1_PG2_PG3_aag 0.0267003)
   (PG1_PG2_PG3_aat 0.0222817)
   (PG1_PG2_PG3_aca 0.0131208)
   (PG1_PG2_PG3_acc 0.0178863)
   (PG1_PG2_PG3_acg 0.010687)
   (PG1_PG2_PG3_act 0.0126563)
   (PG1_PG2_PG3_aga 0.00975258)
   (PG1_PG2_PG3_agc 0.0137065)
   (PG1_PG2_PG3_agg 0.00679175)
   (PG1_PG2_PG3_agt 0.0103158)
   (PG1_PG2_PG3_ata 0.0128152)
   (PG1_PG2_PG3_atc 0.0237653)
   (PG1_PG2_PG3_atg 0.0211823)
   (PG1_PG2_PG3_att 0.0247745)
   (PG1_PG2_PG3_caa 0.0171696)
   (PG1_PG2_PG3_cac 0.0110409)
   (PG1_PG2_PG3_cag 0.020732)
   (PG1_PG2_PG3_cat 0.0106722)
   (PG1_PG2_PG3_cca 0.0111718)
   (PG1_PG2_PG3_ccc 0.0104092)
   (PG1_PG2_PG3_ccg 0.0121998)
   (PG1_PG2_PG3_cct 0.0104308)
   (PG1_PG2_PG3_cga 0.00480916)
   (PG1_PG2_PG3_cgc 0.014609)
   (PG1_PG2_PG3_cgg 0.00816122)
   (PG1_PG2_PG3_cgt 0.008496)
   (PG1_PG2_PG3_cta 0.0073604)
   (PG1_PG2_PG3_ctc 0.0168046)
   (PG1_PG2_PG3_ctg 0.0284979)
   (PG1_PG2_PG3_ctt 0.0151767)
   (PG1_PG2_PG3_gaa 0.0345301)
   (PG1_PG2_PG3_gac 0.0252951)
   (PG1_PG2_PG3_gag 0.0306069)
   (PG1_PG2_PG3_gat 0.0285968)
   (PG1_PG2_PG3_gca 0.0165274)
   (PG1_PG2_PG3_gcc 0.0268302)
   (PG1_PG2_PG3_gcg 0.0183002)
   (PG1_PG2_PG3_gct 0.0189193)
   (PG1_PG2_PG3_gga 0.0163905)
   (PG1_PG2_PG3_ggc 0.0236667)
   (PG1_PG2_PG3_ggg 0.0102233)
   (PG1_PG2_PG3_ggt 0.0168863)
   (PG1_PG2_PG3_gta 0.0109267)
   (PG1_PG2_PG3_gtc 0.0184274)
   (PG1_PG2_PG3_gtg 0.0226397)
   (PG1_PG2_PG3_gtt 0.0190245)
   (PG1_PG2_PG3_taa 0)
   (PG1_PG2_PG3_tac 0.0167001)
   (PG1_PG2_PG3_tag 0)
   (PG1_PG2_PG3_tat 0.0172412)
   (PG1_PG2_PG3_tca 0.0103732)
   (PG1_PG2_PG3_tcc 0.0107613)
   (PG1_PG2_PG3_tcg 0.00872276)
   (PG1_PG2_PG3_tct 0.0118006)
   (PG1_PG2_PG3_tga 0)
   (PG1_PG2_PG3_tgc 0.00741674)
   (PG1_PG2_PG3_tgg 0.0127443)
   (PG1_PG2_PG3_tgt 0.00644323)
   (PG1_PG2_PG3_tta 0.0161985)
   (PG1_PG2_PG3_ttc 0.0213592)
   (PG1_PG2_PG3_ttg 0.0157228)
   (PG1_PG2_PG3_ttt 0.0214185))
  )  ;; end const-pgroup


;;;;;;;;;;;
;; NCRNA ;;
;;;;;;;;;;;

 ;; constant probability parameters
 (const-pgroup

   ;; transition probabilities

   ;; Pfold sub-model
   ;; (these are "super-normalized", but it shouldn't pose a practical problem)
   ((Trans_S2L 0.131488)
    (Trans_S2B 0.868742)
   )
   ((Trans_F2F 0.787854)
    (Trans_F2B 0.212421)
   )
   ((Trans_L2F 0.105404)
    (Trans_L2U 0.895025)
   )

   ;; equilibria for Markov chains
   ;; gap parameters
  ((initGap 5.1347e-10)
   (~initGap 1))

   ;; stems (LNUC RNUC)
  ((LNUC_RNUC_aa 0.00345524)
   (LNUC_RNUC_ac 0.0035059)
   (LNUC_RNUC_ag 0.00197628)
   (LNUC_RNUC_at 0.159652)
   (LNUC_RNUC_ca 0.00385514)
   (LNUC_RNUC_cc 0.00143573)
   (LNUC_RNUC_cg 0.240649)
   (LNUC_RNUC_ct 0.00299132)
   (LNUC_RNUC_ga 0.00299132)
   (LNUC_RNUC_gc 0.259715)
   (LNUC_RNUC_gg 0.00368786)
   (LNUC_RNUC_gt 0.066477)
   (LNUC_RNUC_ta 0.175921)
   (LNUC_RNUC_tc 0.00147712)
   (LNUC_RNUC_tg 0.0673116)
   (LNUC_RNUC_tt 0.00517581))
  ((LNUC_RNUC_aX 0.00188921)
   (LNUC_RNUC_cX 0.0084023)
   (LNUC_RNUC_gX 0.347962)
   (LNUC_RNUC_tX 0.0252775)
   (LNUC_RNUC_Xa 2.02731e-06)
   (LNUC_RNUC_Xc 2.09746e-05)
   (LNUC_RNUC_Xg 0.610473)
   (LNUC_RNUC_Xt 0.00627573)
   (LNUC_RNUC_XX 2.02591e-06))

   ;; loops (NUC)
  ((NUC_a 0.378667)
   (NUC_c 0.161544)
   (NUC_g 0.207761)
   (NUC_t 0.237994)
   (NUC_X 0.0145484))

 )

 ;; constant rate parameters
 (const-rate
  ;; stems (LNUC RNUC)
  (LNUC_RNUC_aa2ac 0)
  (LNUC_RNUC_aa2ag 0)
  (LNUC_RNUC_aa2at 0.426022)
  (LNUC_RNUC_aa2ca 0.0643017)
  (LNUC_RNUC_aa2cc 0.000514693)
  (LNUC_RNUC_aa2cg 0.213602)
  (LNUC_RNUC_aa2ct 0.00296655)
  (LNUC_RNUC_aa2ga 0.0668467)
  (LNUC_RNUC_aa2gc 0.260255)
  (LNUC_RNUC_aa2gg 0.0111023)
  (LNUC_RNUC_aa2gt 0.121498)
  (LNUC_RNUC_aa2ta 0.820173)
  (LNUC_RNUC_aa2tc 0.00268661)
  (LNUC_RNUC_aa2tg 0.0849643)
  (LNUC_RNUC_aa2tt 0.0106205)
  (LNUC_RNUC_ac2aa 0.0190113)
  (LNUC_RNUC_ac2ag 0.0107912)
  (LNUC_RNUC_ac2at 2.38171)
  (LNUC_RNUC_ac2ca 0.0121744)
  (LNUC_RNUC_ac2cc 1.90612e-05)
  (LNUC_RNUC_ac2cg 0.168404)
  (LNUC_RNUC_ac2ct 0.0019914)
  (LNUC_RNUC_ac2ga 0.0182495)
  (LNUC_RNUC_ac2gc 0.959264)
  (LNUC_RNUC_ac2gg 0.00782334)
  (LNUC_RNUC_ac2gt 0.185952)
  (LNUC_RNUC_ac2ta 0.114626)
  (LNUC_RNUC_ac2tc 0.0489007)
  (LNUC_RNUC_ac2tg 0.0600372)
  (LNUC_RNUC_ac2tt 0.058924)
  (LNUC_RNUC_ag2aa 0.165856)
  (LNUC_RNUC_ag2ac 0.0118006)
  (LNUC_RNUC_ag2at 0.312083)
  (LNUC_RNUC_ag2ca 0.00880171)
  (LNUC_RNUC_ag2cc 0.000397433)
  (LNUC_RNUC_ag2cg 0.350868)
  (LNUC_RNUC_ag2ct 0.00443761)
  (LNUC_RNUC_ag2ga 0.084553)
  (LNUC_RNUC_ag2gc 0.297714)
  (LNUC_RNUC_ag2gg 0.636839)
  (LNUC_RNUC_ag2gt 0.0938174)
  (LNUC_RNUC_ag2ta 0.0798812)
  (LNUC_RNUC_ag2tc 0.0433447)
  (LNUC_RNUC_ag2tg 0.120742)
  (LNUC_RNUC_ag2tt 0.00210057)
  (LNUC_RNUC_at2aa 0.00630189)
  (LNUC_RNUC_at2ac 0.0390644)
  (LNUC_RNUC_at2ag 0.00539186)
  (LNUC_RNUC_at2ca 0.00268102)
  (LNUC_RNUC_at2cc 0.00162426)
  (LNUC_RNUC_at2cg 0.246046)
  (LNUC_RNUC_at2ct 0.00257359)
  (LNUC_RNUC_at2ga 0.00223115)
  (LNUC_RNUC_at2gc 0.696889)
  (LNUC_RNUC_at2gg 0.00266066)
  (LNUC_RNUC_at2gt 0.203768)
  (LNUC_RNUC_at2ta 0.222673)
  (LNUC_RNUC_at2tc 0.00434629)
  (LNUC_RNUC_at2tg 0.0420719)
  (LNUC_RNUC_at2tt 0.015625)
  (LNUC_RNUC_ca2aa 0.0516531)
  (LNUC_RNUC_ca2ac 0.0113356)
  (LNUC_RNUC_ca2ag 0.0156575)
  (LNUC_RNUC_ca2at 0.164026)
  (LNUC_RNUC_ca2cc 0.00880782)
  (LNUC_RNUC_ca2cg 1.63807)
  (LNUC_RNUC_ca2ct 0.0286365)
  (LNUC_RNUC_ca2ga 0.000130198)
  (LNUC_RNUC_ca2gc 0.0190641)
  (LNUC_RNUC_ca2gg 1.83337e-07)
  (LNUC_RNUC_ca2gt 0.0524469)
  (LNUC_RNUC_ca2ta 1.66555)
  (LNUC_RNUC_ca2tc 0.0523379)
  (LNUC_RNUC_ca2tg 0.164368)
  (LNUC_RNUC_ca2tt 0.075363)
  (LNUC_RNUC_cc2aa 0.00189446)
  (LNUC_RNUC_cc2ac 4.36094e-05)
  (LNUC_RNUC_cc2ag 0.00233559)
  (LNUC_RNUC_cc2at 0.371646)
  (LNUC_RNUC_cc2ca 0.0329175)
  (LNUC_RNUC_cc2cg 1.06807)
  (LNUC_RNUC_cc2ct 0.346517)
  (LNUC_RNUC_cc2ga 1.99732e-13)
  (LNUC_RNUC_cc2gc 0.935623)
  (LNUC_RNUC_cc2gg 0.00236163)
  (LNUC_RNUC_cc2gt 0.00109337)
  (LNUC_RNUC_cc2ta 0.039418)
  (LNUC_RNUC_cc2tc 3.86527e-06)
  (LNUC_RNUC_cc2tg 2.55025e-05)
  (LNUC_RNUC_cc2tt 0.484309)
  (LNUC_RNUC_cg2aa 0.00172642)
  (LNUC_RNUC_cg2ac 0.00153771)
  (LNUC_RNUC_cg2ag 0.00354009)
  (LNUC_RNUC_cg2at 0.140535)
  (LNUC_RNUC_cg2ca 0.0157228)
  (LNUC_RNUC_cg2cc 0.00265697)
  (LNUC_RNUC_cg2ct 0.00220195)
  (LNUC_RNUC_cg2ga 0.00423918)
  (LNUC_RNUC_cg2gc 0.180867)
  (LNUC_RNUC_cg2gg 0.00213729)
  (LNUC_RNUC_cg2gt 0.0388484)
  (LNUC_RNUC_cg2ta 0.407536)
  (LNUC_RNUC_cg2tc 0.00106421)
  (LNUC_RNUC_cg2tg 0.165282)
  (LNUC_RNUC_cg2tt 0.00299963)
  (LNUC_RNUC_ct2aa 0.00257003)
  (LNUC_RNUC_ct2ac 0.00138779)
  (LNUC_RNUC_ct2ag 0.0160198)
  (LNUC_RNUC_ct2at 0.169341)
  (LNUC_RNUC_ct2ca 0.0336792)
  (LNUC_RNUC_ct2cc 0.0945355)
  (LNUC_RNUC_ct2cg 0.244516)
  (LNUC_RNUC_ct2ga 0.0112261)
  (LNUC_RNUC_ct2gc 0.0246546)
  (LNUC_RNUC_ct2gg 0)
  (LNUC_RNUC_ct2gt 0.0103588)
  (LNUC_RNUC_ct2ta 0.138889)
  (LNUC_RNUC_ct2tc 0.495515)
  (LNUC_RNUC_ct2tg 0.0148747)
  (LNUC_RNUC_ct2tt 0.11941)
  (LNUC_RNUC_ga2aa 0.0359466)
  (LNUC_RNUC_ga2ac 0.0122167)
  (LNUC_RNUC_ga2ag 0.0555527)
  (LNUC_RNUC_ga2at 0.0816728)
  (LNUC_RNUC_ga2ca 4.8522e-05)
  (LNUC_RNUC_ga2cc 6.70286e-17)
  (LNUC_RNUC_ga2cg 0.265908)
  (LNUC_RNUC_ga2ct 0.00599926)
  (LNUC_RNUC_ga2gc 0.339622)
  (LNUC_RNUC_ga2gg 0.115503)
  (LNUC_RNUC_ga2gt 0.130308)
  (LNUC_RNUC_ga2ta 0.0579519)
  (LNUC_RNUC_ga2tc 0.0381546)
  (LNUC_RNUC_ga2tg 0.0224678)
  (LNUC_RNUC_ga2tt 1.88491e-07)
  (LNUC_RNUC_gc2aa 0.0018879)
  (LNUC_RNUC_gc2ac 0.00647906)
  (LNUC_RNUC_gc2ag 0.00255936)
  (LNUC_RNUC_gc2at 0.34151)
  (LNUC_RNUC_gc2ca 0.000133119)
  (LNUC_RNUC_gc2cc 0.00206163)
  (LNUC_RNUC_gc2cg 0.157672)
  (LNUC_RNUC_gc2ct 0.000187867)
  (LNUC_RNUC_gc2ga 0.00459411)
  (LNUC_RNUC_gc2gg 0.00479584)
  (LNUC_RNUC_gc2gt 0.143986)
  (LNUC_RNUC_gc2ta 0.103378)
  (LNUC_RNUC_gc2tc 0.00484597)
  (LNUC_RNUC_gc2tg 0.0355748)
  (LNUC_RNUC_gc2tt 0.000746795)
  (LNUC_RNUC_gg2aa 0.00638101)
  (LNUC_RNUC_gg2ac 0.00524806)
  (LNUC_RNUC_gg2ag 0.431072)
  (LNUC_RNUC_gg2at 0.0879605)
  (LNUC_RNUC_gg2ca 1.93512e-08)
  (LNUC_RNUC_gg2cc 0.000390335)
  (LNUC_RNUC_gg2cg 0.13277)
  (LNUC_RNUC_gg2ct 0)
  (LNUC_RNUC_gg2ga 0.117848)
  (LNUC_RNUC_gg2gc 0.320634)
  (LNUC_RNUC_gg2gt 0.099995)
  (LNUC_RNUC_gg2ta 0.0551307)
  (LNUC_RNUC_gg2tc 1.54794e-09)
  (LNUC_RNUC_gg2tg 0.23229)
  (LNUC_RNUC_gg2tt 1.91672e-05)
  (LNUC_RNUC_gt2aa 0.00360946)
  (LNUC_RNUC_gt2ac 0.0053287)
  (LNUC_RNUC_gt2ag 0.00343137)
  (LNUC_RNUC_gt2at 0.405845)
  (LNUC_RNUC_gt2ca 0.00175782)
  (LNUC_RNUC_gt2cc 6.34038e-06)
  (LNUC_RNUC_gt2cg 0.138216)
  (LNUC_RNUC_gt2ct 0.000343834)
  (LNUC_RNUC_gt2ga 0.00707527)
  (LNUC_RNUC_gt2gc 0.594604)
  (LNUC_RNUC_gt2gg 0.00581905)
  (LNUC_RNUC_gt2ta 0.117034)
  (LNUC_RNUC_gt2tc 0.0120485)
  (LNUC_RNUC_gt2tg 0.0476626)
  (LNUC_RNUC_gt2tt 0.0112651)
  (LNUC_RNUC_ta2aa 0.0100616)
  (LNUC_RNUC_ta2ac 0.00141009)
  (LNUC_RNUC_ta2ag 0.00116053)
  (LNUC_RNUC_ta2at 0.188809)
  (LNUC_RNUC_ta2ca 0.0247059)
  (LNUC_RNUC_ta2cc 0.000146075)
  (LNUC_RNUC_ta2cg 0.610473)
  (LNUC_RNUC_ta2ct 0.00176148)
  (LNUC_RNUC_ta2ga 0.0013955)
  (LNUC_RNUC_ta2gc 0.176409)
  (LNUC_RNUC_ta2gg 0.00127437)
  (LNUC_RNUC_ta2gt 0.0489007)
  (LNUC_RNUC_ta2tc 0.0162772)
  (LNUC_RNUC_ta2tg 0.183519)
  (LNUC_RNUC_ta2tt 0.0061722)
  (LNUC_RNUC_tc2aa 0.00380734)
  (LNUC_RNUC_tc2ac 0.0618535)
  (LNUC_RNUC_tc2ag 0.121835)
  (LNUC_RNUC_tc2at 0.390123)
  (LNUC_RNUC_tc2ca 0.0886338)
  (LNUC_RNUC_tc2cc 7.66119e-06)
  (LNUC_RNUC_tc2cg 0.217789)
  (LNUC_RNUC_tc2ct 0.645729)
  (LNUC_RNUC_tc2ga 0.0643909)
  (LNUC_RNUC_tc2gc 0.767373)
  (LNUC_RNUC_tc2gg 9.32708e-08)
  (LNUC_RNUC_tc2gt 0.586824)
  (LNUC_RNUC_tc2ta 1.7411)
  (LNUC_RNUC_tc2tg 0.229251)
  (LNUC_RNUC_tc2tt 0.3813)
  (LNUC_RNUC_tg2aa 0.00263497)
  (LNUC_RNUC_tg2ac 0.00221881)
  (LNUC_RNUC_tg2ag 0.00446229)
  (LNUC_RNUC_tg2at 0.0915688)
  (LNUC_RNUC_tg2ca 0.00573894)
  (LNUC_RNUC_tg2cc 1.25136e-07)
  (LNUC_RNUC_tg2cg 0.635516)
  (LNUC_RNUC_tg2ct 0.000516122)
  (LNUC_RNUC_tg2ga 0.00131202)
  (LNUC_RNUC_tg2gc 0.155286)
  (LNUC_RNUC_tg2gg 0.0136401)
  (LNUC_RNUC_tg2gt 0.0511189)
  (LNUC_RNUC_tg2ta 0.467164)
  (LNUC_RNUC_tg2tc 0.00531395)
  (LNUC_RNUC_tg2tt 0.0159976)
  (LNUC_RNUC_tt2aa 0.00653317)
  (LNUC_RNUC_tt2ac 0.0294824)
  (LNUC_RNUC_tt2ag 0.00192891)
  (LNUC_RNUC_tt2at 0.653383)
  (LNUC_RNUC_tt2ca 0.0527385)
  (LNUC_RNUC_tt2cc 0.0747387)
  (LNUC_RNUC_tt2cg 0.214939)
  (LNUC_RNUC_tt2ct 0.0665693)
  (LNUC_RNUC_tt2ga 3.17443e-07)
  (LNUC_RNUC_tt2gc 0.0661094)
  (LNUC_RNUC_tt2gg 2.06858e-05)
  (LNUC_RNUC_tt2gt 0.228616)
  (LNUC_RNUC_tt2ta 0.299785)
  (LNUC_RNUC_tt2tc 0.139274)
  (LNUC_RNUC_tt2tg 0.288371)
  (LNUC_RNUC_aX2XX 0)
  (LNUC_RNUC_Xa2XX 0)
  (LNUC_RNUC_XX2aX 0)
  (LNUC_RNUC_XX2Xa 0)
  (LNUC_RNUC_aa2aX 4.92871e-11)
  (LNUC_RNUC_aa2Xa 1.21167e-14)
  (LNUC_RNUC_aX2aa 0.00821229)
  (LNUC_RNUC_Xa2aa 1.29286e-06)
  (LNUC_RNUC_ac2aX 4.08181e-11)
  (LNUC_RNUC_ac2Xa 1.68179e-14)
  (LNUC_RNUC_aX2ac 0.00750985)
  (LNUC_RNUC_Xc2ac 0.000794866)
  (LNUC_RNUC_Xa2Xc 3.24658e-15)
  (LNUC_RNUC_aX2cX 8.8906e-10)
  (LNUC_RNUC_ag2aX 3.23151e-11)
  (LNUC_RNUC_ag2Xa 1.20413e-14)
  (LNUC_RNUC_aX2ag 0.00834426)
  (LNUC_RNUC_Xg2ag 1.3765)
  (LNUC_RNUC_Xa2Xg 1.6303e-12)
  (LNUC_RNUC_aX2gX 1.40486e-08)
  (LNUC_RNUC_at2aX 2.63757e-11)
  (LNUC_RNUC_at2Xa 1.22433e-14)
  (LNUC_RNUC_aX2at 0.00647008)
  (LNUC_RNUC_Xt2at 0.0308625)
  (LNUC_RNUC_Xa2Xt 6.79745e-14)
  (LNUC_RNUC_aX2tX 1.94579e-09)
  (LNUC_RNUC_cX2XX 0)
  (LNUC_RNUC_Xc2XX 0)
  (LNUC_RNUC_XX2cX 0)
  (LNUC_RNUC_XX2Xc 0)
  (LNUC_RNUC_ca2cX 1.04629e-10)
  (LNUC_RNUC_ca2Xc 4.3867e-12)
  (LNUC_RNUC_cX2ca 0.0334002)
  (LNUC_RNUC_Xa2ca 1.17411e-06)
  (LNUC_RNUC_Xc2Xa 5.76341e-15)
  (LNUC_RNUC_cX2aX 1.09836e-09)
  (LNUC_RNUC_cc2cX 1.20104e-10)
  (LNUC_RNUC_cc2Xc 1.60563e-12)
  (LNUC_RNUC_cX2cc 0.0301646)
  (LNUC_RNUC_Xc2cc 0.000416903)
  (LNUC_RNUC_cg2cX 7.7447e-11)
  (LNUC_RNUC_cg2Xc 3.95078e-12)
  (LNUC_RNUC_cX2cg 0.0380489)
  (LNUC_RNUC_Xg2cg 1.31677)
  (LNUC_RNUC_Xc2Xg 7.34255e-10)
  (LNUC_RNUC_cX2gX 1.84475e-08)
  (LNUC_RNUC_ct2cX 9.39706e-11)
  (LNUC_RNUC_ct2Xc 4.73754e-12)
  (LNUC_RNUC_cX2ct 0.0323296)
  (LNUC_RNUC_Xt2ct 0.0276994)
  (LNUC_RNUC_Xc2Xt 4.62101e-11)
  (LNUC_RNUC_cX2tX 5.72928e-09)
  (LNUC_RNUC_gX2XX 0)
  (LNUC_RNUC_Xg2XX 0)
  (LNUC_RNUC_XX2gX 0)
  (LNUC_RNUC_XX2Xg 0)
  (LNUC_RNUC_ga2gX 1.15777e-09)
  (LNUC_RNUC_ga2Xg 1.79173e-09)
  (LNUC_RNUC_gX2ga 0.90125)
  (LNUC_RNUC_Xa2ga 1.41085e-06)
  (LNUC_RNUC_Xg2Xa 2.22562e-11)
  (LNUC_RNUC_gX2aX 3.48081e-08)
  (LNUC_RNUC_gc2gX 1.15617e-09)
  (LNUC_RNUC_gc2Xg 1.63054e-09)
  (LNUC_RNUC_gX2gc 0.889459)
  (LNUC_RNUC_Xc2gc 0.00105247)
  (LNUC_RNUC_Xg2Xc 6.23484e-09)
  (LNUC_RNUC_gX2cX 3.79585e-08)
  (LNUC_RNUC_gg2gX 1.36448e-09)
  (LNUC_RNUC_gg2Xg 1.46444e-09)
  (LNUC_RNUC_gX2gg 0.948027)
  (LNUC_RNUC_Xg2gg 1.45498)
  (LNUC_RNUC_gt2gX 1.21197e-09)
  (LNUC_RNUC_gt2Xg 1.74032e-09)
  (LNUC_RNUC_gX2gt 0.857971)
  (LNUC_RNUC_Xt2gt 0.0316862)
  (LNUC_RNUC_Xg2Xt 9.92058e-08)
  (LNUC_RNUC_gX2tX 1.45045e-07)
  (LNUC_RNUC_tX2XX 0)
  (LNUC_RNUC_Xt2XX 0)
  (LNUC_RNUC_XX2tX 0)
  (LNUC_RNUC_XX2Xt 0)
  (LNUC_RNUC_ta2tX 1.6705e-10)
  (LNUC_RNUC_ta2Xt 8.48085e-11)
  (LNUC_RNUC_tX2ta 0.0886952)
  (LNUC_RNUC_Xa2ta 1.54818e-06)
  (LNUC_RNUC_Xt2Xa 3.61017e-13)
  (LNUC_RNUC_tX2aX 3.29745e-09)
  (LNUC_RNUC_tc2tX 1.56839e-10)
  (LNUC_RNUC_tc2Xt 7.25617e-11)
  (LNUC_RNUC_tX2tc 0.0922697)
  (LNUC_RNUC_Xc2tc 0.000554702)
  (LNUC_RNUC_Xt2Xc 1.31429e-10)
  (LNUC_RNUC_tX2cX 8.00751e-09)
  (LNUC_RNUC_tg2tX 1.65322e-10)
  (LNUC_RNUC_tg2Xt 8.05681e-11)
  (LNUC_RNUC_tX2tg 0.0906217)
  (LNUC_RNUC_Xg2tg 1.24143)
  (LNUC_RNUC_Xt2Xg 3.41865e-08)
  (LNUC_RNUC_tX2gX 9.72316e-08)
  (LNUC_RNUC_tt2tX 1.70324e-10)
  (LNUC_RNUC_tt2Xt 9.46242e-11)
  (LNUC_RNUC_tX2tt 0.0806043)
  (LNUC_RNUC_Xt2tt 0.0262962)

  ;; loops (NUC)
  (NUC_a2c 0.104894)
  (NUC_a2g 0.176777)
  (NUC_a2t 0.146401)
  (NUC_c2a 0.294431)
  (NUC_c2g 0.0724435)
  (NUC_c2t 0.520511)
  (NUC_g2a 0.312083)
  (NUC_g2c 0.0440718)
  (NUC_g2t 0.139371)
  (NUC_t2a 0.278355)
  (NUC_t2c 0.351111)
  (NUC_t2g 0.147624)
  (NUC_a2X 0.37449)
  (NUC_c2X 0.747943)
  (NUC_g2X 0.49141)
  (NUC_t2X 0.786762)
  (NUC_X2a 1.6783)
  (NUC_X2c 2.36035)
  (NUC_X2g 1.80876)
  (NUC_X2t 2.91804)

 )







;; Chains


;;;;;;;;;;;;;;;
;; CONSERVED ;;
;;;;;;;;;;;;;;;

 ;; Rate matrix
 (chain
  (update-policy parametric)
  (terminal (CONS))

  ;; initial probability distribution
  (initial (state (a)) (prob CONS_at))
  (initial (state (c)) (prob CONS_gc))
  (initial (state (g)) (prob CONS_gc))
  (initial (state (t)) (prob CONS_at))

  ;; mutation rates
  (mutate (from (a)) (to (c)) (rate conserved_rate_scale * CONS_a2c_t2g))
  (mutate (from (a)) (to (g)) (rate conserved_rate_scale * CONS_a2g_t2c))
  (mutate (from (a)) (to (t)) (rate conserved_rate_scale * CONS_a2t_t2a))
  (mutate (from (c)) (to (a)) (rate conserved_rate_scale * CONS_c2a_g2t))
  (mutate (from (c)) (to (g)) (rate conserved_rate_scale * CONS_c2g_g2c))
  (mutate (from (c)) (to (t)) (rate conserved_rate_scale * CONS_c2t_g2a))
  (mutate (from (g)) (to (a)) (rate conserved_rate_scale * CONS_c2t_g2a))
  (mutate (from (g)) (to (c)) (rate conserved_rate_scale * CONS_c2g_g2c))
  (mutate (from (g)) (to (t)) (rate conserved_rate_scale * CONS_c2a_g2t))
  (mutate (from (t)) (to (a)) (rate conserved_rate_scale * CONS_a2t_t2a))
  (mutate (from (t)) (to (c)) (rate conserved_rate_scale * CONS_a2g_t2c))
  (mutate (from (t)) (to (g)) (rate conserved_rate_scale * CONS_a2c_t2g))
 )  ;; end chain CONS
 

;;;;;;;;;;;;;
;; NEUTRAL ;;
;;;;;;;;;;;;;

 ;; Rate matrix
 (chain
  (update-policy parametric)
  (terminal (X))

  ;; initial probability distribution
  (initial (state (a)) (prob X_at / 2))
  (initial (state (c)) (prob X_gc / 2))
  (initial (state (g)) (prob X_gc / 2))
  (initial (state (t)) (prob X_at / 2))
  (initial (state (_)) (prob X_gap))

  ;; mutation rates
  (mutate (from (a)) (to (c)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (a)) (to (g)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (a)) (to (t)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (a)) (to (_)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (c)) (to (a)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (c)) (to (g)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (c)) (to (t)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (c)) (to (_)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (g)) (to (a)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (g)) (to (c)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (g)) (to (t)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (g)) (to (_)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (t)) (to (a)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (t)) (to (c)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (t)) (to (g)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (t)) (to (_)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (_)) (to (a)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_)) (to (c)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_)) (to (g)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_)) (to (t)) (rate neutral_rate_scale * X_gap2a_gap2t))
 )  ;; end chain X

;;;;;;;;;;;;
;; INTRON ;;
;;;;;;;;;;;;

 ;; identical to NEUTRAL chain over (X), but with intron_rate_scale

 ;; Rate matrix
 (chain
  (update-policy parametric)
  (terminal (XINTRON))

  ;; initial probability distribution
  (initial (state (a)) (prob X_at / 2))
  (initial (state (c)) (prob X_gc / 2))
  (initial (state (g)) (prob X_gc / 2))
  (initial (state (t)) (prob X_at / 2))
  (initial (state (_)) (prob X_gap))

  ;; mutation rates
  (mutate (from (a)) (to (c)) (rate intron_rate_scale * X_a2c_t2g))
  (mutate (from (a)) (to (g)) (rate intron_rate_scale * X_a2g_t2c))
  (mutate (from (a)) (to (t)) (rate intron_rate_scale * X_a2t_t2a))
  (mutate (from (a)) (to (_)) (rate intron_rate_scale * X_a2gap_t2gap))
  (mutate (from (c)) (to (a)) (rate intron_rate_scale * X_c2a_g2t))
  (mutate (from (c)) (to (g)) (rate intron_rate_scale * X_c2g_g2c))
  (mutate (from (c)) (to (t)) (rate intron_rate_scale * X_c2t_g2a))
  (mutate (from (c)) (to (_)) (rate intron_rate_scale * X_c2gap_g2gap))
  (mutate (from (g)) (to (a)) (rate intron_rate_scale * X_c2t_g2a))
  (mutate (from (g)) (to (c)) (rate intron_rate_scale * X_c2g_g2c))
  (mutate (from (g)) (to (t)) (rate intron_rate_scale * X_c2a_g2t))
  (mutate (from (g)) (to (_)) (rate intron_rate_scale * X_c2gap_g2gap))
  (mutate (from (t)) (to (a)) (rate intron_rate_scale * X_a2t_t2a))
  (mutate (from (t)) (to (c)) (rate intron_rate_scale * X_a2g_t2c))
  (mutate (from (t)) (to (g)) (rate intron_rate_scale * X_a2c_t2g))
  (mutate (from (t)) (to (_)) (rate intron_rate_scale * X_a2gap_t2gap))
  (mutate (from (_)) (to (a)) (rate intron_rate_scale * X_gap2a_gap2t))
  (mutate (from (_)) (to (c)) (rate intron_rate_scale * X_gap2c_gap2g))
  (mutate (from (_)) (to (g)) (rate intron_rate_scale * X_gap2c_gap2g))
  (mutate (from (_)) (to (t)) (rate intron_rate_scale * X_gap2a_gap2t))
 )  ;; end chain XINTRON


;;;;;;;;;;;;
;; CODING ;;
;;;;;;;;;;;;

 ;; gene chain
 (chain
  (update-policy parametric)
  (terminal (COD1 COD2 COD3))

  ;; initial probability distribution
  (initial (state (a a a)) (prob COD1_COD2_COD3_aaa))
  (initial (state (c a a)) (prob COD1_COD2_COD3_caa))
  (initial (state (g a a)) (prob COD1_COD2_COD3_gaa))
  (initial (state (t a a)) (prob COD1_COD2_COD3_taa))
  (initial (state (a c a)) (prob COD1_COD2_COD3_aca))
  (initial (state (c c a)) (prob COD1_COD2_COD3_cca))
  (initial (state (g c a)) (prob COD1_COD2_COD3_gca))
  (initial (state (t c a)) (prob COD1_COD2_COD3_tca))
  (initial (state (a g a)) (prob COD1_COD2_COD3_aga))
  (initial (state (c g a)) (prob COD1_COD2_COD3_cga))
  (initial (state (g g a)) (prob COD1_COD2_COD3_gga))
  (initial (state (t g a)) (prob COD1_COD2_COD3_tga))
  (initial (state (a t a)) (prob COD1_COD2_COD3_ata))
  (initial (state (c t a)) (prob COD1_COD2_COD3_cta))
  (initial (state (g t a)) (prob COD1_COD2_COD3_gta))
  (initial (state (t t a)) (prob COD1_COD2_COD3_tta))
  (initial (state (a a c)) (prob COD1_COD2_COD3_aac))
  (initial (state (c a c)) (prob COD1_COD2_COD3_cac))
  (initial (state (g a c)) (prob COD1_COD2_COD3_gac))
  (initial (state (t a c)) (prob COD1_COD2_COD3_tac))
  (initial (state (a c c)) (prob COD1_COD2_COD3_acc))
  (initial (state (c c c)) (prob COD1_COD2_COD3_ccc))
  (initial (state (g c c)) (prob COD1_COD2_COD3_gcc))
  (initial (state (t c c)) (prob COD1_COD2_COD3_tcc))
  (initial (state (a g c)) (prob COD1_COD2_COD3_agc))
  (initial (state (c g c)) (prob COD1_COD2_COD3_cgc))
  (initial (state (g g c)) (prob COD1_COD2_COD3_ggc))
  (initial (state (t g c)) (prob COD1_COD2_COD3_tgc))
  (initial (state (a t c)) (prob COD1_COD2_COD3_atc))
  (initial (state (c t c)) (prob COD1_COD2_COD3_ctc))
  (initial (state (g t c)) (prob COD1_COD2_COD3_gtc))
  (initial (state (t t c)) (prob COD1_COD2_COD3_ttc))
  (initial (state (a a g)) (prob COD1_COD2_COD3_aag))
  (initial (state (c a g)) (prob COD1_COD2_COD3_cag))
  (initial (state (g a g)) (prob COD1_COD2_COD3_gag))
  (initial (state (t a g)) (prob COD1_COD2_COD3_tag))
  (initial (state (a c g)) (prob COD1_COD2_COD3_acg))
  (initial (state (c c g)) (prob COD1_COD2_COD3_ccg))
  (initial (state (g c g)) (prob COD1_COD2_COD3_gcg))
  (initial (state (t c g)) (prob COD1_COD2_COD3_tcg))
  (initial (state (a g g)) (prob COD1_COD2_COD3_agg))
  (initial (state (c g g)) (prob COD1_COD2_COD3_cgg))
  (initial (state (g g g)) (prob COD1_COD2_COD3_ggg))
  (initial (state (t g g)) (prob COD1_COD2_COD3_tgg))
  (initial (state (a t g)) (prob COD1_COD2_COD3_atg))
  (initial (state (c t g)) (prob COD1_COD2_COD3_ctg))
  (initial (state (g t g)) (prob COD1_COD2_COD3_gtg))
  (initial (state (t t g)) (prob COD1_COD2_COD3_ttg))
  (initial (state (a a t)) (prob COD1_COD2_COD3_aat))
  (initial (state (c a t)) (prob COD1_COD2_COD3_cat))
  (initial (state (g a t)) (prob COD1_COD2_COD3_gat))
  (initial (state (t a t)) (prob COD1_COD2_COD3_tat))
  (initial (state (a c t)) (prob COD1_COD2_COD3_act))
  (initial (state (c c t)) (prob COD1_COD2_COD3_cct))
  (initial (state (g c t)) (prob COD1_COD2_COD3_gct))
  (initial (state (t c t)) (prob COD1_COD2_COD3_tct))
  (initial (state (a g t)) (prob COD1_COD2_COD3_agt))
  (initial (state (c g t)) (prob COD1_COD2_COD3_cgt))
  (initial (state (g g t)) (prob COD1_COD2_COD3_ggt))
  (initial (state (t g t)) (prob COD1_COD2_COD3_tgt))
  (initial (state (a t t)) (prob COD1_COD2_COD3_att))
  (initial (state (c t t)) (prob COD1_COD2_COD3_ctt))
  (initial (state (g t t)) (prob COD1_COD2_COD3_gtt))
  (initial (state (t t t)) (prob COD1_COD2_COD3_ttt))

  ;; mutation rates
  (mutate (from (a a a)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2caa))
  (mutate (from (a a a)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2gaa))
  (mutate (from (a a a)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2taa))
  (mutate (from (a a a)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2aca))
  (mutate (from (a a a)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2cca))
  (mutate (from (a a a)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2gca))
  (mutate (from (a a a)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2tca))
  (mutate (from (a a a)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2aga))
  (mutate (from (a a a)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2cga))
  (mutate (from (a a a)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2gga))
  (mutate (from (a a a)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2tga))
  (mutate (from (a a a)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2ata))
  (mutate (from (a a a)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2cta))
  (mutate (from (a a a)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2gta))
  (mutate (from (a a a)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2tta))
  (mutate (from (a a a)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2aac))
  (mutate (from (a a a)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2cac))
  (mutate (from (a a a)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2gac))
  (mutate (from (a a a)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2tac))
  (mutate (from (a a a)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2acc))
  (mutate (from (a a a)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2ccc))
  (mutate (from (a a a)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2gcc))
  (mutate (from (a a a)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2tcc))
  (mutate (from (a a a)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2agc))
  (mutate (from (a a a)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2cgc))
  (mutate (from (a a a)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2ggc))
  (mutate (from (a a a)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2tgc))
  (mutate (from (a a a)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2atc))
  (mutate (from (a a a)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2ctc))
  (mutate (from (a a a)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2gtc))
  (mutate (from (a a a)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2ttc))
  (mutate (from (a a a)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2aag))
  (mutate (from (a a a)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2cag))
  (mutate (from (a a a)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2gag))
  (mutate (from (a a a)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2tag))
  (mutate (from (a a a)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2acg))
  (mutate (from (a a a)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2ccg))
  (mutate (from (a a a)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2gcg))
  (mutate (from (a a a)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2tcg))
  (mutate (from (a a a)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2agg))
  (mutate (from (a a a)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2cgg))
  (mutate (from (a a a)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2ggg))
  (mutate (from (a a a)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2tgg))
  (mutate (from (a a a)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2atg))
  (mutate (from (a a a)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2ctg))
  (mutate (from (a a a)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2gtg))
  (mutate (from (a a a)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2ttg))
  (mutate (from (a a a)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2aat))
  (mutate (from (a a a)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2cat))
  (mutate (from (a a a)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2gat))
  (mutate (from (a a a)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2tat))
  (mutate (from (a a a)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2act))
  (mutate (from (a a a)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2cct))
  (mutate (from (a a a)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2gct))
  (mutate (from (a a a)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2tct))
  (mutate (from (a a a)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2agt))
  (mutate (from (a a a)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2cgt))
  (mutate (from (a a a)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2ggt))
  (mutate (from (a a a)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2tgt))
  (mutate (from (a a a)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2att))
  (mutate (from (a a a)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2ctt))
  (mutate (from (a a a)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2gtt))
  (mutate (from (a a a)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_aaa2ttt))
  (mutate (from (c a a)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_caa2aaa))
  (mutate (from (c a a)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_caa2gaa))
  (mutate (from (c a a)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_caa2taa))
  (mutate (from (c a a)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_caa2aca))
  (mutate (from (c a a)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_caa2cca))
  (mutate (from (c a a)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_caa2gca))
  (mutate (from (c a a)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_caa2tca))
  (mutate (from (c a a)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_caa2aga))
  (mutate (from (c a a)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_caa2cga))
  (mutate (from (c a a)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_caa2gga))
  (mutate (from (c a a)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_caa2tga))
  (mutate (from (c a a)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_caa2ata))
  (mutate (from (c a a)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_caa2cta))
  (mutate (from (c a a)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_caa2gta))
  (mutate (from (c a a)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_caa2tta))
  (mutate (from (c a a)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_caa2aac))
  (mutate (from (c a a)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_caa2cac))
  (mutate (from (c a a)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_caa2gac))
  (mutate (from (c a a)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_caa2tac))
  (mutate (from (c a a)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_caa2acc))
  (mutate (from (c a a)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_caa2ccc))
  (mutate (from (c a a)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_caa2gcc))
  (mutate (from (c a a)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_caa2tcc))
  (mutate (from (c a a)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_caa2agc))
  (mutate (from (c a a)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_caa2cgc))
  (mutate (from (c a a)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_caa2ggc))
  (mutate (from (c a a)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_caa2tgc))
  (mutate (from (c a a)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_caa2atc))
  (mutate (from (c a a)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_caa2ctc))
  (mutate (from (c a a)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_caa2gtc))
  (mutate (from (c a a)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_caa2ttc))
  (mutate (from (c a a)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_caa2aag))
  (mutate (from (c a a)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_caa2cag))
  (mutate (from (c a a)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_caa2gag))
  (mutate (from (c a a)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_caa2tag))
  (mutate (from (c a a)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_caa2acg))
  (mutate (from (c a a)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_caa2ccg))
  (mutate (from (c a a)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_caa2gcg))
  (mutate (from (c a a)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_caa2tcg))
  (mutate (from (c a a)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_caa2agg))
  (mutate (from (c a a)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_caa2cgg))
  (mutate (from (c a a)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_caa2ggg))
  (mutate (from (c a a)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_caa2tgg))
  (mutate (from (c a a)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_caa2atg))
  (mutate (from (c a a)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_caa2ctg))
  (mutate (from (c a a)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_caa2gtg))
  (mutate (from (c a a)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_caa2ttg))
  (mutate (from (c a a)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_caa2aat))
  (mutate (from (c a a)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_caa2cat))
  (mutate (from (c a a)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_caa2gat))
  (mutate (from (c a a)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_caa2tat))
  (mutate (from (c a a)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_caa2act))
  (mutate (from (c a a)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_caa2cct))
  (mutate (from (c a a)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_caa2gct))
  (mutate (from (c a a)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_caa2tct))
  (mutate (from (c a a)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_caa2agt))
  (mutate (from (c a a)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_caa2cgt))
  (mutate (from (c a a)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_caa2ggt))
  (mutate (from (c a a)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_caa2tgt))
  (mutate (from (c a a)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_caa2att))
  (mutate (from (c a a)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_caa2ctt))
  (mutate (from (c a a)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_caa2gtt))
  (mutate (from (c a a)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_caa2ttt))
  (mutate (from (g a a)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2aaa))
  (mutate (from (g a a)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2caa))
  (mutate (from (g a a)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2taa))
  (mutate (from (g a a)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2aca))
  (mutate (from (g a a)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2cca))
  (mutate (from (g a a)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2gca))
  (mutate (from (g a a)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2tca))
  (mutate (from (g a a)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2aga))
  (mutate (from (g a a)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2cga))
  (mutate (from (g a a)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2gga))
  (mutate (from (g a a)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2tga))
  (mutate (from (g a a)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2ata))
  (mutate (from (g a a)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2cta))
  (mutate (from (g a a)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2gta))
  (mutate (from (g a a)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2tta))
  (mutate (from (g a a)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2aac))
  (mutate (from (g a a)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2cac))
  (mutate (from (g a a)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2gac))
  (mutate (from (g a a)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2tac))
  (mutate (from (g a a)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2acc))
  (mutate (from (g a a)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2ccc))
  (mutate (from (g a a)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2gcc))
  (mutate (from (g a a)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2tcc))
  (mutate (from (g a a)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2agc))
  (mutate (from (g a a)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2cgc))
  (mutate (from (g a a)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2ggc))
  (mutate (from (g a a)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2tgc))
  (mutate (from (g a a)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2atc))
  (mutate (from (g a a)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2ctc))
  (mutate (from (g a a)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2gtc))
  (mutate (from (g a a)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2ttc))
  (mutate (from (g a a)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2aag))
  (mutate (from (g a a)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2cag))
  (mutate (from (g a a)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2gag))
  (mutate (from (g a a)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2tag))
  (mutate (from (g a a)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2acg))
  (mutate (from (g a a)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2ccg))
  (mutate (from (g a a)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2gcg))
  (mutate (from (g a a)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2tcg))
  (mutate (from (g a a)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2agg))
  (mutate (from (g a a)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2cgg))
  (mutate (from (g a a)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2ggg))
  (mutate (from (g a a)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2tgg))
  (mutate (from (g a a)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2atg))
  (mutate (from (g a a)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2ctg))
  (mutate (from (g a a)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2gtg))
  (mutate (from (g a a)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2ttg))
  (mutate (from (g a a)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2aat))
  (mutate (from (g a a)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2cat))
  (mutate (from (g a a)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2gat))
  (mutate (from (g a a)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2tat))
  (mutate (from (g a a)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2act))
  (mutate (from (g a a)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2cct))
  (mutate (from (g a a)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2gct))
  (mutate (from (g a a)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2tct))
  (mutate (from (g a a)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2agt))
  (mutate (from (g a a)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2cgt))
  (mutate (from (g a a)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2ggt))
  (mutate (from (g a a)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2tgt))
  (mutate (from (g a a)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2att))
  (mutate (from (g a a)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2ctt))
  (mutate (from (g a a)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2gtt))
  (mutate (from (g a a)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_gaa2ttt))
  (mutate (from (t a a)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_taa2aaa))
  (mutate (from (t a a)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_taa2caa))
  (mutate (from (t a a)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_taa2gaa))
  (mutate (from (t a a)) (to (t a a)) (rate 0))
  (mutate (from (t a a)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_taa2aca))
  (mutate (from (t a a)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_taa2cca))
  (mutate (from (t a a)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_taa2gca))
  (mutate (from (t a a)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_taa2tca))
  (mutate (from (t a a)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_taa2aga))
  (mutate (from (t a a)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_taa2cga))
  (mutate (from (t a a)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_taa2gga))
  (mutate (from (t a a)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_taa2tga))
  (mutate (from (t a a)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_taa2ata))
  (mutate (from (t a a)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_taa2cta))
  (mutate (from (t a a)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_taa2gta))
  (mutate (from (t a a)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_taa2tta))
  (mutate (from (t a a)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_taa2aac))
  (mutate (from (t a a)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_taa2cac))
  (mutate (from (t a a)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_taa2gac))
  (mutate (from (t a a)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_taa2tac))
  (mutate (from (t a a)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_taa2acc))
  (mutate (from (t a a)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_taa2ccc))
  (mutate (from (t a a)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_taa2gcc))
  (mutate (from (t a a)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_taa2tcc))
  (mutate (from (t a a)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_taa2agc))
  (mutate (from (t a a)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_taa2cgc))
  (mutate (from (t a a)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_taa2ggc))
  (mutate (from (t a a)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_taa2tgc))
  (mutate (from (t a a)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_taa2atc))
  (mutate (from (t a a)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_taa2ctc))
  (mutate (from (t a a)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_taa2gtc))
  (mutate (from (t a a)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_taa2ttc))
  (mutate (from (t a a)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_taa2aag))
  (mutate (from (t a a)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_taa2cag))
  (mutate (from (t a a)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_taa2gag))
  (mutate (from (t a a)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_taa2tag))
  (mutate (from (t a a)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_taa2acg))
  (mutate (from (t a a)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_taa2ccg))
  (mutate (from (t a a)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_taa2gcg))
  (mutate (from (t a a)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_taa2tcg))
  (mutate (from (t a a)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_taa2agg))
  (mutate (from (t a a)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_taa2cgg))
  (mutate (from (t a a)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_taa2ggg))
  (mutate (from (t a a)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_taa2tgg))
  (mutate (from (t a a)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_taa2atg))
  (mutate (from (t a a)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_taa2ctg))
  (mutate (from (t a a)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_taa2gtg))
  (mutate (from (t a a)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_taa2ttg))
  (mutate (from (t a a)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_taa2aat))
  (mutate (from (t a a)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_taa2cat))
  (mutate (from (t a a)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_taa2gat))
  (mutate (from (t a a)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_taa2tat))
  (mutate (from (t a a)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_taa2act))
  (mutate (from (t a a)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_taa2cct))
  (mutate (from (t a a)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_taa2gct))
  (mutate (from (t a a)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_taa2tct))
  (mutate (from (t a a)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_taa2agt))
  (mutate (from (t a a)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_taa2cgt))
  (mutate (from (t a a)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_taa2ggt))
  (mutate (from (t a a)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_taa2tgt))
  (mutate (from (t a a)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_taa2att))
  (mutate (from (t a a)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_taa2ctt))
  (mutate (from (t a a)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_taa2gtt))
  (mutate (from (t a a)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_taa2ttt))
  (mutate (from (a c a)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_aca2aaa))
  (mutate (from (a c a)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_aca2caa))
  (mutate (from (a c a)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_aca2gaa))
  (mutate (from (a c a)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_aca2taa))
  (mutate (from (a c a)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_aca2cca))
  (mutate (from (a c a)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_aca2gca))
  (mutate (from (a c a)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_aca2tca))
  (mutate (from (a c a)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_aca2aga))
  (mutate (from (a c a)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_aca2cga))
  (mutate (from (a c a)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_aca2gga))
  (mutate (from (a c a)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_aca2tga))
  (mutate (from (a c a)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_aca2ata))
  (mutate (from (a c a)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_aca2cta))
  (mutate (from (a c a)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_aca2gta))
  (mutate (from (a c a)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_aca2tta))
  (mutate (from (a c a)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_aca2aac))
  (mutate (from (a c a)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_aca2cac))
  (mutate (from (a c a)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_aca2gac))
  (mutate (from (a c a)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_aca2tac))
  (mutate (from (a c a)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_aca2acc))
  (mutate (from (a c a)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_aca2ccc))
  (mutate (from (a c a)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_aca2gcc))
  (mutate (from (a c a)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_aca2tcc))
  (mutate (from (a c a)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_aca2agc))
  (mutate (from (a c a)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_aca2cgc))
  (mutate (from (a c a)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_aca2ggc))
  (mutate (from (a c a)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_aca2tgc))
  (mutate (from (a c a)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_aca2atc))
  (mutate (from (a c a)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_aca2ctc))
  (mutate (from (a c a)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_aca2gtc))
  (mutate (from (a c a)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_aca2ttc))
  (mutate (from (a c a)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_aca2aag))
  (mutate (from (a c a)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_aca2cag))
  (mutate (from (a c a)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_aca2gag))
  (mutate (from (a c a)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_aca2tag))
  (mutate (from (a c a)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_aca2acg))
  (mutate (from (a c a)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_aca2ccg))
  (mutate (from (a c a)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_aca2gcg))
  (mutate (from (a c a)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_aca2tcg))
  (mutate (from (a c a)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_aca2agg))
  (mutate (from (a c a)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_aca2cgg))
  (mutate (from (a c a)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_aca2ggg))
  (mutate (from (a c a)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_aca2tgg))
  (mutate (from (a c a)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_aca2atg))
  (mutate (from (a c a)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_aca2ctg))
  (mutate (from (a c a)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_aca2gtg))
  (mutate (from (a c a)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_aca2ttg))
  (mutate (from (a c a)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_aca2aat))
  (mutate (from (a c a)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_aca2cat))
  (mutate (from (a c a)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_aca2gat))
  (mutate (from (a c a)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_aca2tat))
  (mutate (from (a c a)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_aca2act))
  (mutate (from (a c a)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_aca2cct))
  (mutate (from (a c a)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_aca2gct))
  (mutate (from (a c a)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_aca2tct))
  (mutate (from (a c a)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_aca2agt))
  (mutate (from (a c a)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_aca2cgt))
  (mutate (from (a c a)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_aca2ggt))
  (mutate (from (a c a)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_aca2tgt))
  (mutate (from (a c a)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_aca2att))
  (mutate (from (a c a)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_aca2ctt))
  (mutate (from (a c a)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_aca2gtt))
  (mutate (from (a c a)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_aca2ttt))
  (mutate (from (c c a)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_cca2aaa))
  (mutate (from (c c a)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_cca2caa))
  (mutate (from (c c a)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_cca2gaa))
  (mutate (from (c c a)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_cca2taa))
  (mutate (from (c c a)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_cca2aca))
  (mutate (from (c c a)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_cca2gca))
  (mutate (from (c c a)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_cca2tca))
  (mutate (from (c c a)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_cca2aga))
  (mutate (from (c c a)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_cca2cga))
  (mutate (from (c c a)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_cca2gga))
  (mutate (from (c c a)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_cca2tga))
  (mutate (from (c c a)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_cca2ata))
  (mutate (from (c c a)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_cca2cta))
  (mutate (from (c c a)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_cca2gta))
  (mutate (from (c c a)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_cca2tta))
  (mutate (from (c c a)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_cca2aac))
  (mutate (from (c c a)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_cca2cac))
  (mutate (from (c c a)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_cca2gac))
  (mutate (from (c c a)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_cca2tac))
  (mutate (from (c c a)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_cca2acc))
  (mutate (from (c c a)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_cca2ccc))
  (mutate (from (c c a)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_cca2gcc))
  (mutate (from (c c a)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_cca2tcc))
  (mutate (from (c c a)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_cca2agc))
  (mutate (from (c c a)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_cca2cgc))
  (mutate (from (c c a)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_cca2ggc))
  (mutate (from (c c a)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_cca2tgc))
  (mutate (from (c c a)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_cca2atc))
  (mutate (from (c c a)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_cca2ctc))
  (mutate (from (c c a)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_cca2gtc))
  (mutate (from (c c a)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_cca2ttc))
  (mutate (from (c c a)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_cca2aag))
  (mutate (from (c c a)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_cca2cag))
  (mutate (from (c c a)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_cca2gag))
  (mutate (from (c c a)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_cca2tag))
  (mutate (from (c c a)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_cca2acg))
  (mutate (from (c c a)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_cca2ccg))
  (mutate (from (c c a)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_cca2gcg))
  (mutate (from (c c a)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_cca2tcg))
  (mutate (from (c c a)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_cca2agg))
  (mutate (from (c c a)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_cca2cgg))
  (mutate (from (c c a)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_cca2ggg))
  (mutate (from (c c a)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_cca2tgg))
  (mutate (from (c c a)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_cca2atg))
  (mutate (from (c c a)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_cca2ctg))
  (mutate (from (c c a)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_cca2gtg))
  (mutate (from (c c a)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_cca2ttg))
  (mutate (from (c c a)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_cca2aat))
  (mutate (from (c c a)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_cca2cat))
  (mutate (from (c c a)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_cca2gat))
  (mutate (from (c c a)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_cca2tat))
  (mutate (from (c c a)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_cca2act))
  (mutate (from (c c a)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_cca2cct))
  (mutate (from (c c a)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_cca2gct))
  (mutate (from (c c a)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_cca2tct))
  (mutate (from (c c a)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_cca2agt))
  (mutate (from (c c a)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_cca2cgt))
  (mutate (from (c c a)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_cca2ggt))
  (mutate (from (c c a)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_cca2tgt))
  (mutate (from (c c a)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_cca2att))
  (mutate (from (c c a)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_cca2ctt))
  (mutate (from (c c a)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_cca2gtt))
  (mutate (from (c c a)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_cca2ttt))
  (mutate (from (g c a)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_gca2aaa))
  (mutate (from (g c a)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_gca2caa))
  (mutate (from (g c a)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_gca2gaa))
  (mutate (from (g c a)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_gca2taa))
  (mutate (from (g c a)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_gca2aca))
  (mutate (from (g c a)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_gca2cca))
  (mutate (from (g c a)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_gca2tca))
  (mutate (from (g c a)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_gca2aga))
  (mutate (from (g c a)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_gca2cga))
  (mutate (from (g c a)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_gca2gga))
  (mutate (from (g c a)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_gca2tga))
  (mutate (from (g c a)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_gca2ata))
  (mutate (from (g c a)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_gca2cta))
  (mutate (from (g c a)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_gca2gta))
  (mutate (from (g c a)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_gca2tta))
  (mutate (from (g c a)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_gca2aac))
  (mutate (from (g c a)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_gca2cac))
  (mutate (from (g c a)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_gca2gac))
  (mutate (from (g c a)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_gca2tac))
  (mutate (from (g c a)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_gca2acc))
  (mutate (from (g c a)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_gca2ccc))
  (mutate (from (g c a)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_gca2gcc))
  (mutate (from (g c a)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_gca2tcc))
  (mutate (from (g c a)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_gca2agc))
  (mutate (from (g c a)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_gca2cgc))
  (mutate (from (g c a)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_gca2ggc))
  (mutate (from (g c a)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_gca2tgc))
  (mutate (from (g c a)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_gca2atc))
  (mutate (from (g c a)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_gca2ctc))
  (mutate (from (g c a)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_gca2gtc))
  (mutate (from (g c a)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_gca2ttc))
  (mutate (from (g c a)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_gca2aag))
  (mutate (from (g c a)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_gca2cag))
  (mutate (from (g c a)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_gca2gag))
  (mutate (from (g c a)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_gca2tag))
  (mutate (from (g c a)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_gca2acg))
  (mutate (from (g c a)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_gca2ccg))
  (mutate (from (g c a)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_gca2gcg))
  (mutate (from (g c a)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_gca2tcg))
  (mutate (from (g c a)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_gca2agg))
  (mutate (from (g c a)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_gca2cgg))
  (mutate (from (g c a)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_gca2ggg))
  (mutate (from (g c a)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_gca2tgg))
  (mutate (from (g c a)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_gca2atg))
  (mutate (from (g c a)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_gca2ctg))
  (mutate (from (g c a)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_gca2gtg))
  (mutate (from (g c a)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_gca2ttg))
  (mutate (from (g c a)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_gca2aat))
  (mutate (from (g c a)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_gca2cat))
  (mutate (from (g c a)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_gca2gat))
  (mutate (from (g c a)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_gca2tat))
  (mutate (from (g c a)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_gca2act))
  (mutate (from (g c a)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_gca2cct))
  (mutate (from (g c a)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_gca2gct))
  (mutate (from (g c a)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_gca2tct))
  (mutate (from (g c a)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_gca2agt))
  (mutate (from (g c a)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_gca2cgt))
  (mutate (from (g c a)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_gca2ggt))
  (mutate (from (g c a)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_gca2tgt))
  (mutate (from (g c a)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_gca2att))
  (mutate (from (g c a)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_gca2ctt))
  (mutate (from (g c a)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_gca2gtt))
  (mutate (from (g c a)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_gca2ttt))
  (mutate (from (t c a)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_tca2aaa))
  (mutate (from (t c a)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_tca2caa))
  (mutate (from (t c a)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_tca2gaa))
  (mutate (from (t c a)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_tca2taa))
  (mutate (from (t c a)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_tca2aca))
  (mutate (from (t c a)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_tca2cca))
  (mutate (from (t c a)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_tca2gca))
  (mutate (from (t c a)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_tca2aga))
  (mutate (from (t c a)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_tca2cga))
  (mutate (from (t c a)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_tca2gga))
  (mutate (from (t c a)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_tca2tga))
  (mutate (from (t c a)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_tca2ata))
  (mutate (from (t c a)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_tca2cta))
  (mutate (from (t c a)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_tca2gta))
  (mutate (from (t c a)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_tca2tta))
  (mutate (from (t c a)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_tca2aac))
  (mutate (from (t c a)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_tca2cac))
  (mutate (from (t c a)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_tca2gac))
  (mutate (from (t c a)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_tca2tac))
  (mutate (from (t c a)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_tca2acc))
  (mutate (from (t c a)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_tca2ccc))
  (mutate (from (t c a)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_tca2gcc))
  (mutate (from (t c a)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_tca2tcc))
  (mutate (from (t c a)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_tca2agc))
  (mutate (from (t c a)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_tca2cgc))
  (mutate (from (t c a)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_tca2ggc))
  (mutate (from (t c a)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_tca2tgc))
  (mutate (from (t c a)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_tca2atc))
  (mutate (from (t c a)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_tca2ctc))
  (mutate (from (t c a)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_tca2gtc))
  (mutate (from (t c a)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_tca2ttc))
  (mutate (from (t c a)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_tca2aag))
  (mutate (from (t c a)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_tca2cag))
  (mutate (from (t c a)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_tca2gag))
  (mutate (from (t c a)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_tca2tag))
  (mutate (from (t c a)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_tca2acg))
  (mutate (from (t c a)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_tca2ccg))
  (mutate (from (t c a)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_tca2gcg))
  (mutate (from (t c a)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_tca2tcg))
  (mutate (from (t c a)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_tca2agg))
  (mutate (from (t c a)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_tca2cgg))
  (mutate (from (t c a)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_tca2ggg))
  (mutate (from (t c a)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_tca2tgg))
  (mutate (from (t c a)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_tca2atg))
  (mutate (from (t c a)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_tca2ctg))
  (mutate (from (t c a)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_tca2gtg))
  (mutate (from (t c a)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_tca2ttg))
  (mutate (from (t c a)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_tca2aat))
  (mutate (from (t c a)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_tca2cat))
  (mutate (from (t c a)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_tca2gat))
  (mutate (from (t c a)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_tca2tat))
  (mutate (from (t c a)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_tca2act))
  (mutate (from (t c a)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_tca2cct))
  (mutate (from (t c a)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_tca2gct))
  (mutate (from (t c a)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_tca2tct))
  (mutate (from (t c a)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_tca2agt))
  (mutate (from (t c a)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_tca2cgt))
  (mutate (from (t c a)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_tca2ggt))
  (mutate (from (t c a)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_tca2tgt))
  (mutate (from (t c a)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_tca2att))
  (mutate (from (t c a)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_tca2ctt))
  (mutate (from (t c a)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_tca2gtt))
  (mutate (from (t c a)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_tca2ttt))
  (mutate (from (a g a)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_aga2aaa))
  (mutate (from (a g a)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_aga2caa))
  (mutate (from (a g a)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_aga2gaa))
  (mutate (from (a g a)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_aga2taa))
  (mutate (from (a g a)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_aga2aca))
  (mutate (from (a g a)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_aga2cca))
  (mutate (from (a g a)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_aga2gca))
  (mutate (from (a g a)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_aga2tca))
  (mutate (from (a g a)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_aga2cga))
  (mutate (from (a g a)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_aga2gga))
  (mutate (from (a g a)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_aga2tga))
  (mutate (from (a g a)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_aga2ata))
  (mutate (from (a g a)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_aga2cta))
  (mutate (from (a g a)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_aga2gta))
  (mutate (from (a g a)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_aga2tta))
  (mutate (from (a g a)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_aga2aac))
  (mutate (from (a g a)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_aga2cac))
  (mutate (from (a g a)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_aga2gac))
  (mutate (from (a g a)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_aga2tac))
  (mutate (from (a g a)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_aga2acc))
  (mutate (from (a g a)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_aga2ccc))
  (mutate (from (a g a)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_aga2gcc))
  (mutate (from (a g a)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_aga2tcc))
  (mutate (from (a g a)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_aga2agc))
  (mutate (from (a g a)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_aga2cgc))
  (mutate (from (a g a)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_aga2ggc))
  (mutate (from (a g a)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_aga2tgc))
  (mutate (from (a g a)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_aga2atc))
  (mutate (from (a g a)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_aga2ctc))
  (mutate (from (a g a)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_aga2gtc))
  (mutate (from (a g a)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_aga2ttc))
  (mutate (from (a g a)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_aga2aag))
  (mutate (from (a g a)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_aga2cag))
  (mutate (from (a g a)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_aga2gag))
  (mutate (from (a g a)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_aga2tag))
  (mutate (from (a g a)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_aga2acg))
  (mutate (from (a g a)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_aga2ccg))
  (mutate (from (a g a)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_aga2gcg))
  (mutate (from (a g a)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_aga2tcg))
  (mutate (from (a g a)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_aga2agg))
  (mutate (from (a g a)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_aga2cgg))
  (mutate (from (a g a)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_aga2ggg))
  (mutate (from (a g a)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_aga2tgg))
  (mutate (from (a g a)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_aga2atg))
  (mutate (from (a g a)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_aga2ctg))
  (mutate (from (a g a)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_aga2gtg))
  (mutate (from (a g a)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_aga2ttg))
  (mutate (from (a g a)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_aga2aat))
  (mutate (from (a g a)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_aga2cat))
  (mutate (from (a g a)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_aga2gat))
  (mutate (from (a g a)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_aga2tat))
  (mutate (from (a g a)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_aga2act))
  (mutate (from (a g a)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_aga2cct))
  (mutate (from (a g a)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_aga2gct))
  (mutate (from (a g a)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_aga2tct))
  (mutate (from (a g a)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_aga2agt))
  (mutate (from (a g a)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_aga2cgt))
  (mutate (from (a g a)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_aga2ggt))
  (mutate (from (a g a)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_aga2tgt))
  (mutate (from (a g a)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_aga2att))
  (mutate (from (a g a)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_aga2ctt))
  (mutate (from (a g a)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_aga2gtt))
  (mutate (from (a g a)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_aga2ttt))
  (mutate (from (c g a)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_cga2aaa))
  (mutate (from (c g a)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_cga2caa))
  (mutate (from (c g a)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_cga2gaa))
  (mutate (from (c g a)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_cga2taa))
  (mutate (from (c g a)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_cga2aca))
  (mutate (from (c g a)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_cga2cca))
  (mutate (from (c g a)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_cga2gca))
  (mutate (from (c g a)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_cga2tca))
  (mutate (from (c g a)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_cga2aga))
  (mutate (from (c g a)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_cga2gga))
  (mutate (from (c g a)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_cga2tga))
  (mutate (from (c g a)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_cga2ata))
  (mutate (from (c g a)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_cga2cta))
  (mutate (from (c g a)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_cga2gta))
  (mutate (from (c g a)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_cga2tta))
  (mutate (from (c g a)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_cga2aac))
  (mutate (from (c g a)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_cga2cac))
  (mutate (from (c g a)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_cga2gac))
  (mutate (from (c g a)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_cga2tac))
  (mutate (from (c g a)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_cga2acc))
  (mutate (from (c g a)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_cga2ccc))
  (mutate (from (c g a)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_cga2gcc))
  (mutate (from (c g a)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_cga2tcc))
  (mutate (from (c g a)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_cga2agc))
  (mutate (from (c g a)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_cga2cgc))
  (mutate (from (c g a)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_cga2ggc))
  (mutate (from (c g a)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_cga2tgc))
  (mutate (from (c g a)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_cga2atc))
  (mutate (from (c g a)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_cga2ctc))
  (mutate (from (c g a)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_cga2gtc))
  (mutate (from (c g a)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_cga2ttc))
  (mutate (from (c g a)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_cga2aag))
  (mutate (from (c g a)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_cga2cag))
  (mutate (from (c g a)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_cga2gag))
  (mutate (from (c g a)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_cga2tag))
  (mutate (from (c g a)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_cga2acg))
  (mutate (from (c g a)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_cga2ccg))
  (mutate (from (c g a)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_cga2gcg))
  (mutate (from (c g a)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_cga2tcg))
  (mutate (from (c g a)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_cga2agg))
  (mutate (from (c g a)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_cga2cgg))
  (mutate (from (c g a)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_cga2ggg))
  (mutate (from (c g a)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_cga2tgg))
  (mutate (from (c g a)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_cga2atg))
  (mutate (from (c g a)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_cga2ctg))
  (mutate (from (c g a)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_cga2gtg))
  (mutate (from (c g a)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_cga2ttg))
  (mutate (from (c g a)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_cga2aat))
  (mutate (from (c g a)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_cga2cat))
  (mutate (from (c g a)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_cga2gat))
  (mutate (from (c g a)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_cga2tat))
  (mutate (from (c g a)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_cga2act))
  (mutate (from (c g a)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_cga2cct))
  (mutate (from (c g a)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_cga2gct))
  (mutate (from (c g a)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_cga2tct))
  (mutate (from (c g a)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_cga2agt))
  (mutate (from (c g a)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_cga2cgt))
  (mutate (from (c g a)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_cga2ggt))
  (mutate (from (c g a)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_cga2tgt))
  (mutate (from (c g a)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_cga2att))
  (mutate (from (c g a)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_cga2ctt))
  (mutate (from (c g a)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_cga2gtt))
  (mutate (from (c g a)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_cga2ttt))
  (mutate (from (g g a)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_gga2aaa))
  (mutate (from (g g a)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_gga2caa))
  (mutate (from (g g a)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_gga2gaa))
  (mutate (from (g g a)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_gga2taa))
  (mutate (from (g g a)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_gga2aca))
  (mutate (from (g g a)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_gga2cca))
  (mutate (from (g g a)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_gga2gca))
  (mutate (from (g g a)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_gga2tca))
  (mutate (from (g g a)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_gga2aga))
  (mutate (from (g g a)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_gga2cga))
  (mutate (from (g g a)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_gga2tga))
  (mutate (from (g g a)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_gga2ata))
  (mutate (from (g g a)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_gga2cta))
  (mutate (from (g g a)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_gga2gta))
  (mutate (from (g g a)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_gga2tta))
  (mutate (from (g g a)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_gga2aac))
  (mutate (from (g g a)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_gga2cac))
  (mutate (from (g g a)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_gga2gac))
  (mutate (from (g g a)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_gga2tac))
  (mutate (from (g g a)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_gga2acc))
  (mutate (from (g g a)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_gga2ccc))
  (mutate (from (g g a)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_gga2gcc))
  (mutate (from (g g a)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_gga2tcc))
  (mutate (from (g g a)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_gga2agc))
  (mutate (from (g g a)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_gga2cgc))
  (mutate (from (g g a)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_gga2ggc))
  (mutate (from (g g a)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_gga2tgc))
  (mutate (from (g g a)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_gga2atc))
  (mutate (from (g g a)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_gga2ctc))
  (mutate (from (g g a)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_gga2gtc))
  (mutate (from (g g a)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_gga2ttc))
  (mutate (from (g g a)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_gga2aag))
  (mutate (from (g g a)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_gga2cag))
  (mutate (from (g g a)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_gga2gag))
  (mutate (from (g g a)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_gga2tag))
  (mutate (from (g g a)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_gga2acg))
  (mutate (from (g g a)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_gga2ccg))
  (mutate (from (g g a)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_gga2gcg))
  (mutate (from (g g a)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_gga2tcg))
  (mutate (from (g g a)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_gga2agg))
  (mutate (from (g g a)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_gga2cgg))
  (mutate (from (g g a)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_gga2ggg))
  (mutate (from (g g a)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_gga2tgg))
  (mutate (from (g g a)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_gga2atg))
  (mutate (from (g g a)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_gga2ctg))
  (mutate (from (g g a)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_gga2gtg))
  (mutate (from (g g a)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_gga2ttg))
  (mutate (from (g g a)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_gga2aat))
  (mutate (from (g g a)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_gga2cat))
  (mutate (from (g g a)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_gga2gat))
  (mutate (from (g g a)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_gga2tat))
  (mutate (from (g g a)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_gga2act))
  (mutate (from (g g a)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_gga2cct))
  (mutate (from (g g a)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_gga2gct))
  (mutate (from (g g a)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_gga2tct))
  (mutate (from (g g a)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_gga2agt))
  (mutate (from (g g a)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_gga2cgt))
  (mutate (from (g g a)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_gga2ggt))
  (mutate (from (g g a)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_gga2tgt))
  (mutate (from (g g a)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_gga2att))
  (mutate (from (g g a)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_gga2ctt))
  (mutate (from (g g a)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_gga2gtt))
  (mutate (from (g g a)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_gga2ttt))
  (mutate (from (t g a)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_tga2aaa))
  (mutate (from (t g a)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_tga2caa))
  (mutate (from (t g a)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_tga2gaa))
  (mutate (from (t g a)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_tga2taa))
  (mutate (from (t g a)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_tga2aca))
  (mutate (from (t g a)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_tga2cca))
  (mutate (from (t g a)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_tga2gca))
  (mutate (from (t g a)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_tga2tca))
  (mutate (from (t g a)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_tga2aga))
  (mutate (from (t g a)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_tga2cga))
  (mutate (from (t g a)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_tga2gga))
  (mutate (from (t g a)) (to (t g a)) (rate 0))
  (mutate (from (t g a)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_tga2ata))
  (mutate (from (t g a)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_tga2cta))
  (mutate (from (t g a)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_tga2gta))
  (mutate (from (t g a)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_tga2tta))
  (mutate (from (t g a)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_tga2aac))
  (mutate (from (t g a)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_tga2cac))
  (mutate (from (t g a)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_tga2gac))
  (mutate (from (t g a)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_tga2tac))
  (mutate (from (t g a)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_tga2acc))
  (mutate (from (t g a)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_tga2ccc))
  (mutate (from (t g a)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_tga2gcc))
  (mutate (from (t g a)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_tga2tcc))
  (mutate (from (t g a)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_tga2agc))
  (mutate (from (t g a)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_tga2cgc))
  (mutate (from (t g a)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_tga2ggc))
  (mutate (from (t g a)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_tga2tgc))
  (mutate (from (t g a)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_tga2atc))
  (mutate (from (t g a)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_tga2ctc))
  (mutate (from (t g a)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_tga2gtc))
  (mutate (from (t g a)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_tga2ttc))
  (mutate (from (t g a)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_tga2aag))
  (mutate (from (t g a)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_tga2cag))
  (mutate (from (t g a)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_tga2gag))
  (mutate (from (t g a)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_tga2tag))
  (mutate (from (t g a)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_tga2acg))
  (mutate (from (t g a)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_tga2ccg))
  (mutate (from (t g a)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_tga2gcg))
  (mutate (from (t g a)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_tga2tcg))
  (mutate (from (t g a)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_tga2agg))
  (mutate (from (t g a)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_tga2cgg))
  (mutate (from (t g a)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_tga2ggg))
  (mutate (from (t g a)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_tga2tgg))
  (mutate (from (t g a)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_tga2atg))
  (mutate (from (t g a)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_tga2ctg))
  (mutate (from (t g a)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_tga2gtg))
  (mutate (from (t g a)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_tga2ttg))
  (mutate (from (t g a)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_tga2aat))
  (mutate (from (t g a)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_tga2cat))
  (mutate (from (t g a)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_tga2gat))
  (mutate (from (t g a)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_tga2tat))
  (mutate (from (t g a)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_tga2act))
  (mutate (from (t g a)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_tga2cct))
  (mutate (from (t g a)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_tga2gct))
  (mutate (from (t g a)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_tga2tct))
  (mutate (from (t g a)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_tga2agt))
  (mutate (from (t g a)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_tga2cgt))
  (mutate (from (t g a)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_tga2ggt))
  (mutate (from (t g a)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_tga2tgt))
  (mutate (from (t g a)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_tga2att))
  (mutate (from (t g a)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_tga2ctt))
  (mutate (from (t g a)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_tga2gtt))
  (mutate (from (t g a)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_tga2ttt))
  (mutate (from (a t a)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_ata2aaa))
  (mutate (from (a t a)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_ata2caa))
  (mutate (from (a t a)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_ata2gaa))
  (mutate (from (a t a)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_ata2taa))
  (mutate (from (a t a)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_ata2aca))
  (mutate (from (a t a)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_ata2cca))
  (mutate (from (a t a)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_ata2gca))
  (mutate (from (a t a)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_ata2tca))
  (mutate (from (a t a)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_ata2aga))
  (mutate (from (a t a)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_ata2cga))
  (mutate (from (a t a)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_ata2gga))
  (mutate (from (a t a)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_ata2tga))
  (mutate (from (a t a)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_ata2cta))
  (mutate (from (a t a)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_ata2gta))
  (mutate (from (a t a)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_ata2tta))
  (mutate (from (a t a)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_ata2aac))
  (mutate (from (a t a)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_ata2cac))
  (mutate (from (a t a)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_ata2gac))
  (mutate (from (a t a)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_ata2tac))
  (mutate (from (a t a)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_ata2acc))
  (mutate (from (a t a)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_ata2ccc))
  (mutate (from (a t a)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_ata2gcc))
  (mutate (from (a t a)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_ata2tcc))
  (mutate (from (a t a)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_ata2agc))
  (mutate (from (a t a)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_ata2cgc))
  (mutate (from (a t a)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_ata2ggc))
  (mutate (from (a t a)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_ata2tgc))
  (mutate (from (a t a)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_ata2atc))
  (mutate (from (a t a)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_ata2ctc))
  (mutate (from (a t a)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_ata2gtc))
  (mutate (from (a t a)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_ata2ttc))
  (mutate (from (a t a)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_ata2aag))
  (mutate (from (a t a)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_ata2cag))
  (mutate (from (a t a)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_ata2gag))
  (mutate (from (a t a)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_ata2tag))
  (mutate (from (a t a)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_ata2acg))
  (mutate (from (a t a)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_ata2ccg))
  (mutate (from (a t a)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_ata2gcg))
  (mutate (from (a t a)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_ata2tcg))
  (mutate (from (a t a)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_ata2agg))
  (mutate (from (a t a)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_ata2cgg))
  (mutate (from (a t a)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_ata2ggg))
  (mutate (from (a t a)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_ata2tgg))
  (mutate (from (a t a)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_ata2atg))
  (mutate (from (a t a)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_ata2ctg))
  (mutate (from (a t a)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_ata2gtg))
  (mutate (from (a t a)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_ata2ttg))
  (mutate (from (a t a)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_ata2aat))
  (mutate (from (a t a)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_ata2cat))
  (mutate (from (a t a)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_ata2gat))
  (mutate (from (a t a)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_ata2tat))
  (mutate (from (a t a)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_ata2act))
  (mutate (from (a t a)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_ata2cct))
  (mutate (from (a t a)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_ata2gct))
  (mutate (from (a t a)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_ata2tct))
  (mutate (from (a t a)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_ata2agt))
  (mutate (from (a t a)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_ata2cgt))
  (mutate (from (a t a)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_ata2ggt))
  (mutate (from (a t a)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_ata2tgt))
  (mutate (from (a t a)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_ata2att))
  (mutate (from (a t a)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_ata2ctt))
  (mutate (from (a t a)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_ata2gtt))
  (mutate (from (a t a)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_ata2ttt))
  (mutate (from (c t a)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_cta2aaa))
  (mutate (from (c t a)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_cta2caa))
  (mutate (from (c t a)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_cta2gaa))
  (mutate (from (c t a)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_cta2taa))
  (mutate (from (c t a)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_cta2aca))
  (mutate (from (c t a)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_cta2cca))
  (mutate (from (c t a)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_cta2gca))
  (mutate (from (c t a)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_cta2tca))
  (mutate (from (c t a)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_cta2aga))
  (mutate (from (c t a)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_cta2cga))
  (mutate (from (c t a)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_cta2gga))
  (mutate (from (c t a)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_cta2tga))
  (mutate (from (c t a)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_cta2ata))
  (mutate (from (c t a)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_cta2gta))
  (mutate (from (c t a)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_cta2tta))
  (mutate (from (c t a)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_cta2aac))
  (mutate (from (c t a)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_cta2cac))
  (mutate (from (c t a)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_cta2gac))
  (mutate (from (c t a)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_cta2tac))
  (mutate (from (c t a)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_cta2acc))
  (mutate (from (c t a)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_cta2ccc))
  (mutate (from (c t a)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_cta2gcc))
  (mutate (from (c t a)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_cta2tcc))
  (mutate (from (c t a)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_cta2agc))
  (mutate (from (c t a)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_cta2cgc))
  (mutate (from (c t a)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_cta2ggc))
  (mutate (from (c t a)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_cta2tgc))
  (mutate (from (c t a)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_cta2atc))
  (mutate (from (c t a)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_cta2ctc))
  (mutate (from (c t a)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_cta2gtc))
  (mutate (from (c t a)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_cta2ttc))
  (mutate (from (c t a)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_cta2aag))
  (mutate (from (c t a)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_cta2cag))
  (mutate (from (c t a)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_cta2gag))
  (mutate (from (c t a)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_cta2tag))
  (mutate (from (c t a)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_cta2acg))
  (mutate (from (c t a)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_cta2ccg))
  (mutate (from (c t a)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_cta2gcg))
  (mutate (from (c t a)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_cta2tcg))
  (mutate (from (c t a)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_cta2agg))
  (mutate (from (c t a)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_cta2cgg))
  (mutate (from (c t a)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_cta2ggg))
  (mutate (from (c t a)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_cta2tgg))
  (mutate (from (c t a)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_cta2atg))
  (mutate (from (c t a)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_cta2ctg))
  (mutate (from (c t a)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_cta2gtg))
  (mutate (from (c t a)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_cta2ttg))
  (mutate (from (c t a)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_cta2aat))
  (mutate (from (c t a)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_cta2cat))
  (mutate (from (c t a)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_cta2gat))
  (mutate (from (c t a)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_cta2tat))
  (mutate (from (c t a)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_cta2act))
  (mutate (from (c t a)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_cta2cct))
  (mutate (from (c t a)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_cta2gct))
  (mutate (from (c t a)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_cta2tct))
  (mutate (from (c t a)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_cta2agt))
  (mutate (from (c t a)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_cta2cgt))
  (mutate (from (c t a)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_cta2ggt))
  (mutate (from (c t a)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_cta2tgt))
  (mutate (from (c t a)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_cta2att))
  (mutate (from (c t a)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_cta2ctt))
  (mutate (from (c t a)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_cta2gtt))
  (mutate (from (c t a)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_cta2ttt))
  (mutate (from (g t a)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_gta2aaa))
  (mutate (from (g t a)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_gta2caa))
  (mutate (from (g t a)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_gta2gaa))
  (mutate (from (g t a)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_gta2taa))
  (mutate (from (g t a)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_gta2aca))
  (mutate (from (g t a)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_gta2cca))
  (mutate (from (g t a)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_gta2gca))
  (mutate (from (g t a)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_gta2tca))
  (mutate (from (g t a)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_gta2aga))
  (mutate (from (g t a)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_gta2cga))
  (mutate (from (g t a)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_gta2gga))
  (mutate (from (g t a)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_gta2tga))
  (mutate (from (g t a)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_gta2ata))
  (mutate (from (g t a)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_gta2cta))
  (mutate (from (g t a)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_gta2tta))
  (mutate (from (g t a)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_gta2aac))
  (mutate (from (g t a)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_gta2cac))
  (mutate (from (g t a)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_gta2gac))
  (mutate (from (g t a)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_gta2tac))
  (mutate (from (g t a)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_gta2acc))
  (mutate (from (g t a)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_gta2ccc))
  (mutate (from (g t a)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_gta2gcc))
  (mutate (from (g t a)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_gta2tcc))
  (mutate (from (g t a)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_gta2agc))
  (mutate (from (g t a)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_gta2cgc))
  (mutate (from (g t a)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_gta2ggc))
  (mutate (from (g t a)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_gta2tgc))
  (mutate (from (g t a)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_gta2atc))
  (mutate (from (g t a)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_gta2ctc))
  (mutate (from (g t a)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_gta2gtc))
  (mutate (from (g t a)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_gta2ttc))
  (mutate (from (g t a)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_gta2aag))
  (mutate (from (g t a)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_gta2cag))
  (mutate (from (g t a)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_gta2gag))
  (mutate (from (g t a)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_gta2tag))
  (mutate (from (g t a)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_gta2acg))
  (mutate (from (g t a)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_gta2ccg))
  (mutate (from (g t a)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_gta2gcg))
  (mutate (from (g t a)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_gta2tcg))
  (mutate (from (g t a)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_gta2agg))
  (mutate (from (g t a)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_gta2cgg))
  (mutate (from (g t a)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_gta2ggg))
  (mutate (from (g t a)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_gta2tgg))
  (mutate (from (g t a)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_gta2atg))
  (mutate (from (g t a)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_gta2ctg))
  (mutate (from (g t a)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_gta2gtg))
  (mutate (from (g t a)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_gta2ttg))
  (mutate (from (g t a)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_gta2aat))
  (mutate (from (g t a)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_gta2cat))
  (mutate (from (g t a)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_gta2gat))
  (mutate (from (g t a)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_gta2tat))
  (mutate (from (g t a)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_gta2act))
  (mutate (from (g t a)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_gta2cct))
  (mutate (from (g t a)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_gta2gct))
  (mutate (from (g t a)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_gta2tct))
  (mutate (from (g t a)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_gta2agt))
  (mutate (from (g t a)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_gta2cgt))
  (mutate (from (g t a)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_gta2ggt))
  (mutate (from (g t a)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_gta2tgt))
  (mutate (from (g t a)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_gta2att))
  (mutate (from (g t a)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_gta2ctt))
  (mutate (from (g t a)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_gta2gtt))
  (mutate (from (g t a)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_gta2ttt))
  (mutate (from (t t a)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_tta2aaa))
  (mutate (from (t t a)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_tta2caa))
  (mutate (from (t t a)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_tta2gaa))
  (mutate (from (t t a)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_tta2taa))
  (mutate (from (t t a)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_tta2aca))
  (mutate (from (t t a)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_tta2cca))
  (mutate (from (t t a)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_tta2gca))
  (mutate (from (t t a)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_tta2tca))
  (mutate (from (t t a)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_tta2aga))
  (mutate (from (t t a)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_tta2cga))
  (mutate (from (t t a)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_tta2gga))
  (mutate (from (t t a)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_tta2tga))
  (mutate (from (t t a)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_tta2ata))
  (mutate (from (t t a)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_tta2cta))
  (mutate (from (t t a)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_tta2gta))
  (mutate (from (t t a)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_tta2aac))
  (mutate (from (t t a)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_tta2cac))
  (mutate (from (t t a)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_tta2gac))
  (mutate (from (t t a)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_tta2tac))
  (mutate (from (t t a)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_tta2acc))
  (mutate (from (t t a)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_tta2ccc))
  (mutate (from (t t a)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_tta2gcc))
  (mutate (from (t t a)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_tta2tcc))
  (mutate (from (t t a)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_tta2agc))
  (mutate (from (t t a)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_tta2cgc))
  (mutate (from (t t a)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_tta2ggc))
  (mutate (from (t t a)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_tta2tgc))
  (mutate (from (t t a)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_tta2atc))
  (mutate (from (t t a)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_tta2ctc))
  (mutate (from (t t a)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_tta2gtc))
  (mutate (from (t t a)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_tta2ttc))
  (mutate (from (t t a)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_tta2aag))
  (mutate (from (t t a)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_tta2cag))
  (mutate (from (t t a)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_tta2gag))
  (mutate (from (t t a)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_tta2tag))
  (mutate (from (t t a)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_tta2acg))
  (mutate (from (t t a)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_tta2ccg))
  (mutate (from (t t a)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_tta2gcg))
  (mutate (from (t t a)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_tta2tcg))
  (mutate (from (t t a)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_tta2agg))
  (mutate (from (t t a)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_tta2cgg))
  (mutate (from (t t a)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_tta2ggg))
  (mutate (from (t t a)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_tta2tgg))
  (mutate (from (t t a)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_tta2atg))
  (mutate (from (t t a)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_tta2ctg))
  (mutate (from (t t a)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_tta2gtg))
  (mutate (from (t t a)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_tta2ttg))
  (mutate (from (t t a)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_tta2aat))
  (mutate (from (t t a)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_tta2cat))
  (mutate (from (t t a)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_tta2gat))
  (mutate (from (t t a)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_tta2tat))
  (mutate (from (t t a)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_tta2act))
  (mutate (from (t t a)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_tta2cct))
  (mutate (from (t t a)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_tta2gct))
  (mutate (from (t t a)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_tta2tct))
  (mutate (from (t t a)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_tta2agt))
  (mutate (from (t t a)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_tta2cgt))
  (mutate (from (t t a)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_tta2ggt))
  (mutate (from (t t a)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_tta2tgt))
  (mutate (from (t t a)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_tta2att))
  (mutate (from (t t a)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_tta2ctt))
  (mutate (from (t t a)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_tta2gtt))
  (mutate (from (t t a)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_tta2ttt))
  (mutate (from (a a c)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_aac2aaa))
  (mutate (from (a a c)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_aac2caa))
  (mutate (from (a a c)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_aac2gaa))
  (mutate (from (a a c)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_aac2taa))
  (mutate (from (a a c)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_aac2aca))
  (mutate (from (a a c)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_aac2cca))
  (mutate (from (a a c)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_aac2gca))
  (mutate (from (a a c)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_aac2tca))
  (mutate (from (a a c)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_aac2aga))
  (mutate (from (a a c)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_aac2cga))
  (mutate (from (a a c)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_aac2gga))
  (mutate (from (a a c)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_aac2tga))
  (mutate (from (a a c)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_aac2ata))
  (mutate (from (a a c)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_aac2cta))
  (mutate (from (a a c)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_aac2gta))
  (mutate (from (a a c)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_aac2tta))
  (mutate (from (a a c)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_aac2cac))
  (mutate (from (a a c)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_aac2gac))
  (mutate (from (a a c)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_aac2tac))
  (mutate (from (a a c)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_aac2acc))
  (mutate (from (a a c)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_aac2ccc))
  (mutate (from (a a c)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_aac2gcc))
  (mutate (from (a a c)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_aac2tcc))
  (mutate (from (a a c)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_aac2agc))
  (mutate (from (a a c)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_aac2cgc))
  (mutate (from (a a c)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_aac2ggc))
  (mutate (from (a a c)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_aac2tgc))
  (mutate (from (a a c)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_aac2atc))
  (mutate (from (a a c)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_aac2ctc))
  (mutate (from (a a c)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_aac2gtc))
  (mutate (from (a a c)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_aac2ttc))
  (mutate (from (a a c)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_aac2aag))
  (mutate (from (a a c)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_aac2cag))
  (mutate (from (a a c)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_aac2gag))
  (mutate (from (a a c)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_aac2tag))
  (mutate (from (a a c)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_aac2acg))
  (mutate (from (a a c)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_aac2ccg))
  (mutate (from (a a c)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_aac2gcg))
  (mutate (from (a a c)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_aac2tcg))
  (mutate (from (a a c)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_aac2agg))
  (mutate (from (a a c)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_aac2cgg))
  (mutate (from (a a c)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_aac2ggg))
  (mutate (from (a a c)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_aac2tgg))
  (mutate (from (a a c)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_aac2atg))
  (mutate (from (a a c)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_aac2ctg))
  (mutate (from (a a c)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_aac2gtg))
  (mutate (from (a a c)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_aac2ttg))
  (mutate (from (a a c)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_aac2aat))
  (mutate (from (a a c)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_aac2cat))
  (mutate (from (a a c)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_aac2gat))
  (mutate (from (a a c)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_aac2tat))
  (mutate (from (a a c)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_aac2act))
  (mutate (from (a a c)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_aac2cct))
  (mutate (from (a a c)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_aac2gct))
  (mutate (from (a a c)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_aac2tct))
  (mutate (from (a a c)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_aac2agt))
  (mutate (from (a a c)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_aac2cgt))
  (mutate (from (a a c)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_aac2ggt))
  (mutate (from (a a c)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_aac2tgt))
  (mutate (from (a a c)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_aac2att))
  (mutate (from (a a c)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_aac2ctt))
  (mutate (from (a a c)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_aac2gtt))
  (mutate (from (a a c)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_aac2ttt))
  (mutate (from (c a c)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_cac2aaa))
  (mutate (from (c a c)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_cac2caa))
  (mutate (from (c a c)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_cac2gaa))
  (mutate (from (c a c)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_cac2taa))
  (mutate (from (c a c)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_cac2aca))
  (mutate (from (c a c)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_cac2cca))
  (mutate (from (c a c)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_cac2gca))
  (mutate (from (c a c)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_cac2tca))
  (mutate (from (c a c)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_cac2aga))
  (mutate (from (c a c)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_cac2cga))
  (mutate (from (c a c)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_cac2gga))
  (mutate (from (c a c)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_cac2tga))
  (mutate (from (c a c)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_cac2ata))
  (mutate (from (c a c)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_cac2cta))
  (mutate (from (c a c)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_cac2gta))
  (mutate (from (c a c)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_cac2tta))
  (mutate (from (c a c)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_cac2aac))
  (mutate (from (c a c)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_cac2gac))
  (mutate (from (c a c)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_cac2tac))
  (mutate (from (c a c)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_cac2acc))
  (mutate (from (c a c)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_cac2ccc))
  (mutate (from (c a c)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_cac2gcc))
  (mutate (from (c a c)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_cac2tcc))
  (mutate (from (c a c)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_cac2agc))
  (mutate (from (c a c)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_cac2cgc))
  (mutate (from (c a c)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_cac2ggc))
  (mutate (from (c a c)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_cac2tgc))
  (mutate (from (c a c)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_cac2atc))
  (mutate (from (c a c)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_cac2ctc))
  (mutate (from (c a c)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_cac2gtc))
  (mutate (from (c a c)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_cac2ttc))
  (mutate (from (c a c)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_cac2aag))
  (mutate (from (c a c)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_cac2cag))
  (mutate (from (c a c)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_cac2gag))
  (mutate (from (c a c)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_cac2tag))
  (mutate (from (c a c)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_cac2acg))
  (mutate (from (c a c)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_cac2ccg))
  (mutate (from (c a c)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_cac2gcg))
  (mutate (from (c a c)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_cac2tcg))
  (mutate (from (c a c)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_cac2agg))
  (mutate (from (c a c)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_cac2cgg))
  (mutate (from (c a c)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_cac2ggg))
  (mutate (from (c a c)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_cac2tgg))
  (mutate (from (c a c)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_cac2atg))
  (mutate (from (c a c)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_cac2ctg))
  (mutate (from (c a c)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_cac2gtg))
  (mutate (from (c a c)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_cac2ttg))
  (mutate (from (c a c)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_cac2aat))
  (mutate (from (c a c)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_cac2cat))
  (mutate (from (c a c)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_cac2gat))
  (mutate (from (c a c)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_cac2tat))
  (mutate (from (c a c)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_cac2act))
  (mutate (from (c a c)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_cac2cct))
  (mutate (from (c a c)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_cac2gct))
  (mutate (from (c a c)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_cac2tct))
  (mutate (from (c a c)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_cac2agt))
  (mutate (from (c a c)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_cac2cgt))
  (mutate (from (c a c)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_cac2ggt))
  (mutate (from (c a c)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_cac2tgt))
  (mutate (from (c a c)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_cac2att))
  (mutate (from (c a c)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_cac2ctt))
  (mutate (from (c a c)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_cac2gtt))
  (mutate (from (c a c)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_cac2ttt))
  (mutate (from (g a c)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_gac2aaa))
  (mutate (from (g a c)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_gac2caa))
  (mutate (from (g a c)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_gac2gaa))
  (mutate (from (g a c)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_gac2taa))
  (mutate (from (g a c)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_gac2aca))
  (mutate (from (g a c)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_gac2cca))
  (mutate (from (g a c)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_gac2gca))
  (mutate (from (g a c)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_gac2tca))
  (mutate (from (g a c)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_gac2aga))
  (mutate (from (g a c)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_gac2cga))
  (mutate (from (g a c)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_gac2gga))
  (mutate (from (g a c)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_gac2tga))
  (mutate (from (g a c)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_gac2ata))
  (mutate (from (g a c)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_gac2cta))
  (mutate (from (g a c)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_gac2gta))
  (mutate (from (g a c)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_gac2tta))
  (mutate (from (g a c)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_gac2aac))
  (mutate (from (g a c)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_gac2cac))
  (mutate (from (g a c)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_gac2tac))
  (mutate (from (g a c)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_gac2acc))
  (mutate (from (g a c)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_gac2ccc))
  (mutate (from (g a c)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_gac2gcc))
  (mutate (from (g a c)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_gac2tcc))
  (mutate (from (g a c)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_gac2agc))
  (mutate (from (g a c)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_gac2cgc))
  (mutate (from (g a c)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_gac2ggc))
  (mutate (from (g a c)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_gac2tgc))
  (mutate (from (g a c)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_gac2atc))
  (mutate (from (g a c)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_gac2ctc))
  (mutate (from (g a c)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_gac2gtc))
  (mutate (from (g a c)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_gac2ttc))
  (mutate (from (g a c)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_gac2aag))
  (mutate (from (g a c)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_gac2cag))
  (mutate (from (g a c)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_gac2gag))
  (mutate (from (g a c)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_gac2tag))
  (mutate (from (g a c)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_gac2acg))
  (mutate (from (g a c)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_gac2ccg))
  (mutate (from (g a c)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_gac2gcg))
  (mutate (from (g a c)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_gac2tcg))
  (mutate (from (g a c)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_gac2agg))
  (mutate (from (g a c)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_gac2cgg))
  (mutate (from (g a c)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_gac2ggg))
  (mutate (from (g a c)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_gac2tgg))
  (mutate (from (g a c)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_gac2atg))
  (mutate (from (g a c)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_gac2ctg))
  (mutate (from (g a c)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_gac2gtg))
  (mutate (from (g a c)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_gac2ttg))
  (mutate (from (g a c)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_gac2aat))
  (mutate (from (g a c)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_gac2cat))
  (mutate (from (g a c)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_gac2gat))
  (mutate (from (g a c)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_gac2tat))
  (mutate (from (g a c)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_gac2act))
  (mutate (from (g a c)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_gac2cct))
  (mutate (from (g a c)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_gac2gct))
  (mutate (from (g a c)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_gac2tct))
  (mutate (from (g a c)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_gac2agt))
  (mutate (from (g a c)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_gac2cgt))
  (mutate (from (g a c)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_gac2ggt))
  (mutate (from (g a c)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_gac2tgt))
  (mutate (from (g a c)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_gac2att))
  (mutate (from (g a c)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_gac2ctt))
  (mutate (from (g a c)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_gac2gtt))
  (mutate (from (g a c)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_gac2ttt))
  (mutate (from (t a c)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_tac2aaa))
  (mutate (from (t a c)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_tac2caa))
  (mutate (from (t a c)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_tac2gaa))
  (mutate (from (t a c)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_tac2taa))
  (mutate (from (t a c)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_tac2aca))
  (mutate (from (t a c)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_tac2cca))
  (mutate (from (t a c)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_tac2gca))
  (mutate (from (t a c)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_tac2tca))
  (mutate (from (t a c)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_tac2aga))
  (mutate (from (t a c)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_tac2cga))
  (mutate (from (t a c)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_tac2gga))
  (mutate (from (t a c)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_tac2tga))
  (mutate (from (t a c)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_tac2ata))
  (mutate (from (t a c)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_tac2cta))
  (mutate (from (t a c)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_tac2gta))
  (mutate (from (t a c)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_tac2tta))
  (mutate (from (t a c)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_tac2aac))
  (mutate (from (t a c)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_tac2cac))
  (mutate (from (t a c)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_tac2gac))
  (mutate (from (t a c)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_tac2acc))
  (mutate (from (t a c)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_tac2ccc))
  (mutate (from (t a c)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_tac2gcc))
  (mutate (from (t a c)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_tac2tcc))
  (mutate (from (t a c)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_tac2agc))
  (mutate (from (t a c)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_tac2cgc))
  (mutate (from (t a c)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_tac2ggc))
  (mutate (from (t a c)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_tac2tgc))
  (mutate (from (t a c)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_tac2atc))
  (mutate (from (t a c)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_tac2ctc))
  (mutate (from (t a c)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_tac2gtc))
  (mutate (from (t a c)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_tac2ttc))
  (mutate (from (t a c)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_tac2aag))
  (mutate (from (t a c)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_tac2cag))
  (mutate (from (t a c)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_tac2gag))
  (mutate (from (t a c)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_tac2tag))
  (mutate (from (t a c)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_tac2acg))
  (mutate (from (t a c)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_tac2ccg))
  (mutate (from (t a c)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_tac2gcg))
  (mutate (from (t a c)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_tac2tcg))
  (mutate (from (t a c)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_tac2agg))
  (mutate (from (t a c)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_tac2cgg))
  (mutate (from (t a c)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_tac2ggg))
  (mutate (from (t a c)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_tac2tgg))
  (mutate (from (t a c)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_tac2atg))
  (mutate (from (t a c)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_tac2ctg))
  (mutate (from (t a c)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_tac2gtg))
  (mutate (from (t a c)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_tac2ttg))
  (mutate (from (t a c)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_tac2aat))
  (mutate (from (t a c)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_tac2cat))
  (mutate (from (t a c)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_tac2gat))
  (mutate (from (t a c)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_tac2tat))
  (mutate (from (t a c)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_tac2act))
  (mutate (from (t a c)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_tac2cct))
  (mutate (from (t a c)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_tac2gct))
  (mutate (from (t a c)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_tac2tct))
  (mutate (from (t a c)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_tac2agt))
  (mutate (from (t a c)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_tac2cgt))
  (mutate (from (t a c)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_tac2ggt))
  (mutate (from (t a c)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_tac2tgt))
  (mutate (from (t a c)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_tac2att))
  (mutate (from (t a c)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_tac2ctt))
  (mutate (from (t a c)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_tac2gtt))
  (mutate (from (t a c)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_tac2ttt))
  (mutate (from (a c c)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_acc2aaa))
  (mutate (from (a c c)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_acc2caa))
  (mutate (from (a c c)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_acc2gaa))
  (mutate (from (a c c)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_acc2taa))
  (mutate (from (a c c)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_acc2aca))
  (mutate (from (a c c)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_acc2cca))
  (mutate (from (a c c)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_acc2gca))
  (mutate (from (a c c)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_acc2tca))
  (mutate (from (a c c)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_acc2aga))
  (mutate (from (a c c)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_acc2cga))
  (mutate (from (a c c)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_acc2gga))
  (mutate (from (a c c)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_acc2tga))
  (mutate (from (a c c)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_acc2ata))
  (mutate (from (a c c)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_acc2cta))
  (mutate (from (a c c)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_acc2gta))
  (mutate (from (a c c)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_acc2tta))
  (mutate (from (a c c)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_acc2aac))
  (mutate (from (a c c)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_acc2cac))
  (mutate (from (a c c)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_acc2gac))
  (mutate (from (a c c)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_acc2tac))
  (mutate (from (a c c)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_acc2ccc))
  (mutate (from (a c c)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_acc2gcc))
  (mutate (from (a c c)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_acc2tcc))
  (mutate (from (a c c)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_acc2agc))
  (mutate (from (a c c)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_acc2cgc))
  (mutate (from (a c c)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_acc2ggc))
  (mutate (from (a c c)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_acc2tgc))
  (mutate (from (a c c)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_acc2atc))
  (mutate (from (a c c)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_acc2ctc))
  (mutate (from (a c c)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_acc2gtc))
  (mutate (from (a c c)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_acc2ttc))
  (mutate (from (a c c)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_acc2aag))
  (mutate (from (a c c)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_acc2cag))
  (mutate (from (a c c)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_acc2gag))
  (mutate (from (a c c)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_acc2tag))
  (mutate (from (a c c)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_acc2acg))
  (mutate (from (a c c)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_acc2ccg))
  (mutate (from (a c c)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_acc2gcg))
  (mutate (from (a c c)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_acc2tcg))
  (mutate (from (a c c)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_acc2agg))
  (mutate (from (a c c)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_acc2cgg))
  (mutate (from (a c c)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_acc2ggg))
  (mutate (from (a c c)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_acc2tgg))
  (mutate (from (a c c)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_acc2atg))
  (mutate (from (a c c)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_acc2ctg))
  (mutate (from (a c c)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_acc2gtg))
  (mutate (from (a c c)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_acc2ttg))
  (mutate (from (a c c)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_acc2aat))
  (mutate (from (a c c)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_acc2cat))
  (mutate (from (a c c)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_acc2gat))
  (mutate (from (a c c)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_acc2tat))
  (mutate (from (a c c)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_acc2act))
  (mutate (from (a c c)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_acc2cct))
  (mutate (from (a c c)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_acc2gct))
  (mutate (from (a c c)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_acc2tct))
  (mutate (from (a c c)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_acc2agt))
  (mutate (from (a c c)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_acc2cgt))
  (mutate (from (a c c)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_acc2ggt))
  (mutate (from (a c c)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_acc2tgt))
  (mutate (from (a c c)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_acc2att))
  (mutate (from (a c c)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_acc2ctt))
  (mutate (from (a c c)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_acc2gtt))
  (mutate (from (a c c)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_acc2ttt))
  (mutate (from (c c c)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2aaa))
  (mutate (from (c c c)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2caa))
  (mutate (from (c c c)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2gaa))
  (mutate (from (c c c)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2taa))
  (mutate (from (c c c)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2aca))
  (mutate (from (c c c)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2cca))
  (mutate (from (c c c)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2gca))
  (mutate (from (c c c)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2tca))
  (mutate (from (c c c)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2aga))
  (mutate (from (c c c)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2cga))
  (mutate (from (c c c)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2gga))
  (mutate (from (c c c)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2tga))
  (mutate (from (c c c)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2ata))
  (mutate (from (c c c)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2cta))
  (mutate (from (c c c)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2gta))
  (mutate (from (c c c)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2tta))
  (mutate (from (c c c)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2aac))
  (mutate (from (c c c)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2cac))
  (mutate (from (c c c)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2gac))
  (mutate (from (c c c)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2tac))
  (mutate (from (c c c)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2acc))
  (mutate (from (c c c)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2gcc))
  (mutate (from (c c c)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2tcc))
  (mutate (from (c c c)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2agc))
  (mutate (from (c c c)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2cgc))
  (mutate (from (c c c)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2ggc))
  (mutate (from (c c c)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2tgc))
  (mutate (from (c c c)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2atc))
  (mutate (from (c c c)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2ctc))
  (mutate (from (c c c)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2gtc))
  (mutate (from (c c c)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2ttc))
  (mutate (from (c c c)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2aag))
  (mutate (from (c c c)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2cag))
  (mutate (from (c c c)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2gag))
  (mutate (from (c c c)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2tag))
  (mutate (from (c c c)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2acg))
  (mutate (from (c c c)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2ccg))
  (mutate (from (c c c)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2gcg))
  (mutate (from (c c c)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2tcg))
  (mutate (from (c c c)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2agg))
  (mutate (from (c c c)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2cgg))
  (mutate (from (c c c)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2ggg))
  (mutate (from (c c c)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2tgg))
  (mutate (from (c c c)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2atg))
  (mutate (from (c c c)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2ctg))
  (mutate (from (c c c)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2gtg))
  (mutate (from (c c c)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2ttg))
  (mutate (from (c c c)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2aat))
  (mutate (from (c c c)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2cat))
  (mutate (from (c c c)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2gat))
  (mutate (from (c c c)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2tat))
  (mutate (from (c c c)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2act))
  (mutate (from (c c c)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2cct))
  (mutate (from (c c c)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2gct))
  (mutate (from (c c c)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2tct))
  (mutate (from (c c c)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2agt))
  (mutate (from (c c c)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2cgt))
  (mutate (from (c c c)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2ggt))
  (mutate (from (c c c)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2tgt))
  (mutate (from (c c c)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2att))
  (mutate (from (c c c)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2ctt))
  (mutate (from (c c c)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2gtt))
  (mutate (from (c c c)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_ccc2ttt))
  (mutate (from (g c c)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2aaa))
  (mutate (from (g c c)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2caa))
  (mutate (from (g c c)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2gaa))
  (mutate (from (g c c)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2taa))
  (mutate (from (g c c)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2aca))
  (mutate (from (g c c)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2cca))
  (mutate (from (g c c)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2gca))
  (mutate (from (g c c)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2tca))
  (mutate (from (g c c)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2aga))
  (mutate (from (g c c)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2cga))
  (mutate (from (g c c)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2gga))
  (mutate (from (g c c)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2tga))
  (mutate (from (g c c)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2ata))
  (mutate (from (g c c)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2cta))
  (mutate (from (g c c)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2gta))
  (mutate (from (g c c)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2tta))
  (mutate (from (g c c)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2aac))
  (mutate (from (g c c)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2cac))
  (mutate (from (g c c)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2gac))
  (mutate (from (g c c)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2tac))
  (mutate (from (g c c)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2acc))
  (mutate (from (g c c)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2ccc))
  (mutate (from (g c c)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2tcc))
  (mutate (from (g c c)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2agc))
  (mutate (from (g c c)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2cgc))
  (mutate (from (g c c)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2ggc))
  (mutate (from (g c c)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2tgc))
  (mutate (from (g c c)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2atc))
  (mutate (from (g c c)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2ctc))
  (mutate (from (g c c)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2gtc))
  (mutate (from (g c c)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2ttc))
  (mutate (from (g c c)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2aag))
  (mutate (from (g c c)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2cag))
  (mutate (from (g c c)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2gag))
  (mutate (from (g c c)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2tag))
  (mutate (from (g c c)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2acg))
  (mutate (from (g c c)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2ccg))
  (mutate (from (g c c)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2gcg))
  (mutate (from (g c c)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2tcg))
  (mutate (from (g c c)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2agg))
  (mutate (from (g c c)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2cgg))
  (mutate (from (g c c)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2ggg))
  (mutate (from (g c c)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2tgg))
  (mutate (from (g c c)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2atg))
  (mutate (from (g c c)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2ctg))
  (mutate (from (g c c)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2gtg))
  (mutate (from (g c c)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2ttg))
  (mutate (from (g c c)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2aat))
  (mutate (from (g c c)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2cat))
  (mutate (from (g c c)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2gat))
  (mutate (from (g c c)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2tat))
  (mutate (from (g c c)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2act))
  (mutate (from (g c c)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2cct))
  (mutate (from (g c c)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2gct))
  (mutate (from (g c c)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2tct))
  (mutate (from (g c c)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2agt))
  (mutate (from (g c c)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2cgt))
  (mutate (from (g c c)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2ggt))
  (mutate (from (g c c)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2tgt))
  (mutate (from (g c c)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2att))
  (mutate (from (g c c)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2ctt))
  (mutate (from (g c c)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2gtt))
  (mutate (from (g c c)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_gcc2ttt))
  (mutate (from (t c c)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2aaa))
  (mutate (from (t c c)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2caa))
  (mutate (from (t c c)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2gaa))
  (mutate (from (t c c)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2taa))
  (mutate (from (t c c)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2aca))
  (mutate (from (t c c)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2cca))
  (mutate (from (t c c)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2gca))
  (mutate (from (t c c)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2tca))
  (mutate (from (t c c)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2aga))
  (mutate (from (t c c)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2cga))
  (mutate (from (t c c)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2gga))
  (mutate (from (t c c)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2tga))
  (mutate (from (t c c)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2ata))
  (mutate (from (t c c)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2cta))
  (mutate (from (t c c)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2gta))
  (mutate (from (t c c)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2tta))
  (mutate (from (t c c)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2aac))
  (mutate (from (t c c)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2cac))
  (mutate (from (t c c)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2gac))
  (mutate (from (t c c)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2tac))
  (mutate (from (t c c)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2acc))
  (mutate (from (t c c)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2ccc))
  (mutate (from (t c c)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2gcc))
  (mutate (from (t c c)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2agc))
  (mutate (from (t c c)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2cgc))
  (mutate (from (t c c)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2ggc))
  (mutate (from (t c c)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2tgc))
  (mutate (from (t c c)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2atc))
  (mutate (from (t c c)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2ctc))
  (mutate (from (t c c)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2gtc))
  (mutate (from (t c c)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2ttc))
  (mutate (from (t c c)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2aag))
  (mutate (from (t c c)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2cag))
  (mutate (from (t c c)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2gag))
  (mutate (from (t c c)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2tag))
  (mutate (from (t c c)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2acg))
  (mutate (from (t c c)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2ccg))
  (mutate (from (t c c)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2gcg))
  (mutate (from (t c c)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2tcg))
  (mutate (from (t c c)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2agg))
  (mutate (from (t c c)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2cgg))
  (mutate (from (t c c)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2ggg))
  (mutate (from (t c c)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2tgg))
  (mutate (from (t c c)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2atg))
  (mutate (from (t c c)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2ctg))
  (mutate (from (t c c)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2gtg))
  (mutate (from (t c c)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2ttg))
  (mutate (from (t c c)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2aat))
  (mutate (from (t c c)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2cat))
  (mutate (from (t c c)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2gat))
  (mutate (from (t c c)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2tat))
  (mutate (from (t c c)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2act))
  (mutate (from (t c c)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2cct))
  (mutate (from (t c c)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2gct))
  (mutate (from (t c c)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2tct))
  (mutate (from (t c c)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2agt))
  (mutate (from (t c c)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2cgt))
  (mutate (from (t c c)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2ggt))
  (mutate (from (t c c)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2tgt))
  (mutate (from (t c c)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2att))
  (mutate (from (t c c)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2ctt))
  (mutate (from (t c c)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2gtt))
  (mutate (from (t c c)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_tcc2ttt))
  (mutate (from (a g c)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_agc2aaa))
  (mutate (from (a g c)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_agc2caa))
  (mutate (from (a g c)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_agc2gaa))
  (mutate (from (a g c)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_agc2taa))
  (mutate (from (a g c)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_agc2aca))
  (mutate (from (a g c)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_agc2cca))
  (mutate (from (a g c)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_agc2gca))
  (mutate (from (a g c)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_agc2tca))
  (mutate (from (a g c)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_agc2aga))
  (mutate (from (a g c)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_agc2cga))
  (mutate (from (a g c)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_agc2gga))
  (mutate (from (a g c)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_agc2tga))
  (mutate (from (a g c)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_agc2ata))
  (mutate (from (a g c)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_agc2cta))
  (mutate (from (a g c)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_agc2gta))
  (mutate (from (a g c)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_agc2tta))
  (mutate (from (a g c)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_agc2aac))
  (mutate (from (a g c)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_agc2cac))
  (mutate (from (a g c)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_agc2gac))
  (mutate (from (a g c)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_agc2tac))
  (mutate (from (a g c)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_agc2acc))
  (mutate (from (a g c)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_agc2ccc))
  (mutate (from (a g c)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_agc2gcc))
  (mutate (from (a g c)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_agc2tcc))
  (mutate (from (a g c)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_agc2cgc))
  (mutate (from (a g c)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_agc2ggc))
  (mutate (from (a g c)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_agc2tgc))
  (mutate (from (a g c)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_agc2atc))
  (mutate (from (a g c)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_agc2ctc))
  (mutate (from (a g c)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_agc2gtc))
  (mutate (from (a g c)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_agc2ttc))
  (mutate (from (a g c)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_agc2aag))
  (mutate (from (a g c)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_agc2cag))
  (mutate (from (a g c)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_agc2gag))
  (mutate (from (a g c)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_agc2tag))
  (mutate (from (a g c)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_agc2acg))
  (mutate (from (a g c)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_agc2ccg))
  (mutate (from (a g c)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_agc2gcg))
  (mutate (from (a g c)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_agc2tcg))
  (mutate (from (a g c)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_agc2agg))
  (mutate (from (a g c)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_agc2cgg))
  (mutate (from (a g c)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_agc2ggg))
  (mutate (from (a g c)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_agc2tgg))
  (mutate (from (a g c)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_agc2atg))
  (mutate (from (a g c)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_agc2ctg))
  (mutate (from (a g c)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_agc2gtg))
  (mutate (from (a g c)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_agc2ttg))
  (mutate (from (a g c)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_agc2aat))
  (mutate (from (a g c)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_agc2cat))
  (mutate (from (a g c)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_agc2gat))
  (mutate (from (a g c)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_agc2tat))
  (mutate (from (a g c)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_agc2act))
  (mutate (from (a g c)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_agc2cct))
  (mutate (from (a g c)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_agc2gct))
  (mutate (from (a g c)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_agc2tct))
  (mutate (from (a g c)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_agc2agt))
  (mutate (from (a g c)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_agc2cgt))
  (mutate (from (a g c)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_agc2ggt))
  (mutate (from (a g c)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_agc2tgt))
  (mutate (from (a g c)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_agc2att))
  (mutate (from (a g c)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_agc2ctt))
  (mutate (from (a g c)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_agc2gtt))
  (mutate (from (a g c)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_agc2ttt))
  (mutate (from (c g c)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2aaa))
  (mutate (from (c g c)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2caa))
  (mutate (from (c g c)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2gaa))
  (mutate (from (c g c)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2taa))
  (mutate (from (c g c)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2aca))
  (mutate (from (c g c)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2cca))
  (mutate (from (c g c)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2gca))
  (mutate (from (c g c)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2tca))
  (mutate (from (c g c)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2aga))
  (mutate (from (c g c)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2cga))
  (mutate (from (c g c)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2gga))
  (mutate (from (c g c)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2tga))
  (mutate (from (c g c)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2ata))
  (mutate (from (c g c)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2cta))
  (mutate (from (c g c)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2gta))
  (mutate (from (c g c)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2tta))
  (mutate (from (c g c)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2aac))
  (mutate (from (c g c)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2cac))
  (mutate (from (c g c)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2gac))
  (mutate (from (c g c)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2tac))
  (mutate (from (c g c)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2acc))
  (mutate (from (c g c)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2ccc))
  (mutate (from (c g c)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2gcc))
  (mutate (from (c g c)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2tcc))
  (mutate (from (c g c)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2agc))
  (mutate (from (c g c)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2ggc))
  (mutate (from (c g c)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2tgc))
  (mutate (from (c g c)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2atc))
  (mutate (from (c g c)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2ctc))
  (mutate (from (c g c)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2gtc))
  (mutate (from (c g c)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2ttc))
  (mutate (from (c g c)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2aag))
  (mutate (from (c g c)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2cag))
  (mutate (from (c g c)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2gag))
  (mutate (from (c g c)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2tag))
  (mutate (from (c g c)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2acg))
  (mutate (from (c g c)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2ccg))
  (mutate (from (c g c)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2gcg))
  (mutate (from (c g c)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2tcg))
  (mutate (from (c g c)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2agg))
  (mutate (from (c g c)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2cgg))
  (mutate (from (c g c)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2ggg))
  (mutate (from (c g c)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2tgg))
  (mutate (from (c g c)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2atg))
  (mutate (from (c g c)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2ctg))
  (mutate (from (c g c)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2gtg))
  (mutate (from (c g c)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2ttg))
  (mutate (from (c g c)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2aat))
  (mutate (from (c g c)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2cat))
  (mutate (from (c g c)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2gat))
  (mutate (from (c g c)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2tat))
  (mutate (from (c g c)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2act))
  (mutate (from (c g c)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2cct))
  (mutate (from (c g c)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2gct))
  (mutate (from (c g c)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2tct))
  (mutate (from (c g c)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2agt))
  (mutate (from (c g c)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2cgt))
  (mutate (from (c g c)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2ggt))
  (mutate (from (c g c)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2tgt))
  (mutate (from (c g c)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2att))
  (mutate (from (c g c)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2ctt))
  (mutate (from (c g c)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2gtt))
  (mutate (from (c g c)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_cgc2ttt))
  (mutate (from (g g c)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2aaa))
  (mutate (from (g g c)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2caa))
  (mutate (from (g g c)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2gaa))
  (mutate (from (g g c)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2taa))
  (mutate (from (g g c)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2aca))
  (mutate (from (g g c)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2cca))
  (mutate (from (g g c)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2gca))
  (mutate (from (g g c)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2tca))
  (mutate (from (g g c)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2aga))
  (mutate (from (g g c)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2cga))
  (mutate (from (g g c)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2gga))
  (mutate (from (g g c)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2tga))
  (mutate (from (g g c)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2ata))
  (mutate (from (g g c)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2cta))
  (mutate (from (g g c)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2gta))
  (mutate (from (g g c)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2tta))
  (mutate (from (g g c)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2aac))
  (mutate (from (g g c)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2cac))
  (mutate (from (g g c)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2gac))
  (mutate (from (g g c)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2tac))
  (mutate (from (g g c)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2acc))
  (mutate (from (g g c)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2ccc))
  (mutate (from (g g c)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2gcc))
  (mutate (from (g g c)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2tcc))
  (mutate (from (g g c)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2agc))
  (mutate (from (g g c)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2cgc))
  (mutate (from (g g c)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2tgc))
  (mutate (from (g g c)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2atc))
  (mutate (from (g g c)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2ctc))
  (mutate (from (g g c)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2gtc))
  (mutate (from (g g c)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2ttc))
  (mutate (from (g g c)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2aag))
  (mutate (from (g g c)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2cag))
  (mutate (from (g g c)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2gag))
  (mutate (from (g g c)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2tag))
  (mutate (from (g g c)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2acg))
  (mutate (from (g g c)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2ccg))
  (mutate (from (g g c)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2gcg))
  (mutate (from (g g c)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2tcg))
  (mutate (from (g g c)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2agg))
  (mutate (from (g g c)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2cgg))
  (mutate (from (g g c)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2ggg))
  (mutate (from (g g c)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2tgg))
  (mutate (from (g g c)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2atg))
  (mutate (from (g g c)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2ctg))
  (mutate (from (g g c)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2gtg))
  (mutate (from (g g c)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2ttg))
  (mutate (from (g g c)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2aat))
  (mutate (from (g g c)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2cat))
  (mutate (from (g g c)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2gat))
  (mutate (from (g g c)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2tat))
  (mutate (from (g g c)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2act))
  (mutate (from (g g c)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2cct))
  (mutate (from (g g c)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2gct))
  (mutate (from (g g c)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2tct))
  (mutate (from (g g c)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2agt))
  (mutate (from (g g c)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2cgt))
  (mutate (from (g g c)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2ggt))
  (mutate (from (g g c)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2tgt))
  (mutate (from (g g c)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2att))
  (mutate (from (g g c)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2ctt))
  (mutate (from (g g c)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2gtt))
  (mutate (from (g g c)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_ggc2ttt))
  (mutate (from (t g c)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2aaa))
  (mutate (from (t g c)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2caa))
  (mutate (from (t g c)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2gaa))
  (mutate (from (t g c)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2taa))
  (mutate (from (t g c)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2aca))
  (mutate (from (t g c)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2cca))
  (mutate (from (t g c)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2gca))
  (mutate (from (t g c)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2tca))
  (mutate (from (t g c)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2aga))
  (mutate (from (t g c)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2cga))
  (mutate (from (t g c)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2gga))
  (mutate (from (t g c)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2tga))
  (mutate (from (t g c)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2ata))
  (mutate (from (t g c)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2cta))
  (mutate (from (t g c)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2gta))
  (mutate (from (t g c)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2tta))
  (mutate (from (t g c)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2aac))
  (mutate (from (t g c)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2cac))
  (mutate (from (t g c)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2gac))
  (mutate (from (t g c)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2tac))
  (mutate (from (t g c)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2acc))
  (mutate (from (t g c)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2ccc))
  (mutate (from (t g c)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2gcc))
  (mutate (from (t g c)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2tcc))
  (mutate (from (t g c)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2agc))
  (mutate (from (t g c)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2cgc))
  (mutate (from (t g c)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2ggc))
  (mutate (from (t g c)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2atc))
  (mutate (from (t g c)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2ctc))
  (mutate (from (t g c)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2gtc))
  (mutate (from (t g c)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2ttc))
  (mutate (from (t g c)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2aag))
  (mutate (from (t g c)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2cag))
  (mutate (from (t g c)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2gag))
  (mutate (from (t g c)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2tag))
  (mutate (from (t g c)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2acg))
  (mutate (from (t g c)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2ccg))
  (mutate (from (t g c)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2gcg))
  (mutate (from (t g c)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2tcg))
  (mutate (from (t g c)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2agg))
  (mutate (from (t g c)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2cgg))
  (mutate (from (t g c)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2ggg))
  (mutate (from (t g c)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2tgg))
  (mutate (from (t g c)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2atg))
  (mutate (from (t g c)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2ctg))
  (mutate (from (t g c)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2gtg))
  (mutate (from (t g c)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2ttg))
  (mutate (from (t g c)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2aat))
  (mutate (from (t g c)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2cat))
  (mutate (from (t g c)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2gat))
  (mutate (from (t g c)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2tat))
  (mutate (from (t g c)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2act))
  (mutate (from (t g c)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2cct))
  (mutate (from (t g c)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2gct))
  (mutate (from (t g c)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2tct))
  (mutate (from (t g c)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2agt))
  (mutate (from (t g c)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2cgt))
  (mutate (from (t g c)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2ggt))
  (mutate (from (t g c)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2tgt))
  (mutate (from (t g c)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2att))
  (mutate (from (t g c)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2ctt))
  (mutate (from (t g c)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2gtt))
  (mutate (from (t g c)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_tgc2ttt))
  (mutate (from (a t c)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_atc2aaa))
  (mutate (from (a t c)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_atc2caa))
  (mutate (from (a t c)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_atc2gaa))
  (mutate (from (a t c)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_atc2taa))
  (mutate (from (a t c)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_atc2aca))
  (mutate (from (a t c)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_atc2cca))
  (mutate (from (a t c)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_atc2gca))
  (mutate (from (a t c)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_atc2tca))
  (mutate (from (a t c)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_atc2aga))
  (mutate (from (a t c)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_atc2cga))
  (mutate (from (a t c)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_atc2gga))
  (mutate (from (a t c)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_atc2tga))
  (mutate (from (a t c)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_atc2ata))
  (mutate (from (a t c)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_atc2cta))
  (mutate (from (a t c)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_atc2gta))
  (mutate (from (a t c)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_atc2tta))
  (mutate (from (a t c)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_atc2aac))
  (mutate (from (a t c)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_atc2cac))
  (mutate (from (a t c)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_atc2gac))
  (mutate (from (a t c)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_atc2tac))
  (mutate (from (a t c)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_atc2acc))
  (mutate (from (a t c)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_atc2ccc))
  (mutate (from (a t c)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_atc2gcc))
  (mutate (from (a t c)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_atc2tcc))
  (mutate (from (a t c)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_atc2agc))
  (mutate (from (a t c)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_atc2cgc))
  (mutate (from (a t c)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_atc2ggc))
  (mutate (from (a t c)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_atc2tgc))
  (mutate (from (a t c)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_atc2ctc))
  (mutate (from (a t c)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_atc2gtc))
  (mutate (from (a t c)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_atc2ttc))
  (mutate (from (a t c)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_atc2aag))
  (mutate (from (a t c)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_atc2cag))
  (mutate (from (a t c)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_atc2gag))
  (mutate (from (a t c)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_atc2tag))
  (mutate (from (a t c)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_atc2acg))
  (mutate (from (a t c)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_atc2ccg))
  (mutate (from (a t c)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_atc2gcg))
  (mutate (from (a t c)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_atc2tcg))
  (mutate (from (a t c)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_atc2agg))
  (mutate (from (a t c)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_atc2cgg))
  (mutate (from (a t c)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_atc2ggg))
  (mutate (from (a t c)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_atc2tgg))
  (mutate (from (a t c)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_atc2atg))
  (mutate (from (a t c)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_atc2ctg))
  (mutate (from (a t c)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_atc2gtg))
  (mutate (from (a t c)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_atc2ttg))
  (mutate (from (a t c)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_atc2aat))
  (mutate (from (a t c)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_atc2cat))
  (mutate (from (a t c)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_atc2gat))
  (mutate (from (a t c)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_atc2tat))
  (mutate (from (a t c)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_atc2act))
  (mutate (from (a t c)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_atc2cct))
  (mutate (from (a t c)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_atc2gct))
  (mutate (from (a t c)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_atc2tct))
  (mutate (from (a t c)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_atc2agt))
  (mutate (from (a t c)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_atc2cgt))
  (mutate (from (a t c)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_atc2ggt))
  (mutate (from (a t c)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_atc2tgt))
  (mutate (from (a t c)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_atc2att))
  (mutate (from (a t c)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_atc2ctt))
  (mutate (from (a t c)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_atc2gtt))
  (mutate (from (a t c)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_atc2ttt))
  (mutate (from (c t c)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2aaa))
  (mutate (from (c t c)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2caa))
  (mutate (from (c t c)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2gaa))
  (mutate (from (c t c)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2taa))
  (mutate (from (c t c)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2aca))
  (mutate (from (c t c)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2cca))
  (mutate (from (c t c)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2gca))
  (mutate (from (c t c)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2tca))
  (mutate (from (c t c)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2aga))
  (mutate (from (c t c)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2cga))
  (mutate (from (c t c)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2gga))
  (mutate (from (c t c)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2tga))
  (mutate (from (c t c)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2ata))
  (mutate (from (c t c)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2cta))
  (mutate (from (c t c)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2gta))
  (mutate (from (c t c)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2tta))
  (mutate (from (c t c)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2aac))
  (mutate (from (c t c)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2cac))
  (mutate (from (c t c)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2gac))
  (mutate (from (c t c)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2tac))
  (mutate (from (c t c)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2acc))
  (mutate (from (c t c)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2ccc))
  (mutate (from (c t c)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2gcc))
  (mutate (from (c t c)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2tcc))
  (mutate (from (c t c)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2agc))
  (mutate (from (c t c)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2cgc))
  (mutate (from (c t c)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2ggc))
  (mutate (from (c t c)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2tgc))
  (mutate (from (c t c)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2atc))
  (mutate (from (c t c)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2gtc))
  (mutate (from (c t c)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2ttc))
  (mutate (from (c t c)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2aag))
  (mutate (from (c t c)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2cag))
  (mutate (from (c t c)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2gag))
  (mutate (from (c t c)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2tag))
  (mutate (from (c t c)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2acg))
  (mutate (from (c t c)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2ccg))
  (mutate (from (c t c)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2gcg))
  (mutate (from (c t c)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2tcg))
  (mutate (from (c t c)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2agg))
  (mutate (from (c t c)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2cgg))
  (mutate (from (c t c)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2ggg))
  (mutate (from (c t c)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2tgg))
  (mutate (from (c t c)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2atg))
  (mutate (from (c t c)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2ctg))
  (mutate (from (c t c)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2gtg))
  (mutate (from (c t c)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2ttg))
  (mutate (from (c t c)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2aat))
  (mutate (from (c t c)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2cat))
  (mutate (from (c t c)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2gat))
  (mutate (from (c t c)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2tat))
  (mutate (from (c t c)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2act))
  (mutate (from (c t c)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2cct))
  (mutate (from (c t c)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2gct))
  (mutate (from (c t c)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2tct))
  (mutate (from (c t c)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2agt))
  (mutate (from (c t c)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2cgt))
  (mutate (from (c t c)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2ggt))
  (mutate (from (c t c)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2tgt))
  (mutate (from (c t c)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2att))
  (mutate (from (c t c)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2ctt))
  (mutate (from (c t c)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2gtt))
  (mutate (from (c t c)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_ctc2ttt))
  (mutate (from (g t c)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2aaa))
  (mutate (from (g t c)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2caa))
  (mutate (from (g t c)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2gaa))
  (mutate (from (g t c)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2taa))
  (mutate (from (g t c)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2aca))
  (mutate (from (g t c)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2cca))
  (mutate (from (g t c)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2gca))
  (mutate (from (g t c)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2tca))
  (mutate (from (g t c)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2aga))
  (mutate (from (g t c)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2cga))
  (mutate (from (g t c)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2gga))
  (mutate (from (g t c)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2tga))
  (mutate (from (g t c)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2ata))
  (mutate (from (g t c)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2cta))
  (mutate (from (g t c)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2gta))
  (mutate (from (g t c)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2tta))
  (mutate (from (g t c)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2aac))
  (mutate (from (g t c)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2cac))
  (mutate (from (g t c)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2gac))
  (mutate (from (g t c)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2tac))
  (mutate (from (g t c)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2acc))
  (mutate (from (g t c)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2ccc))
  (mutate (from (g t c)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2gcc))
  (mutate (from (g t c)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2tcc))
  (mutate (from (g t c)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2agc))
  (mutate (from (g t c)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2cgc))
  (mutate (from (g t c)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2ggc))
  (mutate (from (g t c)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2tgc))
  (mutate (from (g t c)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2atc))
  (mutate (from (g t c)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2ctc))
  (mutate (from (g t c)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2ttc))
  (mutate (from (g t c)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2aag))
  (mutate (from (g t c)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2cag))
  (mutate (from (g t c)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2gag))
  (mutate (from (g t c)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2tag))
  (mutate (from (g t c)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2acg))
  (mutate (from (g t c)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2ccg))
  (mutate (from (g t c)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2gcg))
  (mutate (from (g t c)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2tcg))
  (mutate (from (g t c)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2agg))
  (mutate (from (g t c)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2cgg))
  (mutate (from (g t c)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2ggg))
  (mutate (from (g t c)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2tgg))
  (mutate (from (g t c)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2atg))
  (mutate (from (g t c)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2ctg))
  (mutate (from (g t c)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2gtg))
  (mutate (from (g t c)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2ttg))
  (mutate (from (g t c)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2aat))
  (mutate (from (g t c)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2cat))
  (mutate (from (g t c)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2gat))
  (mutate (from (g t c)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2tat))
  (mutate (from (g t c)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2act))
  (mutate (from (g t c)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2cct))
  (mutate (from (g t c)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2gct))
  (mutate (from (g t c)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2tct))
  (mutate (from (g t c)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2agt))
  (mutate (from (g t c)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2cgt))
  (mutate (from (g t c)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2ggt))
  (mutate (from (g t c)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2tgt))
  (mutate (from (g t c)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2att))
  (mutate (from (g t c)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2ctt))
  (mutate (from (g t c)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2gtt))
  (mutate (from (g t c)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_gtc2ttt))
  (mutate (from (t t c)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2aaa))
  (mutate (from (t t c)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2caa))
  (mutate (from (t t c)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2gaa))
  (mutate (from (t t c)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2taa))
  (mutate (from (t t c)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2aca))
  (mutate (from (t t c)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2cca))
  (mutate (from (t t c)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2gca))
  (mutate (from (t t c)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2tca))
  (mutate (from (t t c)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2aga))
  (mutate (from (t t c)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2cga))
  (mutate (from (t t c)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2gga))
  (mutate (from (t t c)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2tga))
  (mutate (from (t t c)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2ata))
  (mutate (from (t t c)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2cta))
  (mutate (from (t t c)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2gta))
  (mutate (from (t t c)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2tta))
  (mutate (from (t t c)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2aac))
  (mutate (from (t t c)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2cac))
  (mutate (from (t t c)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2gac))
  (mutate (from (t t c)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2tac))
  (mutate (from (t t c)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2acc))
  (mutate (from (t t c)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2ccc))
  (mutate (from (t t c)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2gcc))
  (mutate (from (t t c)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2tcc))
  (mutate (from (t t c)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2agc))
  (mutate (from (t t c)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2cgc))
  (mutate (from (t t c)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2ggc))
  (mutate (from (t t c)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2tgc))
  (mutate (from (t t c)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2atc))
  (mutate (from (t t c)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2ctc))
  (mutate (from (t t c)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2gtc))
  (mutate (from (t t c)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2aag))
  (mutate (from (t t c)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2cag))
  (mutate (from (t t c)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2gag))
  (mutate (from (t t c)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2tag))
  (mutate (from (t t c)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2acg))
  (mutate (from (t t c)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2ccg))
  (mutate (from (t t c)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2gcg))
  (mutate (from (t t c)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2tcg))
  (mutate (from (t t c)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2agg))
  (mutate (from (t t c)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2cgg))
  (mutate (from (t t c)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2ggg))
  (mutate (from (t t c)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2tgg))
  (mutate (from (t t c)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2atg))
  (mutate (from (t t c)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2ctg))
  (mutate (from (t t c)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2gtg))
  (mutate (from (t t c)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2ttg))
  (mutate (from (t t c)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2aat))
  (mutate (from (t t c)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2cat))
  (mutate (from (t t c)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2gat))
  (mutate (from (t t c)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2tat))
  (mutate (from (t t c)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2act))
  (mutate (from (t t c)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2cct))
  (mutate (from (t t c)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2gct))
  (mutate (from (t t c)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2tct))
  (mutate (from (t t c)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2agt))
  (mutate (from (t t c)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2cgt))
  (mutate (from (t t c)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2ggt))
  (mutate (from (t t c)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2tgt))
  (mutate (from (t t c)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2att))
  (mutate (from (t t c)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2ctt))
  (mutate (from (t t c)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2gtt))
  (mutate (from (t t c)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_ttc2ttt))
  (mutate (from (a a g)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_aag2aaa))
  (mutate (from (a a g)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_aag2caa))
  (mutate (from (a a g)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_aag2gaa))
  (mutate (from (a a g)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_aag2taa))
  (mutate (from (a a g)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_aag2aca))
  (mutate (from (a a g)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_aag2cca))
  (mutate (from (a a g)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_aag2gca))
  (mutate (from (a a g)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_aag2tca))
  (mutate (from (a a g)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_aag2aga))
  (mutate (from (a a g)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_aag2cga))
  (mutate (from (a a g)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_aag2gga))
  (mutate (from (a a g)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_aag2tga))
  (mutate (from (a a g)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_aag2ata))
  (mutate (from (a a g)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_aag2cta))
  (mutate (from (a a g)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_aag2gta))
  (mutate (from (a a g)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_aag2tta))
  (mutate (from (a a g)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_aag2aac))
  (mutate (from (a a g)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_aag2cac))
  (mutate (from (a a g)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_aag2gac))
  (mutate (from (a a g)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_aag2tac))
  (mutate (from (a a g)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_aag2acc))
  (mutate (from (a a g)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_aag2ccc))
  (mutate (from (a a g)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_aag2gcc))
  (mutate (from (a a g)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_aag2tcc))
  (mutate (from (a a g)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_aag2agc))
  (mutate (from (a a g)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_aag2cgc))
  (mutate (from (a a g)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_aag2ggc))
  (mutate (from (a a g)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_aag2tgc))
  (mutate (from (a a g)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_aag2atc))
  (mutate (from (a a g)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_aag2ctc))
  (mutate (from (a a g)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_aag2gtc))
  (mutate (from (a a g)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_aag2ttc))
  (mutate (from (a a g)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_aag2cag))
  (mutate (from (a a g)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_aag2gag))
  (mutate (from (a a g)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_aag2tag))
  (mutate (from (a a g)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_aag2acg))
  (mutate (from (a a g)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_aag2ccg))
  (mutate (from (a a g)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_aag2gcg))
  (mutate (from (a a g)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_aag2tcg))
  (mutate (from (a a g)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_aag2agg))
  (mutate (from (a a g)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_aag2cgg))
  (mutate (from (a a g)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_aag2ggg))
  (mutate (from (a a g)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_aag2tgg))
  (mutate (from (a a g)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_aag2atg))
  (mutate (from (a a g)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_aag2ctg))
  (mutate (from (a a g)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_aag2gtg))
  (mutate (from (a a g)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_aag2ttg))
  (mutate (from (a a g)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_aag2aat))
  (mutate (from (a a g)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_aag2cat))
  (mutate (from (a a g)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_aag2gat))
  (mutate (from (a a g)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_aag2tat))
  (mutate (from (a a g)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_aag2act))
  (mutate (from (a a g)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_aag2cct))
  (mutate (from (a a g)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_aag2gct))
  (mutate (from (a a g)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_aag2tct))
  (mutate (from (a a g)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_aag2agt))
  (mutate (from (a a g)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_aag2cgt))
  (mutate (from (a a g)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_aag2ggt))
  (mutate (from (a a g)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_aag2tgt))
  (mutate (from (a a g)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_aag2att))
  (mutate (from (a a g)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_aag2ctt))
  (mutate (from (a a g)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_aag2gtt))
  (mutate (from (a a g)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_aag2ttt))
  (mutate (from (c a g)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_cag2aaa))
  (mutate (from (c a g)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_cag2caa))
  (mutate (from (c a g)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_cag2gaa))
  (mutate (from (c a g)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_cag2taa))
  (mutate (from (c a g)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_cag2aca))
  (mutate (from (c a g)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_cag2cca))
  (mutate (from (c a g)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_cag2gca))
  (mutate (from (c a g)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_cag2tca))
  (mutate (from (c a g)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_cag2aga))
  (mutate (from (c a g)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_cag2cga))
  (mutate (from (c a g)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_cag2gga))
  (mutate (from (c a g)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_cag2tga))
  (mutate (from (c a g)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_cag2ata))
  (mutate (from (c a g)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_cag2cta))
  (mutate (from (c a g)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_cag2gta))
  (mutate (from (c a g)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_cag2tta))
  (mutate (from (c a g)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_cag2aac))
  (mutate (from (c a g)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_cag2cac))
  (mutate (from (c a g)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_cag2gac))
  (mutate (from (c a g)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_cag2tac))
  (mutate (from (c a g)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_cag2acc))
  (mutate (from (c a g)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_cag2ccc))
  (mutate (from (c a g)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_cag2gcc))
  (mutate (from (c a g)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_cag2tcc))
  (mutate (from (c a g)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_cag2agc))
  (mutate (from (c a g)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_cag2cgc))
  (mutate (from (c a g)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_cag2ggc))
  (mutate (from (c a g)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_cag2tgc))
  (mutate (from (c a g)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_cag2atc))
  (mutate (from (c a g)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_cag2ctc))
  (mutate (from (c a g)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_cag2gtc))
  (mutate (from (c a g)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_cag2ttc))
  (mutate (from (c a g)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_cag2aag))
  (mutate (from (c a g)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_cag2gag))
  (mutate (from (c a g)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_cag2tag))
  (mutate (from (c a g)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_cag2acg))
  (mutate (from (c a g)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_cag2ccg))
  (mutate (from (c a g)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_cag2gcg))
  (mutate (from (c a g)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_cag2tcg))
  (mutate (from (c a g)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_cag2agg))
  (mutate (from (c a g)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_cag2cgg))
  (mutate (from (c a g)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_cag2ggg))
  (mutate (from (c a g)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_cag2tgg))
  (mutate (from (c a g)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_cag2atg))
  (mutate (from (c a g)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_cag2ctg))
  (mutate (from (c a g)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_cag2gtg))
  (mutate (from (c a g)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_cag2ttg))
  (mutate (from (c a g)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_cag2aat))
  (mutate (from (c a g)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_cag2cat))
  (mutate (from (c a g)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_cag2gat))
  (mutate (from (c a g)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_cag2tat))
  (mutate (from (c a g)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_cag2act))
  (mutate (from (c a g)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_cag2cct))
  (mutate (from (c a g)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_cag2gct))
  (mutate (from (c a g)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_cag2tct))
  (mutate (from (c a g)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_cag2agt))
  (mutate (from (c a g)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_cag2cgt))
  (mutate (from (c a g)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_cag2ggt))
  (mutate (from (c a g)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_cag2tgt))
  (mutate (from (c a g)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_cag2att))
  (mutate (from (c a g)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_cag2ctt))
  (mutate (from (c a g)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_cag2gtt))
  (mutate (from (c a g)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_cag2ttt))
  (mutate (from (g a g)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_gag2aaa))
  (mutate (from (g a g)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_gag2caa))
  (mutate (from (g a g)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_gag2gaa))
  (mutate (from (g a g)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_gag2taa))
  (mutate (from (g a g)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_gag2aca))
  (mutate (from (g a g)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_gag2cca))
  (mutate (from (g a g)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_gag2gca))
  (mutate (from (g a g)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_gag2tca))
  (mutate (from (g a g)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_gag2aga))
  (mutate (from (g a g)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_gag2cga))
  (mutate (from (g a g)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_gag2gga))
  (mutate (from (g a g)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_gag2tga))
  (mutate (from (g a g)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_gag2ata))
  (mutate (from (g a g)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_gag2cta))
  (mutate (from (g a g)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_gag2gta))
  (mutate (from (g a g)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_gag2tta))
  (mutate (from (g a g)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_gag2aac))
  (mutate (from (g a g)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_gag2cac))
  (mutate (from (g a g)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_gag2gac))
  (mutate (from (g a g)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_gag2tac))
  (mutate (from (g a g)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_gag2acc))
  (mutate (from (g a g)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_gag2ccc))
  (mutate (from (g a g)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_gag2gcc))
  (mutate (from (g a g)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_gag2tcc))
  (mutate (from (g a g)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_gag2agc))
  (mutate (from (g a g)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_gag2cgc))
  (mutate (from (g a g)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_gag2ggc))
  (mutate (from (g a g)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_gag2tgc))
  (mutate (from (g a g)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_gag2atc))
  (mutate (from (g a g)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_gag2ctc))
  (mutate (from (g a g)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_gag2gtc))
  (mutate (from (g a g)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_gag2ttc))
  (mutate (from (g a g)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_gag2aag))
  (mutate (from (g a g)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_gag2cag))
  (mutate (from (g a g)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_gag2tag))
  (mutate (from (g a g)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_gag2acg))
  (mutate (from (g a g)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_gag2ccg))
  (mutate (from (g a g)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_gag2gcg))
  (mutate (from (g a g)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_gag2tcg))
  (mutate (from (g a g)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_gag2agg))
  (mutate (from (g a g)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_gag2cgg))
  (mutate (from (g a g)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_gag2ggg))
  (mutate (from (g a g)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_gag2tgg))
  (mutate (from (g a g)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_gag2atg))
  (mutate (from (g a g)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_gag2ctg))
  (mutate (from (g a g)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_gag2gtg))
  (mutate (from (g a g)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_gag2ttg))
  (mutate (from (g a g)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_gag2aat))
  (mutate (from (g a g)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_gag2cat))
  (mutate (from (g a g)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_gag2gat))
  (mutate (from (g a g)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_gag2tat))
  (mutate (from (g a g)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_gag2act))
  (mutate (from (g a g)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_gag2cct))
  (mutate (from (g a g)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_gag2gct))
  (mutate (from (g a g)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_gag2tct))
  (mutate (from (g a g)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_gag2agt))
  (mutate (from (g a g)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_gag2cgt))
  (mutate (from (g a g)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_gag2ggt))
  (mutate (from (g a g)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_gag2tgt))
  (mutate (from (g a g)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_gag2att))
  (mutate (from (g a g)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_gag2ctt))
  (mutate (from (g a g)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_gag2gtt))
  (mutate (from (g a g)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_gag2ttt))
  (mutate (from (t a g)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_tag2aaa))
  (mutate (from (t a g)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_tag2caa))
  (mutate (from (t a g)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_tag2gaa))
  (mutate (from (t a g)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_tag2taa))
  (mutate (from (t a g)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_tag2aca))
  (mutate (from (t a g)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_tag2cca))
  (mutate (from (t a g)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_tag2gca))
  (mutate (from (t a g)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_tag2tca))
  (mutate (from (t a g)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_tag2aga))
  (mutate (from (t a g)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_tag2cga))
  (mutate (from (t a g)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_tag2gga))
  (mutate (from (t a g)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_tag2tga))
  (mutate (from (t a g)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_tag2ata))
  (mutate (from (t a g)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_tag2cta))
  (mutate (from (t a g)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_tag2gta))
  (mutate (from (t a g)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_tag2tta))
  (mutate (from (t a g)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_tag2aac))
  (mutate (from (t a g)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_tag2cac))
  (mutate (from (t a g)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_tag2gac))
  (mutate (from (t a g)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_tag2tac))
  (mutate (from (t a g)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_tag2acc))
  (mutate (from (t a g)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_tag2ccc))
  (mutate (from (t a g)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_tag2gcc))
  (mutate (from (t a g)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_tag2tcc))
  (mutate (from (t a g)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_tag2agc))
  (mutate (from (t a g)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_tag2cgc))
  (mutate (from (t a g)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_tag2ggc))
  (mutate (from (t a g)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_tag2tgc))
  (mutate (from (t a g)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_tag2atc))
  (mutate (from (t a g)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_tag2ctc))
  (mutate (from (t a g)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_tag2gtc))
  (mutate (from (t a g)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_tag2ttc))
  (mutate (from (t a g)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_tag2aag))
  (mutate (from (t a g)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_tag2cag))
  (mutate (from (t a g)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_tag2gag))
  (mutate (from (t a g)) (to (t a g)) (rate 0))
  (mutate (from (t a g)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_tag2acg))
  (mutate (from (t a g)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_tag2ccg))
  (mutate (from (t a g)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_tag2gcg))
  (mutate (from (t a g)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_tag2tcg))
  (mutate (from (t a g)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_tag2agg))
  (mutate (from (t a g)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_tag2cgg))
  (mutate (from (t a g)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_tag2ggg))
  (mutate (from (t a g)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_tag2tgg))
  (mutate (from (t a g)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_tag2atg))
  (mutate (from (t a g)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_tag2ctg))
  (mutate (from (t a g)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_tag2gtg))
  (mutate (from (t a g)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_tag2ttg))
  (mutate (from (t a g)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_tag2aat))
  (mutate (from (t a g)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_tag2cat))
  (mutate (from (t a g)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_tag2gat))
  (mutate (from (t a g)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_tag2tat))
  (mutate (from (t a g)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_tag2act))
  (mutate (from (t a g)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_tag2cct))
  (mutate (from (t a g)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_tag2gct))
  (mutate (from (t a g)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_tag2tct))
  (mutate (from (t a g)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_tag2agt))
  (mutate (from (t a g)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_tag2cgt))
  (mutate (from (t a g)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_tag2ggt))
  (mutate (from (t a g)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_tag2tgt))
  (mutate (from (t a g)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_tag2att))
  (mutate (from (t a g)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_tag2ctt))
  (mutate (from (t a g)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_tag2gtt))
  (mutate (from (t a g)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_tag2ttt))
  (mutate (from (a c g)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_acg2aaa))
  (mutate (from (a c g)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_acg2caa))
  (mutate (from (a c g)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_acg2gaa))
  (mutate (from (a c g)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_acg2taa))
  (mutate (from (a c g)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_acg2aca))
  (mutate (from (a c g)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_acg2cca))
  (mutate (from (a c g)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_acg2gca))
  (mutate (from (a c g)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_acg2tca))
  (mutate (from (a c g)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_acg2aga))
  (mutate (from (a c g)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_acg2cga))
  (mutate (from (a c g)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_acg2gga))
  (mutate (from (a c g)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_acg2tga))
  (mutate (from (a c g)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_acg2ata))
  (mutate (from (a c g)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_acg2cta))
  (mutate (from (a c g)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_acg2gta))
  (mutate (from (a c g)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_acg2tta))
  (mutate (from (a c g)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_acg2aac))
  (mutate (from (a c g)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_acg2cac))
  (mutate (from (a c g)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_acg2gac))
  (mutate (from (a c g)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_acg2tac))
  (mutate (from (a c g)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_acg2acc))
  (mutate (from (a c g)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_acg2ccc))
  (mutate (from (a c g)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_acg2gcc))
  (mutate (from (a c g)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_acg2tcc))
  (mutate (from (a c g)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_acg2agc))
  (mutate (from (a c g)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_acg2cgc))
  (mutate (from (a c g)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_acg2ggc))
  (mutate (from (a c g)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_acg2tgc))
  (mutate (from (a c g)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_acg2atc))
  (mutate (from (a c g)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_acg2ctc))
  (mutate (from (a c g)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_acg2gtc))
  (mutate (from (a c g)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_acg2ttc))
  (mutate (from (a c g)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_acg2aag))
  (mutate (from (a c g)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_acg2cag))
  (mutate (from (a c g)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_acg2gag))
  (mutate (from (a c g)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_acg2tag))
  (mutate (from (a c g)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_acg2ccg))
  (mutate (from (a c g)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_acg2gcg))
  (mutate (from (a c g)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_acg2tcg))
  (mutate (from (a c g)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_acg2agg))
  (mutate (from (a c g)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_acg2cgg))
  (mutate (from (a c g)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_acg2ggg))
  (mutate (from (a c g)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_acg2tgg))
  (mutate (from (a c g)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_acg2atg))
  (mutate (from (a c g)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_acg2ctg))
  (mutate (from (a c g)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_acg2gtg))
  (mutate (from (a c g)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_acg2ttg))
  (mutate (from (a c g)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_acg2aat))
  (mutate (from (a c g)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_acg2cat))
  (mutate (from (a c g)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_acg2gat))
  (mutate (from (a c g)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_acg2tat))
  (mutate (from (a c g)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_acg2act))
  (mutate (from (a c g)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_acg2cct))
  (mutate (from (a c g)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_acg2gct))
  (mutate (from (a c g)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_acg2tct))
  (mutate (from (a c g)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_acg2agt))
  (mutate (from (a c g)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_acg2cgt))
  (mutate (from (a c g)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_acg2ggt))
  (mutate (from (a c g)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_acg2tgt))
  (mutate (from (a c g)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_acg2att))
  (mutate (from (a c g)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_acg2ctt))
  (mutate (from (a c g)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_acg2gtt))
  (mutate (from (a c g)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_acg2ttt))
  (mutate (from (c c g)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2aaa))
  (mutate (from (c c g)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2caa))
  (mutate (from (c c g)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2gaa))
  (mutate (from (c c g)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2taa))
  (mutate (from (c c g)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2aca))
  (mutate (from (c c g)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2cca))
  (mutate (from (c c g)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2gca))
  (mutate (from (c c g)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2tca))
  (mutate (from (c c g)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2aga))
  (mutate (from (c c g)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2cga))
  (mutate (from (c c g)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2gga))
  (mutate (from (c c g)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2tga))
  (mutate (from (c c g)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2ata))
  (mutate (from (c c g)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2cta))
  (mutate (from (c c g)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2gta))
  (mutate (from (c c g)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2tta))
  (mutate (from (c c g)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2aac))
  (mutate (from (c c g)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2cac))
  (mutate (from (c c g)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2gac))
  (mutate (from (c c g)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2tac))
  (mutate (from (c c g)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2acc))
  (mutate (from (c c g)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2ccc))
  (mutate (from (c c g)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2gcc))
  (mutate (from (c c g)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2tcc))
  (mutate (from (c c g)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2agc))
  (mutate (from (c c g)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2cgc))
  (mutate (from (c c g)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2ggc))
  (mutate (from (c c g)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2tgc))
  (mutate (from (c c g)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2atc))
  (mutate (from (c c g)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2ctc))
  (mutate (from (c c g)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2gtc))
  (mutate (from (c c g)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2ttc))
  (mutate (from (c c g)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2aag))
  (mutate (from (c c g)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2cag))
  (mutate (from (c c g)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2gag))
  (mutate (from (c c g)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2tag))
  (mutate (from (c c g)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2acg))
  (mutate (from (c c g)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2gcg))
  (mutate (from (c c g)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2tcg))
  (mutate (from (c c g)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2agg))
  (mutate (from (c c g)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2cgg))
  (mutate (from (c c g)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2ggg))
  (mutate (from (c c g)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2tgg))
  (mutate (from (c c g)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2atg))
  (mutate (from (c c g)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2ctg))
  (mutate (from (c c g)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2gtg))
  (mutate (from (c c g)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2ttg))
  (mutate (from (c c g)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2aat))
  (mutate (from (c c g)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2cat))
  (mutate (from (c c g)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2gat))
  (mutate (from (c c g)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2tat))
  (mutate (from (c c g)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2act))
  (mutate (from (c c g)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2cct))
  (mutate (from (c c g)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2gct))
  (mutate (from (c c g)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2tct))
  (mutate (from (c c g)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2agt))
  (mutate (from (c c g)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2cgt))
  (mutate (from (c c g)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2ggt))
  (mutate (from (c c g)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2tgt))
  (mutate (from (c c g)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2att))
  (mutate (from (c c g)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2ctt))
  (mutate (from (c c g)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2gtt))
  (mutate (from (c c g)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_ccg2ttt))
  (mutate (from (g c g)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2aaa))
  (mutate (from (g c g)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2caa))
  (mutate (from (g c g)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2gaa))
  (mutate (from (g c g)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2taa))
  (mutate (from (g c g)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2aca))
  (mutate (from (g c g)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2cca))
  (mutate (from (g c g)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2gca))
  (mutate (from (g c g)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2tca))
  (mutate (from (g c g)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2aga))
  (mutate (from (g c g)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2cga))
  (mutate (from (g c g)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2gga))
  (mutate (from (g c g)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2tga))
  (mutate (from (g c g)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2ata))
  (mutate (from (g c g)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2cta))
  (mutate (from (g c g)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2gta))
  (mutate (from (g c g)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2tta))
  (mutate (from (g c g)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2aac))
  (mutate (from (g c g)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2cac))
  (mutate (from (g c g)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2gac))
  (mutate (from (g c g)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2tac))
  (mutate (from (g c g)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2acc))
  (mutate (from (g c g)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2ccc))
  (mutate (from (g c g)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2gcc))
  (mutate (from (g c g)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2tcc))
  (mutate (from (g c g)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2agc))
  (mutate (from (g c g)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2cgc))
  (mutate (from (g c g)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2ggc))
  (mutate (from (g c g)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2tgc))
  (mutate (from (g c g)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2atc))
  (mutate (from (g c g)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2ctc))
  (mutate (from (g c g)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2gtc))
  (mutate (from (g c g)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2ttc))
  (mutate (from (g c g)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2aag))
  (mutate (from (g c g)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2cag))
  (mutate (from (g c g)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2gag))
  (mutate (from (g c g)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2tag))
  (mutate (from (g c g)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2acg))
  (mutate (from (g c g)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2ccg))
  (mutate (from (g c g)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2tcg))
  (mutate (from (g c g)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2agg))
  (mutate (from (g c g)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2cgg))
  (mutate (from (g c g)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2ggg))
  (mutate (from (g c g)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2tgg))
  (mutate (from (g c g)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2atg))
  (mutate (from (g c g)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2ctg))
  (mutate (from (g c g)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2gtg))
  (mutate (from (g c g)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2ttg))
  (mutate (from (g c g)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2aat))
  (mutate (from (g c g)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2cat))
  (mutate (from (g c g)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2gat))
  (mutate (from (g c g)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2tat))
  (mutate (from (g c g)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2act))
  (mutate (from (g c g)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2cct))
  (mutate (from (g c g)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2gct))
  (mutate (from (g c g)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2tct))
  (mutate (from (g c g)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2agt))
  (mutate (from (g c g)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2cgt))
  (mutate (from (g c g)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2ggt))
  (mutate (from (g c g)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2tgt))
  (mutate (from (g c g)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2att))
  (mutate (from (g c g)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2ctt))
  (mutate (from (g c g)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2gtt))
  (mutate (from (g c g)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_gcg2ttt))
  (mutate (from (t c g)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2aaa))
  (mutate (from (t c g)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2caa))
  (mutate (from (t c g)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2gaa))
  (mutate (from (t c g)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2taa))
  (mutate (from (t c g)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2aca))
  (mutate (from (t c g)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2cca))
  (mutate (from (t c g)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2gca))
  (mutate (from (t c g)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2tca))
  (mutate (from (t c g)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2aga))
  (mutate (from (t c g)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2cga))
  (mutate (from (t c g)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2gga))
  (mutate (from (t c g)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2tga))
  (mutate (from (t c g)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2ata))
  (mutate (from (t c g)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2cta))
  (mutate (from (t c g)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2gta))
  (mutate (from (t c g)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2tta))
  (mutate (from (t c g)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2aac))
  (mutate (from (t c g)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2cac))
  (mutate (from (t c g)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2gac))
  (mutate (from (t c g)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2tac))
  (mutate (from (t c g)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2acc))
  (mutate (from (t c g)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2ccc))
  (mutate (from (t c g)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2gcc))
  (mutate (from (t c g)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2tcc))
  (mutate (from (t c g)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2agc))
  (mutate (from (t c g)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2cgc))
  (mutate (from (t c g)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2ggc))
  (mutate (from (t c g)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2tgc))
  (mutate (from (t c g)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2atc))
  (mutate (from (t c g)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2ctc))
  (mutate (from (t c g)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2gtc))
  (mutate (from (t c g)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2ttc))
  (mutate (from (t c g)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2aag))
  (mutate (from (t c g)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2cag))
  (mutate (from (t c g)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2gag))
  (mutate (from (t c g)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2tag))
  (mutate (from (t c g)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2acg))
  (mutate (from (t c g)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2ccg))
  (mutate (from (t c g)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2gcg))
  (mutate (from (t c g)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2agg))
  (mutate (from (t c g)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2cgg))
  (mutate (from (t c g)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2ggg))
  (mutate (from (t c g)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2tgg))
  (mutate (from (t c g)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2atg))
  (mutate (from (t c g)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2ctg))
  (mutate (from (t c g)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2gtg))
  (mutate (from (t c g)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2ttg))
  (mutate (from (t c g)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2aat))
  (mutate (from (t c g)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2cat))
  (mutate (from (t c g)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2gat))
  (mutate (from (t c g)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2tat))
  (mutate (from (t c g)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2act))
  (mutate (from (t c g)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2cct))
  (mutate (from (t c g)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2gct))
  (mutate (from (t c g)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2tct))
  (mutate (from (t c g)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2agt))
  (mutate (from (t c g)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2cgt))
  (mutate (from (t c g)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2ggt))
  (mutate (from (t c g)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2tgt))
  (mutate (from (t c g)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2att))
  (mutate (from (t c g)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2ctt))
  (mutate (from (t c g)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2gtt))
  (mutate (from (t c g)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_tcg2ttt))
  (mutate (from (a g g)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_agg2aaa))
  (mutate (from (a g g)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_agg2caa))
  (mutate (from (a g g)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_agg2gaa))
  (mutate (from (a g g)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_agg2taa))
  (mutate (from (a g g)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_agg2aca))
  (mutate (from (a g g)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_agg2cca))
  (mutate (from (a g g)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_agg2gca))
  (mutate (from (a g g)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_agg2tca))
  (mutate (from (a g g)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_agg2aga))
  (mutate (from (a g g)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_agg2cga))
  (mutate (from (a g g)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_agg2gga))
  (mutate (from (a g g)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_agg2tga))
  (mutate (from (a g g)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_agg2ata))
  (mutate (from (a g g)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_agg2cta))
  (mutate (from (a g g)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_agg2gta))
  (mutate (from (a g g)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_agg2tta))
  (mutate (from (a g g)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_agg2aac))
  (mutate (from (a g g)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_agg2cac))
  (mutate (from (a g g)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_agg2gac))
  (mutate (from (a g g)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_agg2tac))
  (mutate (from (a g g)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_agg2acc))
  (mutate (from (a g g)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_agg2ccc))
  (mutate (from (a g g)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_agg2gcc))
  (mutate (from (a g g)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_agg2tcc))
  (mutate (from (a g g)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_agg2agc))
  (mutate (from (a g g)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_agg2cgc))
  (mutate (from (a g g)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_agg2ggc))
  (mutate (from (a g g)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_agg2tgc))
  (mutate (from (a g g)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_agg2atc))
  (mutate (from (a g g)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_agg2ctc))
  (mutate (from (a g g)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_agg2gtc))
  (mutate (from (a g g)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_agg2ttc))
  (mutate (from (a g g)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_agg2aag))
  (mutate (from (a g g)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_agg2cag))
  (mutate (from (a g g)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_agg2gag))
  (mutate (from (a g g)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_agg2tag))
  (mutate (from (a g g)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_agg2acg))
  (mutate (from (a g g)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_agg2ccg))
  (mutate (from (a g g)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_agg2gcg))
  (mutate (from (a g g)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_agg2tcg))
  (mutate (from (a g g)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_agg2cgg))
  (mutate (from (a g g)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_agg2ggg))
  (mutate (from (a g g)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_agg2tgg))
  (mutate (from (a g g)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_agg2atg))
  (mutate (from (a g g)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_agg2ctg))
  (mutate (from (a g g)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_agg2gtg))
  (mutate (from (a g g)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_agg2ttg))
  (mutate (from (a g g)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_agg2aat))
  (mutate (from (a g g)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_agg2cat))
  (mutate (from (a g g)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_agg2gat))
  (mutate (from (a g g)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_agg2tat))
  (mutate (from (a g g)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_agg2act))
  (mutate (from (a g g)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_agg2cct))
  (mutate (from (a g g)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_agg2gct))
  (mutate (from (a g g)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_agg2tct))
  (mutate (from (a g g)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_agg2agt))
  (mutate (from (a g g)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_agg2cgt))
  (mutate (from (a g g)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_agg2ggt))
  (mutate (from (a g g)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_agg2tgt))
  (mutate (from (a g g)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_agg2att))
  (mutate (from (a g g)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_agg2ctt))
  (mutate (from (a g g)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_agg2gtt))
  (mutate (from (a g g)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_agg2ttt))
  (mutate (from (c g g)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2aaa))
  (mutate (from (c g g)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2caa))
  (mutate (from (c g g)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2gaa))
  (mutate (from (c g g)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2taa))
  (mutate (from (c g g)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2aca))
  (mutate (from (c g g)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2cca))
  (mutate (from (c g g)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2gca))
  (mutate (from (c g g)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2tca))
  (mutate (from (c g g)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2aga))
  (mutate (from (c g g)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2cga))
  (mutate (from (c g g)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2gga))
  (mutate (from (c g g)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2tga))
  (mutate (from (c g g)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2ata))
  (mutate (from (c g g)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2cta))
  (mutate (from (c g g)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2gta))
  (mutate (from (c g g)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2tta))
  (mutate (from (c g g)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2aac))
  (mutate (from (c g g)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2cac))
  (mutate (from (c g g)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2gac))
  (mutate (from (c g g)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2tac))
  (mutate (from (c g g)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2acc))
  (mutate (from (c g g)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2ccc))
  (mutate (from (c g g)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2gcc))
  (mutate (from (c g g)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2tcc))
  (mutate (from (c g g)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2agc))
  (mutate (from (c g g)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2cgc))
  (mutate (from (c g g)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2ggc))
  (mutate (from (c g g)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2tgc))
  (mutate (from (c g g)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2atc))
  (mutate (from (c g g)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2ctc))
  (mutate (from (c g g)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2gtc))
  (mutate (from (c g g)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2ttc))
  (mutate (from (c g g)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2aag))
  (mutate (from (c g g)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2cag))
  (mutate (from (c g g)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2gag))
  (mutate (from (c g g)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2tag))
  (mutate (from (c g g)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2acg))
  (mutate (from (c g g)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2ccg))
  (mutate (from (c g g)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2gcg))
  (mutate (from (c g g)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2tcg))
  (mutate (from (c g g)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2agg))
  (mutate (from (c g g)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2ggg))
  (mutate (from (c g g)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2tgg))
  (mutate (from (c g g)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2atg))
  (mutate (from (c g g)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2ctg))
  (mutate (from (c g g)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2gtg))
  (mutate (from (c g g)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2ttg))
  (mutate (from (c g g)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2aat))
  (mutate (from (c g g)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2cat))
  (mutate (from (c g g)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2gat))
  (mutate (from (c g g)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2tat))
  (mutate (from (c g g)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2act))
  (mutate (from (c g g)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2cct))
  (mutate (from (c g g)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2gct))
  (mutate (from (c g g)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2tct))
  (mutate (from (c g g)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2agt))
  (mutate (from (c g g)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2cgt))
  (mutate (from (c g g)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2ggt))
  (mutate (from (c g g)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2tgt))
  (mutate (from (c g g)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2att))
  (mutate (from (c g g)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2ctt))
  (mutate (from (c g g)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2gtt))
  (mutate (from (c g g)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_cgg2ttt))
  (mutate (from (g g g)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2aaa))
  (mutate (from (g g g)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2caa))
  (mutate (from (g g g)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2gaa))
  (mutate (from (g g g)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2taa))
  (mutate (from (g g g)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2aca))
  (mutate (from (g g g)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2cca))
  (mutate (from (g g g)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2gca))
  (mutate (from (g g g)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2tca))
  (mutate (from (g g g)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2aga))
  (mutate (from (g g g)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2cga))
  (mutate (from (g g g)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2gga))
  (mutate (from (g g g)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2tga))
  (mutate (from (g g g)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2ata))
  (mutate (from (g g g)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2cta))
  (mutate (from (g g g)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2gta))
  (mutate (from (g g g)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2tta))
  (mutate (from (g g g)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2aac))
  (mutate (from (g g g)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2cac))
  (mutate (from (g g g)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2gac))
  (mutate (from (g g g)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2tac))
  (mutate (from (g g g)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2acc))
  (mutate (from (g g g)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2ccc))
  (mutate (from (g g g)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2gcc))
  (mutate (from (g g g)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2tcc))
  (mutate (from (g g g)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2agc))
  (mutate (from (g g g)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2cgc))
  (mutate (from (g g g)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2ggc))
  (mutate (from (g g g)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2tgc))
  (mutate (from (g g g)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2atc))
  (mutate (from (g g g)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2ctc))
  (mutate (from (g g g)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2gtc))
  (mutate (from (g g g)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2ttc))
  (mutate (from (g g g)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2aag))
  (mutate (from (g g g)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2cag))
  (mutate (from (g g g)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2gag))
  (mutate (from (g g g)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2tag))
  (mutate (from (g g g)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2acg))
  (mutate (from (g g g)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2ccg))
  (mutate (from (g g g)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2gcg))
  (mutate (from (g g g)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2tcg))
  (mutate (from (g g g)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2agg))
  (mutate (from (g g g)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2cgg))
  (mutate (from (g g g)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2tgg))
  (mutate (from (g g g)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2atg))
  (mutate (from (g g g)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2ctg))
  (mutate (from (g g g)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2gtg))
  (mutate (from (g g g)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2ttg))
  (mutate (from (g g g)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2aat))
  (mutate (from (g g g)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2cat))
  (mutate (from (g g g)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2gat))
  (mutate (from (g g g)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2tat))
  (mutate (from (g g g)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2act))
  (mutate (from (g g g)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2cct))
  (mutate (from (g g g)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2gct))
  (mutate (from (g g g)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2tct))
  (mutate (from (g g g)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2agt))
  (mutate (from (g g g)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2cgt))
  (mutate (from (g g g)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2ggt))
  (mutate (from (g g g)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2tgt))
  (mutate (from (g g g)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2att))
  (mutate (from (g g g)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2ctt))
  (mutate (from (g g g)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2gtt))
  (mutate (from (g g g)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_ggg2ttt))
  (mutate (from (t g g)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2aaa))
  (mutate (from (t g g)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2caa))
  (mutate (from (t g g)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2gaa))
  (mutate (from (t g g)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2taa))
  (mutate (from (t g g)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2aca))
  (mutate (from (t g g)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2cca))
  (mutate (from (t g g)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2gca))
  (mutate (from (t g g)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2tca))
  (mutate (from (t g g)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2aga))
  (mutate (from (t g g)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2cga))
  (mutate (from (t g g)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2gga))
  (mutate (from (t g g)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2tga))
  (mutate (from (t g g)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2ata))
  (mutate (from (t g g)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2cta))
  (mutate (from (t g g)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2gta))
  (mutate (from (t g g)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2tta))
  (mutate (from (t g g)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2aac))
  (mutate (from (t g g)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2cac))
  (mutate (from (t g g)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2gac))
  (mutate (from (t g g)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2tac))
  (mutate (from (t g g)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2acc))
  (mutate (from (t g g)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2ccc))
  (mutate (from (t g g)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2gcc))
  (mutate (from (t g g)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2tcc))
  (mutate (from (t g g)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2agc))
  (mutate (from (t g g)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2cgc))
  (mutate (from (t g g)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2ggc))
  (mutate (from (t g g)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2tgc))
  (mutate (from (t g g)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2atc))
  (mutate (from (t g g)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2ctc))
  (mutate (from (t g g)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2gtc))
  (mutate (from (t g g)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2ttc))
  (mutate (from (t g g)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2aag))
  (mutate (from (t g g)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2cag))
  (mutate (from (t g g)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2gag))
  (mutate (from (t g g)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2tag))
  (mutate (from (t g g)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2acg))
  (mutate (from (t g g)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2ccg))
  (mutate (from (t g g)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2gcg))
  (mutate (from (t g g)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2tcg))
  (mutate (from (t g g)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2agg))
  (mutate (from (t g g)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2cgg))
  (mutate (from (t g g)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2ggg))
  (mutate (from (t g g)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2atg))
  (mutate (from (t g g)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2ctg))
  (mutate (from (t g g)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2gtg))
  (mutate (from (t g g)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2ttg))
  (mutate (from (t g g)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2aat))
  (mutate (from (t g g)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2cat))
  (mutate (from (t g g)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2gat))
  (mutate (from (t g g)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2tat))
  (mutate (from (t g g)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2act))
  (mutate (from (t g g)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2cct))
  (mutate (from (t g g)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2gct))
  (mutate (from (t g g)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2tct))
  (mutate (from (t g g)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2agt))
  (mutate (from (t g g)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2cgt))
  (mutate (from (t g g)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2ggt))
  (mutate (from (t g g)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2tgt))
  (mutate (from (t g g)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2att))
  (mutate (from (t g g)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2ctt))
  (mutate (from (t g g)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2gtt))
  (mutate (from (t g g)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_tgg2ttt))
  (mutate (from (a t g)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_atg2aaa))
  (mutate (from (a t g)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_atg2caa))
  (mutate (from (a t g)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_atg2gaa))
  (mutate (from (a t g)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_atg2taa))
  (mutate (from (a t g)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_atg2aca))
  (mutate (from (a t g)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_atg2cca))
  (mutate (from (a t g)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_atg2gca))
  (mutate (from (a t g)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_atg2tca))
  (mutate (from (a t g)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_atg2aga))
  (mutate (from (a t g)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_atg2cga))
  (mutate (from (a t g)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_atg2gga))
  (mutate (from (a t g)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_atg2tga))
  (mutate (from (a t g)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_atg2ata))
  (mutate (from (a t g)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_atg2cta))
  (mutate (from (a t g)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_atg2gta))
  (mutate (from (a t g)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_atg2tta))
  (mutate (from (a t g)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_atg2aac))
  (mutate (from (a t g)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_atg2cac))
  (mutate (from (a t g)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_atg2gac))
  (mutate (from (a t g)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_atg2tac))
  (mutate (from (a t g)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_atg2acc))
  (mutate (from (a t g)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_atg2ccc))
  (mutate (from (a t g)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_atg2gcc))
  (mutate (from (a t g)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_atg2tcc))
  (mutate (from (a t g)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_atg2agc))
  (mutate (from (a t g)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_atg2cgc))
  (mutate (from (a t g)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_atg2ggc))
  (mutate (from (a t g)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_atg2tgc))
  (mutate (from (a t g)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_atg2atc))
  (mutate (from (a t g)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_atg2ctc))
  (mutate (from (a t g)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_atg2gtc))
  (mutate (from (a t g)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_atg2ttc))
  (mutate (from (a t g)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_atg2aag))
  (mutate (from (a t g)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_atg2cag))
  (mutate (from (a t g)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_atg2gag))
  (mutate (from (a t g)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_atg2tag))
  (mutate (from (a t g)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_atg2acg))
  (mutate (from (a t g)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_atg2ccg))
  (mutate (from (a t g)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_atg2gcg))
  (mutate (from (a t g)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_atg2tcg))
  (mutate (from (a t g)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_atg2agg))
  (mutate (from (a t g)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_atg2cgg))
  (mutate (from (a t g)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_atg2ggg))
  (mutate (from (a t g)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_atg2tgg))
  (mutate (from (a t g)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_atg2ctg))
  (mutate (from (a t g)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_atg2gtg))
  (mutate (from (a t g)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_atg2ttg))
  (mutate (from (a t g)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_atg2aat))
  (mutate (from (a t g)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_atg2cat))
  (mutate (from (a t g)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_atg2gat))
  (mutate (from (a t g)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_atg2tat))
  (mutate (from (a t g)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_atg2act))
  (mutate (from (a t g)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_atg2cct))
  (mutate (from (a t g)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_atg2gct))
  (mutate (from (a t g)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_atg2tct))
  (mutate (from (a t g)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_atg2agt))
  (mutate (from (a t g)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_atg2cgt))
  (mutate (from (a t g)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_atg2ggt))
  (mutate (from (a t g)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_atg2tgt))
  (mutate (from (a t g)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_atg2att))
  (mutate (from (a t g)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_atg2ctt))
  (mutate (from (a t g)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_atg2gtt))
  (mutate (from (a t g)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_atg2ttt))
  (mutate (from (c t g)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2aaa))
  (mutate (from (c t g)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2caa))
  (mutate (from (c t g)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2gaa))
  (mutate (from (c t g)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2taa))
  (mutate (from (c t g)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2aca))
  (mutate (from (c t g)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2cca))
  (mutate (from (c t g)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2gca))
  (mutate (from (c t g)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2tca))
  (mutate (from (c t g)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2aga))
  (mutate (from (c t g)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2cga))
  (mutate (from (c t g)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2gga))
  (mutate (from (c t g)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2tga))
  (mutate (from (c t g)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2ata))
  (mutate (from (c t g)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2cta))
  (mutate (from (c t g)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2gta))
  (mutate (from (c t g)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2tta))
  (mutate (from (c t g)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2aac))
  (mutate (from (c t g)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2cac))
  (mutate (from (c t g)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2gac))
  (mutate (from (c t g)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2tac))
  (mutate (from (c t g)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2acc))
  (mutate (from (c t g)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2ccc))
  (mutate (from (c t g)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2gcc))
  (mutate (from (c t g)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2tcc))
  (mutate (from (c t g)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2agc))
  (mutate (from (c t g)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2cgc))
  (mutate (from (c t g)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2ggc))
  (mutate (from (c t g)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2tgc))
  (mutate (from (c t g)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2atc))
  (mutate (from (c t g)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2ctc))
  (mutate (from (c t g)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2gtc))
  (mutate (from (c t g)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2ttc))
  (mutate (from (c t g)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2aag))
  (mutate (from (c t g)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2cag))
  (mutate (from (c t g)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2gag))
  (mutate (from (c t g)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2tag))
  (mutate (from (c t g)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2acg))
  (mutate (from (c t g)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2ccg))
  (mutate (from (c t g)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2gcg))
  (mutate (from (c t g)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2tcg))
  (mutate (from (c t g)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2agg))
  (mutate (from (c t g)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2cgg))
  (mutate (from (c t g)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2ggg))
  (mutate (from (c t g)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2tgg))
  (mutate (from (c t g)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2atg))
  (mutate (from (c t g)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2gtg))
  (mutate (from (c t g)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2ttg))
  (mutate (from (c t g)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2aat))
  (mutate (from (c t g)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2cat))
  (mutate (from (c t g)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2gat))
  (mutate (from (c t g)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2tat))
  (mutate (from (c t g)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2act))
  (mutate (from (c t g)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2cct))
  (mutate (from (c t g)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2gct))
  (mutate (from (c t g)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2tct))
  (mutate (from (c t g)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2agt))
  (mutate (from (c t g)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2cgt))
  (mutate (from (c t g)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2ggt))
  (mutate (from (c t g)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2tgt))
  (mutate (from (c t g)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2att))
  (mutate (from (c t g)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2ctt))
  (mutate (from (c t g)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2gtt))
  (mutate (from (c t g)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_ctg2ttt))
  (mutate (from (g t g)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2aaa))
  (mutate (from (g t g)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2caa))
  (mutate (from (g t g)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2gaa))
  (mutate (from (g t g)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2taa))
  (mutate (from (g t g)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2aca))
  (mutate (from (g t g)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2cca))
  (mutate (from (g t g)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2gca))
  (mutate (from (g t g)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2tca))
  (mutate (from (g t g)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2aga))
  (mutate (from (g t g)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2cga))
  (mutate (from (g t g)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2gga))
  (mutate (from (g t g)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2tga))
  (mutate (from (g t g)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2ata))
  (mutate (from (g t g)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2cta))
  (mutate (from (g t g)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2gta))
  (mutate (from (g t g)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2tta))
  (mutate (from (g t g)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2aac))
  (mutate (from (g t g)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2cac))
  (mutate (from (g t g)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2gac))
  (mutate (from (g t g)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2tac))
  (mutate (from (g t g)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2acc))
  (mutate (from (g t g)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2ccc))
  (mutate (from (g t g)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2gcc))
  (mutate (from (g t g)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2tcc))
  (mutate (from (g t g)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2agc))
  (mutate (from (g t g)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2cgc))
  (mutate (from (g t g)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2ggc))
  (mutate (from (g t g)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2tgc))
  (mutate (from (g t g)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2atc))
  (mutate (from (g t g)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2ctc))
  (mutate (from (g t g)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2gtc))
  (mutate (from (g t g)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2ttc))
  (mutate (from (g t g)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2aag))
  (mutate (from (g t g)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2cag))
  (mutate (from (g t g)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2gag))
  (mutate (from (g t g)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2tag))
  (mutate (from (g t g)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2acg))
  (mutate (from (g t g)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2ccg))
  (mutate (from (g t g)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2gcg))
  (mutate (from (g t g)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2tcg))
  (mutate (from (g t g)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2agg))
  (mutate (from (g t g)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2cgg))
  (mutate (from (g t g)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2ggg))
  (mutate (from (g t g)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2tgg))
  (mutate (from (g t g)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2atg))
  (mutate (from (g t g)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2ctg))
  (mutate (from (g t g)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2ttg))
  (mutate (from (g t g)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2aat))
  (mutate (from (g t g)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2cat))
  (mutate (from (g t g)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2gat))
  (mutate (from (g t g)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2tat))
  (mutate (from (g t g)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2act))
  (mutate (from (g t g)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2cct))
  (mutate (from (g t g)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2gct))
  (mutate (from (g t g)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2tct))
  (mutate (from (g t g)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2agt))
  (mutate (from (g t g)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2cgt))
  (mutate (from (g t g)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2ggt))
  (mutate (from (g t g)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2tgt))
  (mutate (from (g t g)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2att))
  (mutate (from (g t g)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2ctt))
  (mutate (from (g t g)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2gtt))
  (mutate (from (g t g)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_gtg2ttt))
  (mutate (from (t t g)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2aaa))
  (mutate (from (t t g)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2caa))
  (mutate (from (t t g)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2gaa))
  (mutate (from (t t g)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2taa))
  (mutate (from (t t g)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2aca))
  (mutate (from (t t g)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2cca))
  (mutate (from (t t g)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2gca))
  (mutate (from (t t g)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2tca))
  (mutate (from (t t g)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2aga))
  (mutate (from (t t g)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2cga))
  (mutate (from (t t g)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2gga))
  (mutate (from (t t g)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2tga))
  (mutate (from (t t g)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2ata))
  (mutate (from (t t g)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2cta))
  (mutate (from (t t g)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2gta))
  (mutate (from (t t g)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2tta))
  (mutate (from (t t g)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2aac))
  (mutate (from (t t g)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2cac))
  (mutate (from (t t g)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2gac))
  (mutate (from (t t g)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2tac))
  (mutate (from (t t g)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2acc))
  (mutate (from (t t g)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2ccc))
  (mutate (from (t t g)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2gcc))
  (mutate (from (t t g)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2tcc))
  (mutate (from (t t g)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2agc))
  (mutate (from (t t g)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2cgc))
  (mutate (from (t t g)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2ggc))
  (mutate (from (t t g)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2tgc))
  (mutate (from (t t g)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2atc))
  (mutate (from (t t g)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2ctc))
  (mutate (from (t t g)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2gtc))
  (mutate (from (t t g)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2ttc))
  (mutate (from (t t g)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2aag))
  (mutate (from (t t g)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2cag))
  (mutate (from (t t g)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2gag))
  (mutate (from (t t g)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2tag))
  (mutate (from (t t g)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2acg))
  (mutate (from (t t g)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2ccg))
  (mutate (from (t t g)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2gcg))
  (mutate (from (t t g)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2tcg))
  (mutate (from (t t g)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2agg))
  (mutate (from (t t g)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2cgg))
  (mutate (from (t t g)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2ggg))
  (mutate (from (t t g)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2tgg))
  (mutate (from (t t g)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2atg))
  (mutate (from (t t g)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2ctg))
  (mutate (from (t t g)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2gtg))
  (mutate (from (t t g)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2aat))
  (mutate (from (t t g)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2cat))
  (mutate (from (t t g)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2gat))
  (mutate (from (t t g)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2tat))
  (mutate (from (t t g)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2act))
  (mutate (from (t t g)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2cct))
  (mutate (from (t t g)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2gct))
  (mutate (from (t t g)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2tct))
  (mutate (from (t t g)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2agt))
  (mutate (from (t t g)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2cgt))
  (mutate (from (t t g)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2ggt))
  (mutate (from (t t g)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2tgt))
  (mutate (from (t t g)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2att))
  (mutate (from (t t g)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2ctt))
  (mutate (from (t t g)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2gtt))
  (mutate (from (t t g)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_ttg2ttt))
  (mutate (from (a a t)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_aat2aaa))
  (mutate (from (a a t)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_aat2caa))
  (mutate (from (a a t)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_aat2gaa))
  (mutate (from (a a t)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_aat2taa))
  (mutate (from (a a t)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_aat2aca))
  (mutate (from (a a t)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_aat2cca))
  (mutate (from (a a t)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_aat2gca))
  (mutate (from (a a t)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_aat2tca))
  (mutate (from (a a t)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_aat2aga))
  (mutate (from (a a t)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_aat2cga))
  (mutate (from (a a t)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_aat2gga))
  (mutate (from (a a t)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_aat2tga))
  (mutate (from (a a t)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_aat2ata))
  (mutate (from (a a t)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_aat2cta))
  (mutate (from (a a t)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_aat2gta))
  (mutate (from (a a t)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_aat2tta))
  (mutate (from (a a t)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_aat2aac))
  (mutate (from (a a t)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_aat2cac))
  (mutate (from (a a t)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_aat2gac))
  (mutate (from (a a t)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_aat2tac))
  (mutate (from (a a t)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_aat2acc))
  (mutate (from (a a t)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_aat2ccc))
  (mutate (from (a a t)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_aat2gcc))
  (mutate (from (a a t)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_aat2tcc))
  (mutate (from (a a t)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_aat2agc))
  (mutate (from (a a t)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_aat2cgc))
  (mutate (from (a a t)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_aat2ggc))
  (mutate (from (a a t)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_aat2tgc))
  (mutate (from (a a t)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_aat2atc))
  (mutate (from (a a t)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_aat2ctc))
  (mutate (from (a a t)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_aat2gtc))
  (mutate (from (a a t)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_aat2ttc))
  (mutate (from (a a t)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_aat2aag))
  (mutate (from (a a t)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_aat2cag))
  (mutate (from (a a t)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_aat2gag))
  (mutate (from (a a t)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_aat2tag))
  (mutate (from (a a t)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_aat2acg))
  (mutate (from (a a t)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_aat2ccg))
  (mutate (from (a a t)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_aat2gcg))
  (mutate (from (a a t)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_aat2tcg))
  (mutate (from (a a t)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_aat2agg))
  (mutate (from (a a t)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_aat2cgg))
  (mutate (from (a a t)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_aat2ggg))
  (mutate (from (a a t)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_aat2tgg))
  (mutate (from (a a t)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_aat2atg))
  (mutate (from (a a t)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_aat2ctg))
  (mutate (from (a a t)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_aat2gtg))
  (mutate (from (a a t)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_aat2ttg))
  (mutate (from (a a t)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_aat2cat))
  (mutate (from (a a t)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_aat2gat))
  (mutate (from (a a t)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_aat2tat))
  (mutate (from (a a t)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_aat2act))
  (mutate (from (a a t)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_aat2cct))
  (mutate (from (a a t)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_aat2gct))
  (mutate (from (a a t)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_aat2tct))
  (mutate (from (a a t)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_aat2agt))
  (mutate (from (a a t)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_aat2cgt))
  (mutate (from (a a t)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_aat2ggt))
  (mutate (from (a a t)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_aat2tgt))
  (mutate (from (a a t)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_aat2att))
  (mutate (from (a a t)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_aat2ctt))
  (mutate (from (a a t)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_aat2gtt))
  (mutate (from (a a t)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_aat2ttt))
  (mutate (from (c a t)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_cat2aaa))
  (mutate (from (c a t)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_cat2caa))
  (mutate (from (c a t)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_cat2gaa))
  (mutate (from (c a t)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_cat2taa))
  (mutate (from (c a t)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_cat2aca))
  (mutate (from (c a t)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_cat2cca))
  (mutate (from (c a t)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_cat2gca))
  (mutate (from (c a t)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_cat2tca))
  (mutate (from (c a t)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_cat2aga))
  (mutate (from (c a t)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_cat2cga))
  (mutate (from (c a t)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_cat2gga))
  (mutate (from (c a t)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_cat2tga))
  (mutate (from (c a t)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_cat2ata))
  (mutate (from (c a t)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_cat2cta))
  (mutate (from (c a t)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_cat2gta))
  (mutate (from (c a t)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_cat2tta))
  (mutate (from (c a t)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_cat2aac))
  (mutate (from (c a t)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_cat2cac))
  (mutate (from (c a t)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_cat2gac))
  (mutate (from (c a t)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_cat2tac))
  (mutate (from (c a t)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_cat2acc))
  (mutate (from (c a t)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_cat2ccc))
  (mutate (from (c a t)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_cat2gcc))
  (mutate (from (c a t)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_cat2tcc))
  (mutate (from (c a t)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_cat2agc))
  (mutate (from (c a t)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_cat2cgc))
  (mutate (from (c a t)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_cat2ggc))
  (mutate (from (c a t)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_cat2tgc))
  (mutate (from (c a t)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_cat2atc))
  (mutate (from (c a t)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_cat2ctc))
  (mutate (from (c a t)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_cat2gtc))
  (mutate (from (c a t)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_cat2ttc))
  (mutate (from (c a t)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_cat2aag))
  (mutate (from (c a t)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_cat2cag))
  (mutate (from (c a t)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_cat2gag))
  (mutate (from (c a t)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_cat2tag))
  (mutate (from (c a t)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_cat2acg))
  (mutate (from (c a t)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_cat2ccg))
  (mutate (from (c a t)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_cat2gcg))
  (mutate (from (c a t)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_cat2tcg))
  (mutate (from (c a t)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_cat2agg))
  (mutate (from (c a t)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_cat2cgg))
  (mutate (from (c a t)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_cat2ggg))
  (mutate (from (c a t)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_cat2tgg))
  (mutate (from (c a t)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_cat2atg))
  (mutate (from (c a t)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_cat2ctg))
  (mutate (from (c a t)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_cat2gtg))
  (mutate (from (c a t)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_cat2ttg))
  (mutate (from (c a t)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_cat2aat))
  (mutate (from (c a t)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_cat2gat))
  (mutate (from (c a t)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_cat2tat))
  (mutate (from (c a t)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_cat2act))
  (mutate (from (c a t)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_cat2cct))
  (mutate (from (c a t)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_cat2gct))
  (mutate (from (c a t)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_cat2tct))
  (mutate (from (c a t)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_cat2agt))
  (mutate (from (c a t)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_cat2cgt))
  (mutate (from (c a t)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_cat2ggt))
  (mutate (from (c a t)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_cat2tgt))
  (mutate (from (c a t)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_cat2att))
  (mutate (from (c a t)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_cat2ctt))
  (mutate (from (c a t)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_cat2gtt))
  (mutate (from (c a t)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_cat2ttt))
  (mutate (from (g a t)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_gat2aaa))
  (mutate (from (g a t)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_gat2caa))
  (mutate (from (g a t)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_gat2gaa))
  (mutate (from (g a t)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_gat2taa))
  (mutate (from (g a t)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_gat2aca))
  (mutate (from (g a t)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_gat2cca))
  (mutate (from (g a t)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_gat2gca))
  (mutate (from (g a t)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_gat2tca))
  (mutate (from (g a t)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_gat2aga))
  (mutate (from (g a t)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_gat2cga))
  (mutate (from (g a t)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_gat2gga))
  (mutate (from (g a t)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_gat2tga))
  (mutate (from (g a t)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_gat2ata))
  (mutate (from (g a t)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_gat2cta))
  (mutate (from (g a t)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_gat2gta))
  (mutate (from (g a t)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_gat2tta))
  (mutate (from (g a t)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_gat2aac))
  (mutate (from (g a t)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_gat2cac))
  (mutate (from (g a t)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_gat2gac))
  (mutate (from (g a t)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_gat2tac))
  (mutate (from (g a t)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_gat2acc))
  (mutate (from (g a t)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_gat2ccc))
  (mutate (from (g a t)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_gat2gcc))
  (mutate (from (g a t)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_gat2tcc))
  (mutate (from (g a t)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_gat2agc))
  (mutate (from (g a t)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_gat2cgc))
  (mutate (from (g a t)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_gat2ggc))
  (mutate (from (g a t)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_gat2tgc))
  (mutate (from (g a t)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_gat2atc))
  (mutate (from (g a t)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_gat2ctc))
  (mutate (from (g a t)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_gat2gtc))
  (mutate (from (g a t)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_gat2ttc))
  (mutate (from (g a t)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_gat2aag))
  (mutate (from (g a t)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_gat2cag))
  (mutate (from (g a t)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_gat2gag))
  (mutate (from (g a t)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_gat2tag))
  (mutate (from (g a t)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_gat2acg))
  (mutate (from (g a t)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_gat2ccg))
  (mutate (from (g a t)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_gat2gcg))
  (mutate (from (g a t)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_gat2tcg))
  (mutate (from (g a t)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_gat2agg))
  (mutate (from (g a t)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_gat2cgg))
  (mutate (from (g a t)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_gat2ggg))
  (mutate (from (g a t)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_gat2tgg))
  (mutate (from (g a t)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_gat2atg))
  (mutate (from (g a t)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_gat2ctg))
  (mutate (from (g a t)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_gat2gtg))
  (mutate (from (g a t)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_gat2ttg))
  (mutate (from (g a t)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_gat2aat))
  (mutate (from (g a t)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_gat2cat))
  (mutate (from (g a t)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_gat2tat))
  (mutate (from (g a t)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_gat2act))
  (mutate (from (g a t)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_gat2cct))
  (mutate (from (g a t)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_gat2gct))
  (mutate (from (g a t)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_gat2tct))
  (mutate (from (g a t)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_gat2agt))
  (mutate (from (g a t)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_gat2cgt))
  (mutate (from (g a t)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_gat2ggt))
  (mutate (from (g a t)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_gat2tgt))
  (mutate (from (g a t)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_gat2att))
  (mutate (from (g a t)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_gat2ctt))
  (mutate (from (g a t)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_gat2gtt))
  (mutate (from (g a t)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_gat2ttt))
  (mutate (from (t a t)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_tat2aaa))
  (mutate (from (t a t)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_tat2caa))
  (mutate (from (t a t)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_tat2gaa))
  (mutate (from (t a t)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_tat2taa))
  (mutate (from (t a t)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_tat2aca))
  (mutate (from (t a t)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_tat2cca))
  (mutate (from (t a t)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_tat2gca))
  (mutate (from (t a t)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_tat2tca))
  (mutate (from (t a t)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_tat2aga))
  (mutate (from (t a t)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_tat2cga))
  (mutate (from (t a t)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_tat2gga))
  (mutate (from (t a t)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_tat2tga))
  (mutate (from (t a t)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_tat2ata))
  (mutate (from (t a t)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_tat2cta))
  (mutate (from (t a t)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_tat2gta))
  (mutate (from (t a t)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_tat2tta))
  (mutate (from (t a t)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_tat2aac))
  (mutate (from (t a t)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_tat2cac))
  (mutate (from (t a t)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_tat2gac))
  (mutate (from (t a t)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_tat2tac))
  (mutate (from (t a t)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_tat2acc))
  (mutate (from (t a t)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_tat2ccc))
  (mutate (from (t a t)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_tat2gcc))
  (mutate (from (t a t)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_tat2tcc))
  (mutate (from (t a t)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_tat2agc))
  (mutate (from (t a t)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_tat2cgc))
  (mutate (from (t a t)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_tat2ggc))
  (mutate (from (t a t)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_tat2tgc))
  (mutate (from (t a t)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_tat2atc))
  (mutate (from (t a t)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_tat2ctc))
  (mutate (from (t a t)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_tat2gtc))
  (mutate (from (t a t)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_tat2ttc))
  (mutate (from (t a t)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_tat2aag))
  (mutate (from (t a t)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_tat2cag))
  (mutate (from (t a t)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_tat2gag))
  (mutate (from (t a t)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_tat2tag))
  (mutate (from (t a t)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_tat2acg))
  (mutate (from (t a t)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_tat2ccg))
  (mutate (from (t a t)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_tat2gcg))
  (mutate (from (t a t)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_tat2tcg))
  (mutate (from (t a t)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_tat2agg))
  (mutate (from (t a t)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_tat2cgg))
  (mutate (from (t a t)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_tat2ggg))
  (mutate (from (t a t)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_tat2tgg))
  (mutate (from (t a t)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_tat2atg))
  (mutate (from (t a t)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_tat2ctg))
  (mutate (from (t a t)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_tat2gtg))
  (mutate (from (t a t)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_tat2ttg))
  (mutate (from (t a t)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_tat2aat))
  (mutate (from (t a t)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_tat2cat))
  (mutate (from (t a t)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_tat2gat))
  (mutate (from (t a t)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_tat2act))
  (mutate (from (t a t)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_tat2cct))
  (mutate (from (t a t)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_tat2gct))
  (mutate (from (t a t)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_tat2tct))
  (mutate (from (t a t)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_tat2agt))
  (mutate (from (t a t)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_tat2cgt))
  (mutate (from (t a t)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_tat2ggt))
  (mutate (from (t a t)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_tat2tgt))
  (mutate (from (t a t)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_tat2att))
  (mutate (from (t a t)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_tat2ctt))
  (mutate (from (t a t)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_tat2gtt))
  (mutate (from (t a t)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_tat2ttt))
  (mutate (from (a c t)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_act2aaa))
  (mutate (from (a c t)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_act2caa))
  (mutate (from (a c t)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_act2gaa))
  (mutate (from (a c t)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_act2taa))
  (mutate (from (a c t)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_act2aca))
  (mutate (from (a c t)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_act2cca))
  (mutate (from (a c t)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_act2gca))
  (mutate (from (a c t)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_act2tca))
  (mutate (from (a c t)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_act2aga))
  (mutate (from (a c t)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_act2cga))
  (mutate (from (a c t)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_act2gga))
  (mutate (from (a c t)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_act2tga))
  (mutate (from (a c t)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_act2ata))
  (mutate (from (a c t)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_act2cta))
  (mutate (from (a c t)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_act2gta))
  (mutate (from (a c t)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_act2tta))
  (mutate (from (a c t)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_act2aac))
  (mutate (from (a c t)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_act2cac))
  (mutate (from (a c t)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_act2gac))
  (mutate (from (a c t)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_act2tac))
  (mutate (from (a c t)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_act2acc))
  (mutate (from (a c t)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_act2ccc))
  (mutate (from (a c t)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_act2gcc))
  (mutate (from (a c t)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_act2tcc))
  (mutate (from (a c t)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_act2agc))
  (mutate (from (a c t)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_act2cgc))
  (mutate (from (a c t)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_act2ggc))
  (mutate (from (a c t)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_act2tgc))
  (mutate (from (a c t)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_act2atc))
  (mutate (from (a c t)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_act2ctc))
  (mutate (from (a c t)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_act2gtc))
  (mutate (from (a c t)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_act2ttc))
  (mutate (from (a c t)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_act2aag))
  (mutate (from (a c t)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_act2cag))
  (mutate (from (a c t)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_act2gag))
  (mutate (from (a c t)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_act2tag))
  (mutate (from (a c t)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_act2acg))
  (mutate (from (a c t)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_act2ccg))
  (mutate (from (a c t)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_act2gcg))
  (mutate (from (a c t)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_act2tcg))
  (mutate (from (a c t)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_act2agg))
  (mutate (from (a c t)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_act2cgg))
  (mutate (from (a c t)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_act2ggg))
  (mutate (from (a c t)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_act2tgg))
  (mutate (from (a c t)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_act2atg))
  (mutate (from (a c t)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_act2ctg))
  (mutate (from (a c t)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_act2gtg))
  (mutate (from (a c t)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_act2ttg))
  (mutate (from (a c t)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_act2aat))
  (mutate (from (a c t)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_act2cat))
  (mutate (from (a c t)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_act2gat))
  (mutate (from (a c t)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_act2tat))
  (mutate (from (a c t)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_act2cct))
  (mutate (from (a c t)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_act2gct))
  (mutate (from (a c t)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_act2tct))
  (mutate (from (a c t)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_act2agt))
  (mutate (from (a c t)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_act2cgt))
  (mutate (from (a c t)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_act2ggt))
  (mutate (from (a c t)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_act2tgt))
  (mutate (from (a c t)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_act2att))
  (mutate (from (a c t)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_act2ctt))
  (mutate (from (a c t)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_act2gtt))
  (mutate (from (a c t)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_act2ttt))
  (mutate (from (c c t)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_cct2aaa))
  (mutate (from (c c t)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_cct2caa))
  (mutate (from (c c t)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_cct2gaa))
  (mutate (from (c c t)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_cct2taa))
  (mutate (from (c c t)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_cct2aca))
  (mutate (from (c c t)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_cct2cca))
  (mutate (from (c c t)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_cct2gca))
  (mutate (from (c c t)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_cct2tca))
  (mutate (from (c c t)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_cct2aga))
  (mutate (from (c c t)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_cct2cga))
  (mutate (from (c c t)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_cct2gga))
  (mutate (from (c c t)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_cct2tga))
  (mutate (from (c c t)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_cct2ata))
  (mutate (from (c c t)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_cct2cta))
  (mutate (from (c c t)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_cct2gta))
  (mutate (from (c c t)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_cct2tta))
  (mutate (from (c c t)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_cct2aac))
  (mutate (from (c c t)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_cct2cac))
  (mutate (from (c c t)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_cct2gac))
  (mutate (from (c c t)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_cct2tac))
  (mutate (from (c c t)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_cct2acc))
  (mutate (from (c c t)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_cct2ccc))
  (mutate (from (c c t)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_cct2gcc))
  (mutate (from (c c t)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_cct2tcc))
  (mutate (from (c c t)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_cct2agc))
  (mutate (from (c c t)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_cct2cgc))
  (mutate (from (c c t)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_cct2ggc))
  (mutate (from (c c t)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_cct2tgc))
  (mutate (from (c c t)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_cct2atc))
  (mutate (from (c c t)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_cct2ctc))
  (mutate (from (c c t)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_cct2gtc))
  (mutate (from (c c t)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_cct2ttc))
  (mutate (from (c c t)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_cct2aag))
  (mutate (from (c c t)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_cct2cag))
  (mutate (from (c c t)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_cct2gag))
  (mutate (from (c c t)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_cct2tag))
  (mutate (from (c c t)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_cct2acg))
  (mutate (from (c c t)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_cct2ccg))
  (mutate (from (c c t)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_cct2gcg))
  (mutate (from (c c t)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_cct2tcg))
  (mutate (from (c c t)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_cct2agg))
  (mutate (from (c c t)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_cct2cgg))
  (mutate (from (c c t)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_cct2ggg))
  (mutate (from (c c t)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_cct2tgg))
  (mutate (from (c c t)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_cct2atg))
  (mutate (from (c c t)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_cct2ctg))
  (mutate (from (c c t)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_cct2gtg))
  (mutate (from (c c t)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_cct2ttg))
  (mutate (from (c c t)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_cct2aat))
  (mutate (from (c c t)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_cct2cat))
  (mutate (from (c c t)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_cct2gat))
  (mutate (from (c c t)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_cct2tat))
  (mutate (from (c c t)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_cct2act))
  (mutate (from (c c t)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_cct2gct))
  (mutate (from (c c t)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_cct2tct))
  (mutate (from (c c t)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_cct2agt))
  (mutate (from (c c t)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_cct2cgt))
  (mutate (from (c c t)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_cct2ggt))
  (mutate (from (c c t)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_cct2tgt))
  (mutate (from (c c t)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_cct2att))
  (mutate (from (c c t)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_cct2ctt))
  (mutate (from (c c t)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_cct2gtt))
  (mutate (from (c c t)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_cct2ttt))
  (mutate (from (g c t)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_gct2aaa))
  (mutate (from (g c t)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_gct2caa))
  (mutate (from (g c t)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_gct2gaa))
  (mutate (from (g c t)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_gct2taa))
  (mutate (from (g c t)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_gct2aca))
  (mutate (from (g c t)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_gct2cca))
  (mutate (from (g c t)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_gct2gca))
  (mutate (from (g c t)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_gct2tca))
  (mutate (from (g c t)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_gct2aga))
  (mutate (from (g c t)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_gct2cga))
  (mutate (from (g c t)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_gct2gga))
  (mutate (from (g c t)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_gct2tga))
  (mutate (from (g c t)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_gct2ata))
  (mutate (from (g c t)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_gct2cta))
  (mutate (from (g c t)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_gct2gta))
  (mutate (from (g c t)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_gct2tta))
  (mutate (from (g c t)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_gct2aac))
  (mutate (from (g c t)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_gct2cac))
  (mutate (from (g c t)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_gct2gac))
  (mutate (from (g c t)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_gct2tac))
  (mutate (from (g c t)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_gct2acc))
  (mutate (from (g c t)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_gct2ccc))
  (mutate (from (g c t)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_gct2gcc))
  (mutate (from (g c t)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_gct2tcc))
  (mutate (from (g c t)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_gct2agc))
  (mutate (from (g c t)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_gct2cgc))
  (mutate (from (g c t)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_gct2ggc))
  (mutate (from (g c t)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_gct2tgc))
  (mutate (from (g c t)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_gct2atc))
  (mutate (from (g c t)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_gct2ctc))
  (mutate (from (g c t)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_gct2gtc))
  (mutate (from (g c t)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_gct2ttc))
  (mutate (from (g c t)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_gct2aag))
  (mutate (from (g c t)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_gct2cag))
  (mutate (from (g c t)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_gct2gag))
  (mutate (from (g c t)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_gct2tag))
  (mutate (from (g c t)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_gct2acg))
  (mutate (from (g c t)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_gct2ccg))
  (mutate (from (g c t)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_gct2gcg))
  (mutate (from (g c t)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_gct2tcg))
  (mutate (from (g c t)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_gct2agg))
  (mutate (from (g c t)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_gct2cgg))
  (mutate (from (g c t)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_gct2ggg))
  (mutate (from (g c t)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_gct2tgg))
  (mutate (from (g c t)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_gct2atg))
  (mutate (from (g c t)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_gct2ctg))
  (mutate (from (g c t)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_gct2gtg))
  (mutate (from (g c t)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_gct2ttg))
  (mutate (from (g c t)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_gct2aat))
  (mutate (from (g c t)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_gct2cat))
  (mutate (from (g c t)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_gct2gat))
  (mutate (from (g c t)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_gct2tat))
  (mutate (from (g c t)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_gct2act))
  (mutate (from (g c t)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_gct2cct))
  (mutate (from (g c t)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_gct2tct))
  (mutate (from (g c t)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_gct2agt))
  (mutate (from (g c t)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_gct2cgt))
  (mutate (from (g c t)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_gct2ggt))
  (mutate (from (g c t)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_gct2tgt))
  (mutate (from (g c t)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_gct2att))
  (mutate (from (g c t)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_gct2ctt))
  (mutate (from (g c t)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_gct2gtt))
  (mutate (from (g c t)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_gct2ttt))
  (mutate (from (t c t)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_tct2aaa))
  (mutate (from (t c t)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_tct2caa))
  (mutate (from (t c t)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_tct2gaa))
  (mutate (from (t c t)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_tct2taa))
  (mutate (from (t c t)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_tct2aca))
  (mutate (from (t c t)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_tct2cca))
  (mutate (from (t c t)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_tct2gca))
  (mutate (from (t c t)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_tct2tca))
  (mutate (from (t c t)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_tct2aga))
  (mutate (from (t c t)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_tct2cga))
  (mutate (from (t c t)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_tct2gga))
  (mutate (from (t c t)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_tct2tga))
  (mutate (from (t c t)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_tct2ata))
  (mutate (from (t c t)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_tct2cta))
  (mutate (from (t c t)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_tct2gta))
  (mutate (from (t c t)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_tct2tta))
  (mutate (from (t c t)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_tct2aac))
  (mutate (from (t c t)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_tct2cac))
  (mutate (from (t c t)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_tct2gac))
  (mutate (from (t c t)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_tct2tac))
  (mutate (from (t c t)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_tct2acc))
  (mutate (from (t c t)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_tct2ccc))
  (mutate (from (t c t)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_tct2gcc))
  (mutate (from (t c t)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_tct2tcc))
  (mutate (from (t c t)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_tct2agc))
  (mutate (from (t c t)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_tct2cgc))
  (mutate (from (t c t)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_tct2ggc))
  (mutate (from (t c t)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_tct2tgc))
  (mutate (from (t c t)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_tct2atc))
  (mutate (from (t c t)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_tct2ctc))
  (mutate (from (t c t)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_tct2gtc))
  (mutate (from (t c t)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_tct2ttc))
  (mutate (from (t c t)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_tct2aag))
  (mutate (from (t c t)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_tct2cag))
  (mutate (from (t c t)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_tct2gag))
  (mutate (from (t c t)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_tct2tag))
  (mutate (from (t c t)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_tct2acg))
  (mutate (from (t c t)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_tct2ccg))
  (mutate (from (t c t)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_tct2gcg))
  (mutate (from (t c t)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_tct2tcg))
  (mutate (from (t c t)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_tct2agg))
  (mutate (from (t c t)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_tct2cgg))
  (mutate (from (t c t)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_tct2ggg))
  (mutate (from (t c t)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_tct2tgg))
  (mutate (from (t c t)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_tct2atg))
  (mutate (from (t c t)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_tct2ctg))
  (mutate (from (t c t)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_tct2gtg))
  (mutate (from (t c t)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_tct2ttg))
  (mutate (from (t c t)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_tct2aat))
  (mutate (from (t c t)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_tct2cat))
  (mutate (from (t c t)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_tct2gat))
  (mutate (from (t c t)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_tct2tat))
  (mutate (from (t c t)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_tct2act))
  (mutate (from (t c t)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_tct2cct))
  (mutate (from (t c t)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_tct2gct))
  (mutate (from (t c t)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_tct2agt))
  (mutate (from (t c t)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_tct2cgt))
  (mutate (from (t c t)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_tct2ggt))
  (mutate (from (t c t)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_tct2tgt))
  (mutate (from (t c t)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_tct2att))
  (mutate (from (t c t)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_tct2ctt))
  (mutate (from (t c t)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_tct2gtt))
  (mutate (from (t c t)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_tct2ttt))
  (mutate (from (a g t)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_agt2aaa))
  (mutate (from (a g t)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_agt2caa))
  (mutate (from (a g t)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_agt2gaa))
  (mutate (from (a g t)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_agt2taa))
  (mutate (from (a g t)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_agt2aca))
  (mutate (from (a g t)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_agt2cca))
  (mutate (from (a g t)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_agt2gca))
  (mutate (from (a g t)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_agt2tca))
  (mutate (from (a g t)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_agt2aga))
  (mutate (from (a g t)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_agt2cga))
  (mutate (from (a g t)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_agt2gga))
  (mutate (from (a g t)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_agt2tga))
  (mutate (from (a g t)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_agt2ata))
  (mutate (from (a g t)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_agt2cta))
  (mutate (from (a g t)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_agt2gta))
  (mutate (from (a g t)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_agt2tta))
  (mutate (from (a g t)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_agt2aac))
  (mutate (from (a g t)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_agt2cac))
  (mutate (from (a g t)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_agt2gac))
  (mutate (from (a g t)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_agt2tac))
  (mutate (from (a g t)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_agt2acc))
  (mutate (from (a g t)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_agt2ccc))
  (mutate (from (a g t)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_agt2gcc))
  (mutate (from (a g t)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_agt2tcc))
  (mutate (from (a g t)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_agt2agc))
  (mutate (from (a g t)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_agt2cgc))
  (mutate (from (a g t)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_agt2ggc))
  (mutate (from (a g t)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_agt2tgc))
  (mutate (from (a g t)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_agt2atc))
  (mutate (from (a g t)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_agt2ctc))
  (mutate (from (a g t)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_agt2gtc))
  (mutate (from (a g t)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_agt2ttc))
  (mutate (from (a g t)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_agt2aag))
  (mutate (from (a g t)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_agt2cag))
  (mutate (from (a g t)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_agt2gag))
  (mutate (from (a g t)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_agt2tag))
  (mutate (from (a g t)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_agt2acg))
  (mutate (from (a g t)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_agt2ccg))
  (mutate (from (a g t)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_agt2gcg))
  (mutate (from (a g t)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_agt2tcg))
  (mutate (from (a g t)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_agt2agg))
  (mutate (from (a g t)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_agt2cgg))
  (mutate (from (a g t)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_agt2ggg))
  (mutate (from (a g t)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_agt2tgg))
  (mutate (from (a g t)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_agt2atg))
  (mutate (from (a g t)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_agt2ctg))
  (mutate (from (a g t)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_agt2gtg))
  (mutate (from (a g t)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_agt2ttg))
  (mutate (from (a g t)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_agt2aat))
  (mutate (from (a g t)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_agt2cat))
  (mutate (from (a g t)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_agt2gat))
  (mutate (from (a g t)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_agt2tat))
  (mutate (from (a g t)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_agt2act))
  (mutate (from (a g t)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_agt2cct))
  (mutate (from (a g t)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_agt2gct))
  (mutate (from (a g t)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_agt2tct))
  (mutate (from (a g t)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_agt2cgt))
  (mutate (from (a g t)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_agt2ggt))
  (mutate (from (a g t)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_agt2tgt))
  (mutate (from (a g t)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_agt2att))
  (mutate (from (a g t)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_agt2ctt))
  (mutate (from (a g t)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_agt2gtt))
  (mutate (from (a g t)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_agt2ttt))
  (mutate (from (c g t)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2aaa))
  (mutate (from (c g t)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2caa))
  (mutate (from (c g t)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2gaa))
  (mutate (from (c g t)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2taa))
  (mutate (from (c g t)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2aca))
  (mutate (from (c g t)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2cca))
  (mutate (from (c g t)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2gca))
  (mutate (from (c g t)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2tca))
  (mutate (from (c g t)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2aga))
  (mutate (from (c g t)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2cga))
  (mutate (from (c g t)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2gga))
  (mutate (from (c g t)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2tga))
  (mutate (from (c g t)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2ata))
  (mutate (from (c g t)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2cta))
  (mutate (from (c g t)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2gta))
  (mutate (from (c g t)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2tta))
  (mutate (from (c g t)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2aac))
  (mutate (from (c g t)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2cac))
  (mutate (from (c g t)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2gac))
  (mutate (from (c g t)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2tac))
  (mutate (from (c g t)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2acc))
  (mutate (from (c g t)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2ccc))
  (mutate (from (c g t)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2gcc))
  (mutate (from (c g t)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2tcc))
  (mutate (from (c g t)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2agc))
  (mutate (from (c g t)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2cgc))
  (mutate (from (c g t)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2ggc))
  (mutate (from (c g t)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2tgc))
  (mutate (from (c g t)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2atc))
  (mutate (from (c g t)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2ctc))
  (mutate (from (c g t)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2gtc))
  (mutate (from (c g t)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2ttc))
  (mutate (from (c g t)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2aag))
  (mutate (from (c g t)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2cag))
  (mutate (from (c g t)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2gag))
  (mutate (from (c g t)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2tag))
  (mutate (from (c g t)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2acg))
  (mutate (from (c g t)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2ccg))
  (mutate (from (c g t)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2gcg))
  (mutate (from (c g t)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2tcg))
  (mutate (from (c g t)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2agg))
  (mutate (from (c g t)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2cgg))
  (mutate (from (c g t)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2ggg))
  (mutate (from (c g t)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2tgg))
  (mutate (from (c g t)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2atg))
  (mutate (from (c g t)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2ctg))
  (mutate (from (c g t)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2gtg))
  (mutate (from (c g t)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2ttg))
  (mutate (from (c g t)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2aat))
  (mutate (from (c g t)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2cat))
  (mutate (from (c g t)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2gat))
  (mutate (from (c g t)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2tat))
  (mutate (from (c g t)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2act))
  (mutate (from (c g t)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2cct))
  (mutate (from (c g t)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2gct))
  (mutate (from (c g t)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2tct))
  (mutate (from (c g t)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2agt))
  (mutate (from (c g t)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2ggt))
  (mutate (from (c g t)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2tgt))
  (mutate (from (c g t)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2att))
  (mutate (from (c g t)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2ctt))
  (mutate (from (c g t)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2gtt))
  (mutate (from (c g t)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_cgt2ttt))
  (mutate (from (g g t)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2aaa))
  (mutate (from (g g t)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2caa))
  (mutate (from (g g t)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2gaa))
  (mutate (from (g g t)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2taa))
  (mutate (from (g g t)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2aca))
  (mutate (from (g g t)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2cca))
  (mutate (from (g g t)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2gca))
  (mutate (from (g g t)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2tca))
  (mutate (from (g g t)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2aga))
  (mutate (from (g g t)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2cga))
  (mutate (from (g g t)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2gga))
  (mutate (from (g g t)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2tga))
  (mutate (from (g g t)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2ata))
  (mutate (from (g g t)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2cta))
  (mutate (from (g g t)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2gta))
  (mutate (from (g g t)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2tta))
  (mutate (from (g g t)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2aac))
  (mutate (from (g g t)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2cac))
  (mutate (from (g g t)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2gac))
  (mutate (from (g g t)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2tac))
  (mutate (from (g g t)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2acc))
  (mutate (from (g g t)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2ccc))
  (mutate (from (g g t)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2gcc))
  (mutate (from (g g t)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2tcc))
  (mutate (from (g g t)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2agc))
  (mutate (from (g g t)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2cgc))
  (mutate (from (g g t)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2ggc))
  (mutate (from (g g t)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2tgc))
  (mutate (from (g g t)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2atc))
  (mutate (from (g g t)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2ctc))
  (mutate (from (g g t)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2gtc))
  (mutate (from (g g t)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2ttc))
  (mutate (from (g g t)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2aag))
  (mutate (from (g g t)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2cag))
  (mutate (from (g g t)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2gag))
  (mutate (from (g g t)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2tag))
  (mutate (from (g g t)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2acg))
  (mutate (from (g g t)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2ccg))
  (mutate (from (g g t)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2gcg))
  (mutate (from (g g t)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2tcg))
  (mutate (from (g g t)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2agg))
  (mutate (from (g g t)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2cgg))
  (mutate (from (g g t)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2ggg))
  (mutate (from (g g t)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2tgg))
  (mutate (from (g g t)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2atg))
  (mutate (from (g g t)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2ctg))
  (mutate (from (g g t)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2gtg))
  (mutate (from (g g t)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2ttg))
  (mutate (from (g g t)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2aat))
  (mutate (from (g g t)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2cat))
  (mutate (from (g g t)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2gat))
  (mutate (from (g g t)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2tat))
  (mutate (from (g g t)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2act))
  (mutate (from (g g t)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2cct))
  (mutate (from (g g t)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2gct))
  (mutate (from (g g t)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2tct))
  (mutate (from (g g t)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2agt))
  (mutate (from (g g t)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2cgt))
  (mutate (from (g g t)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2tgt))
  (mutate (from (g g t)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2att))
  (mutate (from (g g t)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2ctt))
  (mutate (from (g g t)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2gtt))
  (mutate (from (g g t)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_ggt2ttt))
  (mutate (from (t g t)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2aaa))
  (mutate (from (t g t)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2caa))
  (mutate (from (t g t)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2gaa))
  (mutate (from (t g t)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2taa))
  (mutate (from (t g t)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2aca))
  (mutate (from (t g t)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2cca))
  (mutate (from (t g t)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2gca))
  (mutate (from (t g t)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2tca))
  (mutate (from (t g t)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2aga))
  (mutate (from (t g t)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2cga))
  (mutate (from (t g t)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2gga))
  (mutate (from (t g t)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2tga))
  (mutate (from (t g t)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2ata))
  (mutate (from (t g t)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2cta))
  (mutate (from (t g t)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2gta))
  (mutate (from (t g t)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2tta))
  (mutate (from (t g t)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2aac))
  (mutate (from (t g t)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2cac))
  (mutate (from (t g t)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2gac))
  (mutate (from (t g t)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2tac))
  (mutate (from (t g t)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2acc))
  (mutate (from (t g t)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2ccc))
  (mutate (from (t g t)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2gcc))
  (mutate (from (t g t)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2tcc))
  (mutate (from (t g t)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2agc))
  (mutate (from (t g t)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2cgc))
  (mutate (from (t g t)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2ggc))
  (mutate (from (t g t)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2tgc))
  (mutate (from (t g t)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2atc))
  (mutate (from (t g t)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2ctc))
  (mutate (from (t g t)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2gtc))
  (mutate (from (t g t)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2ttc))
  (mutate (from (t g t)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2aag))
  (mutate (from (t g t)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2cag))
  (mutate (from (t g t)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2gag))
  (mutate (from (t g t)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2tag))
  (mutate (from (t g t)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2acg))
  (mutate (from (t g t)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2ccg))
  (mutate (from (t g t)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2gcg))
  (mutate (from (t g t)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2tcg))
  (mutate (from (t g t)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2agg))
  (mutate (from (t g t)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2cgg))
  (mutate (from (t g t)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2ggg))
  (mutate (from (t g t)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2tgg))
  (mutate (from (t g t)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2atg))
  (mutate (from (t g t)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2ctg))
  (mutate (from (t g t)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2gtg))
  (mutate (from (t g t)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2ttg))
  (mutate (from (t g t)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2aat))
  (mutate (from (t g t)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2cat))
  (mutate (from (t g t)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2gat))
  (mutate (from (t g t)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2tat))
  (mutate (from (t g t)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2act))
  (mutate (from (t g t)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2cct))
  (mutate (from (t g t)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2gct))
  (mutate (from (t g t)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2tct))
  (mutate (from (t g t)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2agt))
  (mutate (from (t g t)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2cgt))
  (mutate (from (t g t)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2ggt))
  (mutate (from (t g t)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2att))
  (mutate (from (t g t)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2ctt))
  (mutate (from (t g t)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2gtt))
  (mutate (from (t g t)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_tgt2ttt))
  (mutate (from (a t t)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_att2aaa))
  (mutate (from (a t t)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_att2caa))
  (mutate (from (a t t)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_att2gaa))
  (mutate (from (a t t)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_att2taa))
  (mutate (from (a t t)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_att2aca))
  (mutate (from (a t t)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_att2cca))
  (mutate (from (a t t)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_att2gca))
  (mutate (from (a t t)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_att2tca))
  (mutate (from (a t t)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_att2aga))
  (mutate (from (a t t)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_att2cga))
  (mutate (from (a t t)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_att2gga))
  (mutate (from (a t t)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_att2tga))
  (mutate (from (a t t)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_att2ata))
  (mutate (from (a t t)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_att2cta))
  (mutate (from (a t t)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_att2gta))
  (mutate (from (a t t)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_att2tta))
  (mutate (from (a t t)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_att2aac))
  (mutate (from (a t t)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_att2cac))
  (mutate (from (a t t)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_att2gac))
  (mutate (from (a t t)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_att2tac))
  (mutate (from (a t t)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_att2acc))
  (mutate (from (a t t)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_att2ccc))
  (mutate (from (a t t)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_att2gcc))
  (mutate (from (a t t)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_att2tcc))
  (mutate (from (a t t)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_att2agc))
  (mutate (from (a t t)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_att2cgc))
  (mutate (from (a t t)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_att2ggc))
  (mutate (from (a t t)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_att2tgc))
  (mutate (from (a t t)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_att2atc))
  (mutate (from (a t t)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_att2ctc))
  (mutate (from (a t t)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_att2gtc))
  (mutate (from (a t t)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_att2ttc))
  (mutate (from (a t t)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_att2aag))
  (mutate (from (a t t)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_att2cag))
  (mutate (from (a t t)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_att2gag))
  (mutate (from (a t t)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_att2tag))
  (mutate (from (a t t)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_att2acg))
  (mutate (from (a t t)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_att2ccg))
  (mutate (from (a t t)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_att2gcg))
  (mutate (from (a t t)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_att2tcg))
  (mutate (from (a t t)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_att2agg))
  (mutate (from (a t t)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_att2cgg))
  (mutate (from (a t t)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_att2ggg))
  (mutate (from (a t t)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_att2tgg))
  (mutate (from (a t t)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_att2atg))
  (mutate (from (a t t)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_att2ctg))
  (mutate (from (a t t)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_att2gtg))
  (mutate (from (a t t)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_att2ttg))
  (mutate (from (a t t)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_att2aat))
  (mutate (from (a t t)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_att2cat))
  (mutate (from (a t t)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_att2gat))
  (mutate (from (a t t)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_att2tat))
  (mutate (from (a t t)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_att2act))
  (mutate (from (a t t)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_att2cct))
  (mutate (from (a t t)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_att2gct))
  (mutate (from (a t t)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_att2tct))
  (mutate (from (a t t)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_att2agt))
  (mutate (from (a t t)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_att2cgt))
  (mutate (from (a t t)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_att2ggt))
  (mutate (from (a t t)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_att2tgt))
  (mutate (from (a t t)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_att2ctt))
  (mutate (from (a t t)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_att2gtt))
  (mutate (from (a t t)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_att2ttt))
  (mutate (from (c t t)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2aaa))
  (mutate (from (c t t)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2caa))
  (mutate (from (c t t)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2gaa))
  (mutate (from (c t t)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2taa))
  (mutate (from (c t t)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2aca))
  (mutate (from (c t t)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2cca))
  (mutate (from (c t t)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2gca))
  (mutate (from (c t t)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2tca))
  (mutate (from (c t t)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2aga))
  (mutate (from (c t t)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2cga))
  (mutate (from (c t t)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2gga))
  (mutate (from (c t t)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2tga))
  (mutate (from (c t t)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2ata))
  (mutate (from (c t t)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2cta))
  (mutate (from (c t t)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2gta))
  (mutate (from (c t t)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2tta))
  (mutate (from (c t t)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2aac))
  (mutate (from (c t t)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2cac))
  (mutate (from (c t t)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2gac))
  (mutate (from (c t t)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2tac))
  (mutate (from (c t t)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2acc))
  (mutate (from (c t t)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2ccc))
  (mutate (from (c t t)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2gcc))
  (mutate (from (c t t)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2tcc))
  (mutate (from (c t t)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2agc))
  (mutate (from (c t t)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2cgc))
  (mutate (from (c t t)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2ggc))
  (mutate (from (c t t)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2tgc))
  (mutate (from (c t t)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2atc))
  (mutate (from (c t t)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2ctc))
  (mutate (from (c t t)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2gtc))
  (mutate (from (c t t)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2ttc))
  (mutate (from (c t t)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2aag))
  (mutate (from (c t t)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2cag))
  (mutate (from (c t t)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2gag))
  (mutate (from (c t t)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2tag))
  (mutate (from (c t t)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2acg))
  (mutate (from (c t t)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2ccg))
  (mutate (from (c t t)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2gcg))
  (mutate (from (c t t)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2tcg))
  (mutate (from (c t t)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2agg))
  (mutate (from (c t t)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2cgg))
  (mutate (from (c t t)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2ggg))
  (mutate (from (c t t)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2tgg))
  (mutate (from (c t t)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2atg))
  (mutate (from (c t t)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2ctg))
  (mutate (from (c t t)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2gtg))
  (mutate (from (c t t)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2ttg))
  (mutate (from (c t t)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2aat))
  (mutate (from (c t t)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2cat))
  (mutate (from (c t t)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2gat))
  (mutate (from (c t t)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2tat))
  (mutate (from (c t t)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2act))
  (mutate (from (c t t)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2cct))
  (mutate (from (c t t)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2gct))
  (mutate (from (c t t)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2tct))
  (mutate (from (c t t)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2agt))
  (mutate (from (c t t)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2cgt))
  (mutate (from (c t t)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2ggt))
  (mutate (from (c t t)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2tgt))
  (mutate (from (c t t)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2att))
  (mutate (from (c t t)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2gtt))
  (mutate (from (c t t)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_ctt2ttt))
  (mutate (from (g t t)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2aaa))
  (mutate (from (g t t)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2caa))
  (mutate (from (g t t)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2gaa))
  (mutate (from (g t t)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2taa))
  (mutate (from (g t t)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2aca))
  (mutate (from (g t t)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2cca))
  (mutate (from (g t t)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2gca))
  (mutate (from (g t t)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2tca))
  (mutate (from (g t t)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2aga))
  (mutate (from (g t t)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2cga))
  (mutate (from (g t t)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2gga))
  (mutate (from (g t t)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2tga))
  (mutate (from (g t t)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2ata))
  (mutate (from (g t t)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2cta))
  (mutate (from (g t t)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2gta))
  (mutate (from (g t t)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2tta))
  (mutate (from (g t t)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2aac))
  (mutate (from (g t t)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2cac))
  (mutate (from (g t t)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2gac))
  (mutate (from (g t t)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2tac))
  (mutate (from (g t t)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2acc))
  (mutate (from (g t t)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2ccc))
  (mutate (from (g t t)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2gcc))
  (mutate (from (g t t)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2tcc))
  (mutate (from (g t t)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2agc))
  (mutate (from (g t t)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2cgc))
  (mutate (from (g t t)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2ggc))
  (mutate (from (g t t)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2tgc))
  (mutate (from (g t t)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2atc))
  (mutate (from (g t t)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2ctc))
  (mutate (from (g t t)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2gtc))
  (mutate (from (g t t)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2ttc))
  (mutate (from (g t t)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2aag))
  (mutate (from (g t t)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2cag))
  (mutate (from (g t t)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2gag))
  (mutate (from (g t t)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2tag))
  (mutate (from (g t t)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2acg))
  (mutate (from (g t t)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2ccg))
  (mutate (from (g t t)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2gcg))
  (mutate (from (g t t)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2tcg))
  (mutate (from (g t t)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2agg))
  (mutate (from (g t t)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2cgg))
  (mutate (from (g t t)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2ggg))
  (mutate (from (g t t)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2tgg))
  (mutate (from (g t t)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2atg))
  (mutate (from (g t t)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2ctg))
  (mutate (from (g t t)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2gtg))
  (mutate (from (g t t)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2ttg))
  (mutate (from (g t t)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2aat))
  (mutate (from (g t t)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2cat))
  (mutate (from (g t t)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2gat))
  (mutate (from (g t t)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2tat))
  (mutate (from (g t t)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2act))
  (mutate (from (g t t)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2cct))
  (mutate (from (g t t)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2gct))
  (mutate (from (g t t)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2tct))
  (mutate (from (g t t)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2agt))
  (mutate (from (g t t)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2cgt))
  (mutate (from (g t t)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2ggt))
  (mutate (from (g t t)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2tgt))
  (mutate (from (g t t)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2att))
  (mutate (from (g t t)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2ctt))
  (mutate (from (g t t)) (to (t t t)) (rate coding_rate_scale * COD1_COD2_COD3_gtt2ttt))
  (mutate (from (t t t)) (to (a a a)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2aaa))
  (mutate (from (t t t)) (to (c a a)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2caa))
  (mutate (from (t t t)) (to (g a a)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2gaa))
  (mutate (from (t t t)) (to (t a a)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2taa))
  (mutate (from (t t t)) (to (a c a)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2aca))
  (mutate (from (t t t)) (to (c c a)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2cca))
  (mutate (from (t t t)) (to (g c a)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2gca))
  (mutate (from (t t t)) (to (t c a)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2tca))
  (mutate (from (t t t)) (to (a g a)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2aga))
  (mutate (from (t t t)) (to (c g a)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2cga))
  (mutate (from (t t t)) (to (g g a)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2gga))
  (mutate (from (t t t)) (to (t g a)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2tga))
  (mutate (from (t t t)) (to (a t a)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2ata))
  (mutate (from (t t t)) (to (c t a)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2cta))
  (mutate (from (t t t)) (to (g t a)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2gta))
  (mutate (from (t t t)) (to (t t a)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2tta))
  (mutate (from (t t t)) (to (a a c)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2aac))
  (mutate (from (t t t)) (to (c a c)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2cac))
  (mutate (from (t t t)) (to (g a c)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2gac))
  (mutate (from (t t t)) (to (t a c)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2tac))
  (mutate (from (t t t)) (to (a c c)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2acc))
  (mutate (from (t t t)) (to (c c c)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2ccc))
  (mutate (from (t t t)) (to (g c c)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2gcc))
  (mutate (from (t t t)) (to (t c c)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2tcc))
  (mutate (from (t t t)) (to (a g c)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2agc))
  (mutate (from (t t t)) (to (c g c)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2cgc))
  (mutate (from (t t t)) (to (g g c)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2ggc))
  (mutate (from (t t t)) (to (t g c)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2tgc))
  (mutate (from (t t t)) (to (a t c)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2atc))
  (mutate (from (t t t)) (to (c t c)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2ctc))
  (mutate (from (t t t)) (to (g t c)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2gtc))
  (mutate (from (t t t)) (to (t t c)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2ttc))
  (mutate (from (t t t)) (to (a a g)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2aag))
  (mutate (from (t t t)) (to (c a g)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2cag))
  (mutate (from (t t t)) (to (g a g)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2gag))
  (mutate (from (t t t)) (to (t a g)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2tag))
  (mutate (from (t t t)) (to (a c g)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2acg))
  (mutate (from (t t t)) (to (c c g)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2ccg))
  (mutate (from (t t t)) (to (g c g)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2gcg))
  (mutate (from (t t t)) (to (t c g)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2tcg))
  (mutate (from (t t t)) (to (a g g)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2agg))
  (mutate (from (t t t)) (to (c g g)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2cgg))
  (mutate (from (t t t)) (to (g g g)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2ggg))
  (mutate (from (t t t)) (to (t g g)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2tgg))
  (mutate (from (t t t)) (to (a t g)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2atg))
  (mutate (from (t t t)) (to (c t g)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2ctg))
  (mutate (from (t t t)) (to (g t g)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2gtg))
  (mutate (from (t t t)) (to (t t g)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2ttg))
  (mutate (from (t t t)) (to (a a t)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2aat))
  (mutate (from (t t t)) (to (c a t)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2cat))
  (mutate (from (t t t)) (to (g a t)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2gat))
  (mutate (from (t t t)) (to (t a t)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2tat))
  (mutate (from (t t t)) (to (a c t)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2act))
  (mutate (from (t t t)) (to (c c t)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2cct))
  (mutate (from (t t t)) (to (g c t)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2gct))
  (mutate (from (t t t)) (to (t c t)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2tct))
  (mutate (from (t t t)) (to (a g t)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2agt))
  (mutate (from (t t t)) (to (c g t)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2cgt))
  (mutate (from (t t t)) (to (g g t)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2ggt))
  (mutate (from (t t t)) (to (t g t)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2tgt))
  (mutate (from (t t t)) (to (a t t)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2att))
  (mutate (from (t t t)) (to (c t t)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2ctt))
  (mutate (from (t t t)) (to (g t t)) (rate coding_rate_scale * COD1_COD2_COD3_ttt2gtt))

  (mutate (from (a a a)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_aaa2gapgapgap))
  (mutate (from (a a c)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_aac2gapgapgap))
  (mutate (from (a a g)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_aag2gapgapgap))
  (mutate (from (a a t)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_aat2gapgapgap))
  (mutate (from (a c a)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_aca2gapgapgap))
  (mutate (from (a c c)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_acc2gapgapgap))
  (mutate (from (a c g)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_acg2gapgapgap))
  (mutate (from (a c t)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_act2gapgapgap))
  (mutate (from (a g a)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_aga2gapgapgap))
  (mutate (from (a g c)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_agc2gapgapgap))
  (mutate (from (a g g)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_agg2gapgapgap))
  (mutate (from (a g t)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_agt2gapgapgap))
  (mutate (from (a t a)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_ata2gapgapgap))
  (mutate (from (a t c)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_atc2gapgapgap))
  (mutate (from (a t g)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_atg2gapgapgap))
  (mutate (from (a t t)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_att2gapgapgap))
  (mutate (from (c a a)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_caa2gapgapgap))
  (mutate (from (c a c)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_cac2gapgapgap))
  (mutate (from (c a g)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_cag2gapgapgap))
  (mutate (from (c a t)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_cat2gapgapgap))
  (mutate (from (c c a)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_cca2gapgapgap))
  (mutate (from (c c c)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_ccc2gapgapgap))
  (mutate (from (c c g)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_ccg2gapgapgap))
  (mutate (from (c c t)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_cct2gapgapgap))
  (mutate (from (c g a)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_cga2gapgapgap))
  (mutate (from (c g c)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_cgc2gapgapgap))
  (mutate (from (c g g)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_cgg2gapgapgap))
  (mutate (from (c g t)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_cgt2gapgapgap))
  (mutate (from (c t a)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_cta2gapgapgap))
  (mutate (from (c t c)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_ctc2gapgapgap))
  (mutate (from (c t g)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_ctg2gapgapgap))
  (mutate (from (c t t)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_ctt2gapgapgap))
  (mutate (from (g a a)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_gaa2gapgapgap))
  (mutate (from (g a c)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_gac2gapgapgap))
  (mutate (from (g a g)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_gag2gapgapgap))
  (mutate (from (g a t)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_gat2gapgapgap))
  (mutate (from (g c a)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_gca2gapgapgap))
  (mutate (from (g c c)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_gcc2gapgapgap))
  (mutate (from (g c g)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_gcg2gapgapgap))
  (mutate (from (g c t)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_gct2gapgapgap))
  (mutate (from (g g a)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_gga2gapgapgap))
  (mutate (from (g g c)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_ggc2gapgapgap))
  (mutate (from (g g g)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_ggg2gapgapgap))
  (mutate (from (g g t)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_ggt2gapgapgap))
  (mutate (from (g t a)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_gta2gapgapgap))
  (mutate (from (g t c)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_gtc2gapgapgap))
  (mutate (from (g t g)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_gtg2gapgapgap))
  (mutate (from (g t t)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_gtt2gapgapgap))
  (mutate (from (t a a)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_taa2gapgapgap))
  (mutate (from (t a c)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_tac2gapgapgap))
  (mutate (from (t a g)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_tag2gapgapgap))
  (mutate (from (t a t)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_tat2gapgapgap))
  (mutate (from (t c a)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_tca2gapgapgap))
  (mutate (from (t c c)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_tcc2gapgapgap))
  (mutate (from (t c g)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_tcg2gapgapgap))
  (mutate (from (t c t)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_tct2gapgapgap))
  (mutate (from (t g a)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_tga2gapgapgap))
  (mutate (from (t g c)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_tgc2gapgapgap))
  (mutate (from (t g g)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_tgg2gapgapgap))
  (mutate (from (t g t)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_tgt2gapgapgap))
  (mutate (from (t t a)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_tta2gapgapgap))
  (mutate (from (t t c)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_ttc2gapgapgap))
  (mutate (from (t t g)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_ttg2gapgapgap))
  (mutate (from (t t t)) (to (_ _ _)) (rate coding_rate_scale * coding_gap_rate * COD1_COD2_COD3_ttt2gapgapgap))

 )  ;; end chain COD1 COD2 COD3

;;;;;;;;;;;;;;;;
;; PSEUDOGENE ;;
;;;;;;;;;;;;;;;;

 ;; pseudogene chain
 (chain
  (update-policy parametric)
  (terminal (PG1 PG2 PG3))

  ;; initial probability distribution
  (initial (state (a a a)) (prob PG1_PG2_PG3_aaa))
  (initial (state (c a a)) (prob PG1_PG2_PG3_caa))
  (initial (state (g a a)) (prob PG1_PG2_PG3_gaa))
  (initial (state (t a a)) (prob PG1_PG2_PG3_taa))
  (initial (state (a c a)) (prob PG1_PG2_PG3_aca))
  (initial (state (c c a)) (prob PG1_PG2_PG3_cca))
  (initial (state (g c a)) (prob PG1_PG2_PG3_gca))
  (initial (state (t c a)) (prob PG1_PG2_PG3_tca))
  (initial (state (a g a)) (prob PG1_PG2_PG3_aga))
  (initial (state (c g a)) (prob PG1_PG2_PG3_cga))
  (initial (state (g g a)) (prob PG1_PG2_PG3_gga))
  (initial (state (t g a)) (prob PG1_PG2_PG3_tga))
  (initial (state (a t a)) (prob PG1_PG2_PG3_ata))
  (initial (state (c t a)) (prob PG1_PG2_PG3_cta))
  (initial (state (g t a)) (prob PG1_PG2_PG3_gta))
  (initial (state (t t a)) (prob PG1_PG2_PG3_tta))
  (initial (state (a a c)) (prob PG1_PG2_PG3_aac))
  (initial (state (c a c)) (prob PG1_PG2_PG3_cac))
  (initial (state (g a c)) (prob PG1_PG2_PG3_gac))
  (initial (state (t a c)) (prob PG1_PG2_PG3_tac))
  (initial (state (a c c)) (prob PG1_PG2_PG3_acc))
  (initial (state (c c c)) (prob PG1_PG2_PG3_ccc))
  (initial (state (g c c)) (prob PG1_PG2_PG3_gcc))
  (initial (state (t c c)) (prob PG1_PG2_PG3_tcc))
  (initial (state (a g c)) (prob PG1_PG2_PG3_agc))
  (initial (state (c g c)) (prob PG1_PG2_PG3_cgc))
  (initial (state (g g c)) (prob PG1_PG2_PG3_ggc))
  (initial (state (t g c)) (prob PG1_PG2_PG3_tgc))
  (initial (state (a t c)) (prob PG1_PG2_PG3_atc))
  (initial (state (c t c)) (prob PG1_PG2_PG3_ctc))
  (initial (state (g t c)) (prob PG1_PG2_PG3_gtc))
  (initial (state (t t c)) (prob PG1_PG2_PG3_ttc))
  (initial (state (a a g)) (prob PG1_PG2_PG3_aag))
  (initial (state (c a g)) (prob PG1_PG2_PG3_cag))
  (initial (state (g a g)) (prob PG1_PG2_PG3_gag))
  (initial (state (t a g)) (prob PG1_PG2_PG3_tag))
  (initial (state (a c g)) (prob PG1_PG2_PG3_acg))
  (initial (state (c c g)) (prob PG1_PG2_PG3_ccg))
  (initial (state (g c g)) (prob PG1_PG2_PG3_gcg))
  (initial (state (t c g)) (prob PG1_PG2_PG3_tcg))
  (initial (state (a g g)) (prob PG1_PG2_PG3_agg))
  (initial (state (c g g)) (prob PG1_PG2_PG3_cgg))
  (initial (state (g g g)) (prob PG1_PG2_PG3_ggg))
  (initial (state (t g g)) (prob PG1_PG2_PG3_tgg))
  (initial (state (a t g)) (prob PG1_PG2_PG3_atg))
  (initial (state (c t g)) (prob PG1_PG2_PG3_ctg))
  (initial (state (g t g)) (prob PG1_PG2_PG3_gtg))
  (initial (state (t t g)) (prob PG1_PG2_PG3_ttg))
  (initial (state (a a t)) (prob PG1_PG2_PG3_aat))
  (initial (state (c a t)) (prob PG1_PG2_PG3_cat))
  (initial (state (g a t)) (prob PG1_PG2_PG3_gat))
  (initial (state (t a t)) (prob PG1_PG2_PG3_tat))
  (initial (state (a c t)) (prob PG1_PG2_PG3_act))
  (initial (state (c c t)) (prob PG1_PG2_PG3_cct))
  (initial (state (g c t)) (prob PG1_PG2_PG3_gct))
  (initial (state (t c t)) (prob PG1_PG2_PG3_tct))
  (initial (state (a g t)) (prob PG1_PG2_PG3_agt))
  (initial (state (c g t)) (prob PG1_PG2_PG3_cgt))
  (initial (state (g g t)) (prob PG1_PG2_PG3_ggt))
  (initial (state (t g t)) (prob PG1_PG2_PG3_tgt))
  (initial (state (a t t)) (prob PG1_PG2_PG3_att))
  (initial (state (c t t)) (prob PG1_PG2_PG3_ctt))
  (initial (state (g t t)) (prob PG1_PG2_PG3_gtt))
  (initial (state (t t t)) (prob PG1_PG2_PG3_ttt))

  ;; mutation rates
  (mutate (from (a a a)) (to (a a c)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (a a a)) (to (a a g)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (a a a)) (to (a a t)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (a a a)) (to (a a _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (a a a)) (to (a c a)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (a a a)) (to (a g a)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (a a a)) (to (a t a)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (a a a)) (to (a _ a)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (a a a)) (to (c a a)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (a a a)) (to (g a a)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (a a a)) (to (t a a)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (a a a)) (to (_ a a)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (a a c)) (to (a a a)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (a a c)) (to (a a g)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (a a c)) (to (a a t)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (a a c)) (to (a a _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (a a c)) (to (a c c)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (a a c)) (to (a g c)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (a a c)) (to (a t c)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (a a c)) (to (a _ c)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (a a c)) (to (c a c)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (a a c)) (to (g a c)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (a a c)) (to (t a c)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (a a c)) (to (_ a c)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (a a g)) (to (a a a)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (a a g)) (to (a a c)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (a a g)) (to (a a t)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (a a g)) (to (a a _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (a a g)) (to (a c g)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (a a g)) (to (a g g)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (a a g)) (to (a t g)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (a a g)) (to (a _ g)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (a a g)) (to (c a g)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (a a g)) (to (g a g)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (a a g)) (to (t a g)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (a a g)) (to (_ a g)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (a a t)) (to (a a a)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (a a t)) (to (a a c)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (a a t)) (to (a a g)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (a a t)) (to (a a _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (a a t)) (to (a c t)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (a a t)) (to (a g t)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (a a t)) (to (a t t)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (a a t)) (to (a _ t)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (a a t)) (to (c a t)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (a a t)) (to (g a t)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (a a t)) (to (t a t)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (a a t)) (to (_ a t)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (a a _)) (to (a a a)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (a a _)) (to (a a c)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (a a _)) (to (a a g)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (a a _)) (to (a a t)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (a a _)) (to (a c _)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (a a _)) (to (a g _)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (a a _)) (to (a t _)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (a a _)) (to (a _ _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (a a _)) (to (c a _)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (a a _)) (to (g a _)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (a a _)) (to (t a _)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (a a _)) (to (_ a _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (a c a)) (to (a a a)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (a c a)) (to (a c c)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (a c a)) (to (a c g)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (a c a)) (to (a c t)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (a c a)) (to (a c _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (a c a)) (to (a g a)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (a c a)) (to (a t a)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (a c a)) (to (a _ a)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (a c a)) (to (c c a)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (a c a)) (to (g c a)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (a c a)) (to (t c a)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (a c a)) (to (_ c a)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (a c c)) (to (a a c)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (a c c)) (to (a c a)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (a c c)) (to (a c g)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (a c c)) (to (a c t)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (a c c)) (to (a c _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (a c c)) (to (a g c)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (a c c)) (to (a t c)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (a c c)) (to (a _ c)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (a c c)) (to (c c c)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (a c c)) (to (g c c)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (a c c)) (to (t c c)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (a c c)) (to (_ c c)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (a c g)) (to (a a g)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (a c g)) (to (a c a)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (a c g)) (to (a c c)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (a c g)) (to (a c t)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (a c g)) (to (a c _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (a c g)) (to (a g g)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (a c g)) (to (a t g)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (a c g)) (to (a _ g)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (a c g)) (to (c c g)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (a c g)) (to (g c g)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (a c g)) (to (t c g)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (a c g)) (to (_ c g)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (a c t)) (to (a a t)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (a c t)) (to (a c a)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (a c t)) (to (a c c)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (a c t)) (to (a c g)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (a c t)) (to (a c _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (a c t)) (to (a g t)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (a c t)) (to (a t t)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (a c t)) (to (a _ t)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (a c t)) (to (c c t)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (a c t)) (to (g c t)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (a c t)) (to (t c t)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (a c t)) (to (_ c t)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (a c _)) (to (a a _)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (a c _)) (to (a c a)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (a c _)) (to (a c c)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (a c _)) (to (a c g)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (a c _)) (to (a c t)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (a c _)) (to (a g _)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (a c _)) (to (a t _)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (a c _)) (to (a _ _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (a c _)) (to (c c _)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (a c _)) (to (g c _)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (a c _)) (to (t c _)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (a c _)) (to (_ c _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (a g a)) (to (a a a)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (a g a)) (to (a c a)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (a g a)) (to (a g c)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (a g a)) (to (a g g)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (a g a)) (to (a g t)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (a g a)) (to (a g _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (a g a)) (to (a t a)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (a g a)) (to (a _ a)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (a g a)) (to (c g a)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (a g a)) (to (g g a)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (a g a)) (to (t g a)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (a g a)) (to (_ g a)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (a g c)) (to (a a c)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (a g c)) (to (a c c)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (a g c)) (to (a g a)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (a g c)) (to (a g g)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (a g c)) (to (a g t)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (a g c)) (to (a g _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (a g c)) (to (a t c)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (a g c)) (to (a _ c)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (a g c)) (to (c g c)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (a g c)) (to (g g c)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (a g c)) (to (t g c)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (a g c)) (to (_ g c)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (a g g)) (to (a a g)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (a g g)) (to (a c g)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (a g g)) (to (a g a)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (a g g)) (to (a g c)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (a g g)) (to (a g t)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (a g g)) (to (a g _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (a g g)) (to (a t g)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (a g g)) (to (a _ g)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (a g g)) (to (c g g)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (a g g)) (to (g g g)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (a g g)) (to (t g g)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (a g g)) (to (_ g g)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (a g t)) (to (a a t)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (a g t)) (to (a c t)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (a g t)) (to (a g a)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (a g t)) (to (a g c)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (a g t)) (to (a g g)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (a g t)) (to (a g _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (a g t)) (to (a t t)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (a g t)) (to (a _ t)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (a g t)) (to (c g t)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (a g t)) (to (g g t)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (a g t)) (to (t g t)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (a g t)) (to (_ g t)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (a g _)) (to (a a _)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (a g _)) (to (a c _)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (a g _)) (to (a g a)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (a g _)) (to (a g c)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (a g _)) (to (a g g)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (a g _)) (to (a g t)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (a g _)) (to (a t _)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (a g _)) (to (a _ _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (a g _)) (to (c g _)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (a g _)) (to (g g _)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (a g _)) (to (t g _)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (a g _)) (to (_ g _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (a t a)) (to (a a a)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (a t a)) (to (a c a)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (a t a)) (to (a g a)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (a t a)) (to (a t c)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (a t a)) (to (a t g)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (a t a)) (to (a t t)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (a t a)) (to (a t _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (a t a)) (to (a _ a)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (a t a)) (to (c t a)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (a t a)) (to (g t a)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (a t a)) (to (t t a)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (a t a)) (to (_ t a)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (a t c)) (to (a a c)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (a t c)) (to (a c c)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (a t c)) (to (a g c)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (a t c)) (to (a t a)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (a t c)) (to (a t g)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (a t c)) (to (a t t)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (a t c)) (to (a t _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (a t c)) (to (a _ c)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (a t c)) (to (c t c)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (a t c)) (to (g t c)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (a t c)) (to (t t c)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (a t c)) (to (_ t c)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (a t g)) (to (a a g)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (a t g)) (to (a c g)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (a t g)) (to (a g g)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (a t g)) (to (a t a)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (a t g)) (to (a t c)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (a t g)) (to (a t t)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (a t g)) (to (a t _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (a t g)) (to (a _ g)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (a t g)) (to (c t g)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (a t g)) (to (g t g)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (a t g)) (to (t t g)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (a t g)) (to (_ t g)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (a t t)) (to (a a t)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (a t t)) (to (a c t)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (a t t)) (to (a g t)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (a t t)) (to (a t a)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (a t t)) (to (a t c)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (a t t)) (to (a t g)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (a t t)) (to (a t _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (a t t)) (to (a _ t)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (a t t)) (to (c t t)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (a t t)) (to (g t t)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (a t t)) (to (t t t)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (a t t)) (to (_ t t)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (a t _)) (to (a a _)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (a t _)) (to (a c _)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (a t _)) (to (a g _)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (a t _)) (to (a t a)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (a t _)) (to (a t c)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (a t _)) (to (a t g)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (a t _)) (to (a t t)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (a t _)) (to (a _ _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (a t _)) (to (c t _)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (a t _)) (to (g t _)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (a t _)) (to (t t _)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (a t _)) (to (_ t _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (a _ a)) (to (a a a)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (a _ a)) (to (a c a)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (a _ a)) (to (a g a)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (a _ a)) (to (a t a)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (a _ a)) (to (a _ c)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (a _ a)) (to (a _ g)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (a _ a)) (to (a _ t)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (a _ a)) (to (a _ _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (a _ a)) (to (c _ a)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (a _ a)) (to (g _ a)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (a _ a)) (to (t _ a)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (a _ a)) (to (_ _ a)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (a _ c)) (to (a a c)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (a _ c)) (to (a c c)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (a _ c)) (to (a g c)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (a _ c)) (to (a t c)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (a _ c)) (to (a _ a)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (a _ c)) (to (a _ g)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (a _ c)) (to (a _ t)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (a _ c)) (to (a _ _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (a _ c)) (to (c _ c)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (a _ c)) (to (g _ c)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (a _ c)) (to (t _ c)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (a _ c)) (to (_ _ c)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (a _ g)) (to (a a g)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (a _ g)) (to (a c g)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (a _ g)) (to (a g g)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (a _ g)) (to (a t g)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (a _ g)) (to (a _ a)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (a _ g)) (to (a _ c)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (a _ g)) (to (a _ t)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (a _ g)) (to (a _ _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (a _ g)) (to (c _ g)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (a _ g)) (to (g _ g)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (a _ g)) (to (t _ g)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (a _ g)) (to (_ _ g)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (a _ t)) (to (a a t)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (a _ t)) (to (a c t)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (a _ t)) (to (a g t)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (a _ t)) (to (a t t)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (a _ t)) (to (a _ a)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (a _ t)) (to (a _ c)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (a _ t)) (to (a _ g)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (a _ t)) (to (a _ _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (a _ t)) (to (c _ t)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (a _ t)) (to (g _ t)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (a _ t)) (to (t _ t)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (a _ t)) (to (_ _ t)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (a _ _)) (to (a a _)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (a _ _)) (to (a c _)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (a _ _)) (to (a g _)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (a _ _)) (to (a t _)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (a _ _)) (to (a _ a)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (a _ _)) (to (a _ c)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (a _ _)) (to (a _ g)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (a _ _)) (to (a _ t)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (a _ _)) (to (c _ _)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (a _ _)) (to (g _ _)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (a _ _)) (to (t _ _)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (a _ _)) (to (_ _ _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (c a a)) (to (a a a)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (c a a)) (to (c a c)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (c a a)) (to (c a g)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (c a a)) (to (c a t)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (c a a)) (to (c a _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (c a a)) (to (c c a)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (c a a)) (to (c g a)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (c a a)) (to (c t a)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (c a a)) (to (c _ a)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (c a a)) (to (g a a)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (c a a)) (to (t a a)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (c a a)) (to (_ a a)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (c a c)) (to (a a c)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (c a c)) (to (c a a)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (c a c)) (to (c a g)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (c a c)) (to (c a t)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (c a c)) (to (c a _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (c a c)) (to (c c c)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (c a c)) (to (c g c)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (c a c)) (to (c t c)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (c a c)) (to (c _ c)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (c a c)) (to (g a c)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (c a c)) (to (t a c)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (c a c)) (to (_ a c)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (c a g)) (to (a a g)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (c a g)) (to (c a a)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (c a g)) (to (c a c)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (c a g)) (to (c a t)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (c a g)) (to (c a _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (c a g)) (to (c c g)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (c a g)) (to (c g g)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (c a g)) (to (c t g)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (c a g)) (to (c _ g)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (c a g)) (to (g a g)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (c a g)) (to (t a g)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (c a g)) (to (_ a g)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (c a t)) (to (a a t)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (c a t)) (to (c a a)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (c a t)) (to (c a c)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (c a t)) (to (c a g)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (c a t)) (to (c a _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (c a t)) (to (c c t)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (c a t)) (to (c g t)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (c a t)) (to (c t t)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (c a t)) (to (c _ t)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (c a t)) (to (g a t)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (c a t)) (to (t a t)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (c a t)) (to (_ a t)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (c a _)) (to (a a _)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (c a _)) (to (c a a)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (c a _)) (to (c a c)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (c a _)) (to (c a g)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (c a _)) (to (c a t)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (c a _)) (to (c c _)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (c a _)) (to (c g _)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (c a _)) (to (c t _)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (c a _)) (to (c _ _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (c a _)) (to (g a _)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (c a _)) (to (t a _)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (c a _)) (to (_ a _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (c c a)) (to (a c a)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (c c a)) (to (c a a)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (c c a)) (to (c c c)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (c c a)) (to (c c g)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (c c a)) (to (c c t)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (c c a)) (to (c c _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (c c a)) (to (c g a)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (c c a)) (to (c t a)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (c c a)) (to (c _ a)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (c c a)) (to (g c a)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (c c a)) (to (t c a)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (c c a)) (to (_ c a)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (c c c)) (to (a c c)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (c c c)) (to (c a c)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (c c c)) (to (c c a)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (c c c)) (to (c c g)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (c c c)) (to (c c t)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (c c c)) (to (c c _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (c c c)) (to (c g c)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (c c c)) (to (c t c)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (c c c)) (to (c _ c)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (c c c)) (to (g c c)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (c c c)) (to (t c c)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (c c c)) (to (_ c c)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (c c g)) (to (a c g)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (c c g)) (to (c a g)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (c c g)) (to (c c a)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (c c g)) (to (c c c)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (c c g)) (to (c c t)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (c c g)) (to (c c _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (c c g)) (to (c g g)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (c c g)) (to (c t g)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (c c g)) (to (c _ g)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (c c g)) (to (g c g)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (c c g)) (to (t c g)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (c c g)) (to (_ c g)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (c c t)) (to (a c t)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (c c t)) (to (c a t)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (c c t)) (to (c c a)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (c c t)) (to (c c c)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (c c t)) (to (c c g)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (c c t)) (to (c c _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (c c t)) (to (c g t)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (c c t)) (to (c t t)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (c c t)) (to (c _ t)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (c c t)) (to (g c t)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (c c t)) (to (t c t)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (c c t)) (to (_ c t)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (c c _)) (to (a c _)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (c c _)) (to (c a _)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (c c _)) (to (c c a)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (c c _)) (to (c c c)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (c c _)) (to (c c g)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (c c _)) (to (c c t)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (c c _)) (to (c g _)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (c c _)) (to (c t _)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (c c _)) (to (c _ _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (c c _)) (to (g c _)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (c c _)) (to (t c _)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (c c _)) (to (_ c _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (c g a)) (to (a g a)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (c g a)) (to (c a a)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (c g a)) (to (c c a)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (c g a)) (to (c g c)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (c g a)) (to (c g g)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (c g a)) (to (c g t)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (c g a)) (to (c g _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (c g a)) (to (c t a)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (c g a)) (to (c _ a)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (c g a)) (to (g g a)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (c g a)) (to (t g a)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (c g a)) (to (_ g a)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (c g c)) (to (a g c)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (c g c)) (to (c a c)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (c g c)) (to (c c c)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (c g c)) (to (c g a)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (c g c)) (to (c g g)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (c g c)) (to (c g t)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (c g c)) (to (c g _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (c g c)) (to (c t c)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (c g c)) (to (c _ c)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (c g c)) (to (g g c)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (c g c)) (to (t g c)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (c g c)) (to (_ g c)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (c g g)) (to (a g g)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (c g g)) (to (c a g)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (c g g)) (to (c c g)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (c g g)) (to (c g a)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (c g g)) (to (c g c)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (c g g)) (to (c g t)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (c g g)) (to (c g _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (c g g)) (to (c t g)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (c g g)) (to (c _ g)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (c g g)) (to (g g g)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (c g g)) (to (t g g)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (c g g)) (to (_ g g)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (c g t)) (to (a g t)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (c g t)) (to (c a t)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (c g t)) (to (c c t)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (c g t)) (to (c g a)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (c g t)) (to (c g c)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (c g t)) (to (c g g)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (c g t)) (to (c g _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (c g t)) (to (c t t)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (c g t)) (to (c _ t)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (c g t)) (to (g g t)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (c g t)) (to (t g t)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (c g t)) (to (_ g t)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (c g _)) (to (a g _)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (c g _)) (to (c a _)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (c g _)) (to (c c _)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (c g _)) (to (c g a)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (c g _)) (to (c g c)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (c g _)) (to (c g g)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (c g _)) (to (c g t)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (c g _)) (to (c t _)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (c g _)) (to (c _ _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (c g _)) (to (g g _)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (c g _)) (to (t g _)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (c g _)) (to (_ g _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (c t a)) (to (a t a)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (c t a)) (to (c a a)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (c t a)) (to (c c a)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (c t a)) (to (c g a)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (c t a)) (to (c t c)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (c t a)) (to (c t g)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (c t a)) (to (c t t)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (c t a)) (to (c t _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (c t a)) (to (c _ a)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (c t a)) (to (g t a)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (c t a)) (to (t t a)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (c t a)) (to (_ t a)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (c t c)) (to (a t c)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (c t c)) (to (c a c)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (c t c)) (to (c c c)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (c t c)) (to (c g c)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (c t c)) (to (c t a)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (c t c)) (to (c t g)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (c t c)) (to (c t t)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (c t c)) (to (c t _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (c t c)) (to (c _ c)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (c t c)) (to (g t c)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (c t c)) (to (t t c)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (c t c)) (to (_ t c)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (c t g)) (to (a t g)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (c t g)) (to (c a g)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (c t g)) (to (c c g)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (c t g)) (to (c g g)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (c t g)) (to (c t a)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (c t g)) (to (c t c)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (c t g)) (to (c t t)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (c t g)) (to (c t _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (c t g)) (to (c _ g)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (c t g)) (to (g t g)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (c t g)) (to (t t g)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (c t g)) (to (_ t g)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (c t t)) (to (a t t)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (c t t)) (to (c a t)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (c t t)) (to (c c t)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (c t t)) (to (c g t)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (c t t)) (to (c t a)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (c t t)) (to (c t c)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (c t t)) (to (c t g)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (c t t)) (to (c t _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (c t t)) (to (c _ t)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (c t t)) (to (g t t)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (c t t)) (to (t t t)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (c t t)) (to (_ t t)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (c t _)) (to (a t _)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (c t _)) (to (c a _)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (c t _)) (to (c c _)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (c t _)) (to (c g _)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (c t _)) (to (c t a)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (c t _)) (to (c t c)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (c t _)) (to (c t g)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (c t _)) (to (c t t)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (c t _)) (to (c _ _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (c t _)) (to (g t _)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (c t _)) (to (t t _)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (c t _)) (to (_ t _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (c _ a)) (to (a _ a)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (c _ a)) (to (c a a)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (c _ a)) (to (c c a)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (c _ a)) (to (c g a)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (c _ a)) (to (c t a)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (c _ a)) (to (c _ c)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (c _ a)) (to (c _ g)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (c _ a)) (to (c _ t)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (c _ a)) (to (c _ _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (c _ a)) (to (g _ a)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (c _ a)) (to (t _ a)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (c _ a)) (to (_ _ a)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (c _ c)) (to (a _ c)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (c _ c)) (to (c a c)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (c _ c)) (to (c c c)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (c _ c)) (to (c g c)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (c _ c)) (to (c t c)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (c _ c)) (to (c _ a)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (c _ c)) (to (c _ g)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (c _ c)) (to (c _ t)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (c _ c)) (to (c _ _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (c _ c)) (to (g _ c)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (c _ c)) (to (t _ c)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (c _ c)) (to (_ _ c)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (c _ g)) (to (a _ g)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (c _ g)) (to (c a g)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (c _ g)) (to (c c g)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (c _ g)) (to (c g g)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (c _ g)) (to (c t g)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (c _ g)) (to (c _ a)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (c _ g)) (to (c _ c)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (c _ g)) (to (c _ t)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (c _ g)) (to (c _ _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (c _ g)) (to (g _ g)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (c _ g)) (to (t _ g)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (c _ g)) (to (_ _ g)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (c _ t)) (to (a _ t)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (c _ t)) (to (c a t)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (c _ t)) (to (c c t)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (c _ t)) (to (c g t)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (c _ t)) (to (c t t)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (c _ t)) (to (c _ a)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (c _ t)) (to (c _ c)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (c _ t)) (to (c _ g)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (c _ t)) (to (c _ _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (c _ t)) (to (g _ t)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (c _ t)) (to (t _ t)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (c _ t)) (to (_ _ t)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (c _ _)) (to (a _ _)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (c _ _)) (to (c a _)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (c _ _)) (to (c c _)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (c _ _)) (to (c g _)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (c _ _)) (to (c t _)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (c _ _)) (to (c _ a)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (c _ _)) (to (c _ c)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (c _ _)) (to (c _ g)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (c _ _)) (to (c _ t)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (c _ _)) (to (g _ _)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (c _ _)) (to (t _ _)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (c _ _)) (to (_ _ _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (g a a)) (to (a a a)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (g a a)) (to (c a a)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (g a a)) (to (g a c)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (g a a)) (to (g a g)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (g a a)) (to (g a t)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (g a a)) (to (g a _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (g a a)) (to (g c a)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (g a a)) (to (g g a)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (g a a)) (to (g t a)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (g a a)) (to (g _ a)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (g a a)) (to (t a a)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (g a a)) (to (_ a a)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (g a c)) (to (a a c)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (g a c)) (to (c a c)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (g a c)) (to (g a a)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (g a c)) (to (g a g)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (g a c)) (to (g a t)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (g a c)) (to (g a _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (g a c)) (to (g c c)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (g a c)) (to (g g c)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (g a c)) (to (g t c)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (g a c)) (to (g _ c)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (g a c)) (to (t a c)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (g a c)) (to (_ a c)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (g a g)) (to (a a g)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (g a g)) (to (c a g)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (g a g)) (to (g a a)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (g a g)) (to (g a c)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (g a g)) (to (g a t)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (g a g)) (to (g a _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (g a g)) (to (g c g)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (g a g)) (to (g g g)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (g a g)) (to (g t g)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (g a g)) (to (g _ g)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (g a g)) (to (t a g)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (g a g)) (to (_ a g)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (g a t)) (to (a a t)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (g a t)) (to (c a t)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (g a t)) (to (g a a)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (g a t)) (to (g a c)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (g a t)) (to (g a g)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (g a t)) (to (g a _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (g a t)) (to (g c t)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (g a t)) (to (g g t)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (g a t)) (to (g t t)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (g a t)) (to (g _ t)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (g a t)) (to (t a t)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (g a t)) (to (_ a t)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (g a _)) (to (a a _)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (g a _)) (to (c a _)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (g a _)) (to (g a a)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (g a _)) (to (g a c)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (g a _)) (to (g a g)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (g a _)) (to (g a t)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (g a _)) (to (g c _)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (g a _)) (to (g g _)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (g a _)) (to (g t _)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (g a _)) (to (g _ _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (g a _)) (to (t a _)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (g a _)) (to (_ a _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (g c a)) (to (a c a)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (g c a)) (to (c c a)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (g c a)) (to (g a a)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (g c a)) (to (g c c)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (g c a)) (to (g c g)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (g c a)) (to (g c t)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (g c a)) (to (g c _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (g c a)) (to (g g a)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (g c a)) (to (g t a)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (g c a)) (to (g _ a)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (g c a)) (to (t c a)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (g c a)) (to (_ c a)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (g c c)) (to (a c c)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (g c c)) (to (c c c)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (g c c)) (to (g a c)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (g c c)) (to (g c a)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (g c c)) (to (g c g)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (g c c)) (to (g c t)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (g c c)) (to (g c _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (g c c)) (to (g g c)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (g c c)) (to (g t c)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (g c c)) (to (g _ c)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (g c c)) (to (t c c)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (g c c)) (to (_ c c)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (g c g)) (to (a c g)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (g c g)) (to (c c g)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (g c g)) (to (g a g)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (g c g)) (to (g c a)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (g c g)) (to (g c c)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (g c g)) (to (g c t)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (g c g)) (to (g c _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (g c g)) (to (g g g)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (g c g)) (to (g t g)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (g c g)) (to (g _ g)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (g c g)) (to (t c g)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (g c g)) (to (_ c g)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (g c t)) (to (a c t)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (g c t)) (to (c c t)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (g c t)) (to (g a t)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (g c t)) (to (g c a)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (g c t)) (to (g c c)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (g c t)) (to (g c g)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (g c t)) (to (g c _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (g c t)) (to (g g t)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (g c t)) (to (g t t)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (g c t)) (to (g _ t)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (g c t)) (to (t c t)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (g c t)) (to (_ c t)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (g c _)) (to (a c _)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (g c _)) (to (c c _)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (g c _)) (to (g a _)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (g c _)) (to (g c a)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (g c _)) (to (g c c)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (g c _)) (to (g c g)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (g c _)) (to (g c t)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (g c _)) (to (g g _)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (g c _)) (to (g t _)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (g c _)) (to (g _ _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (g c _)) (to (t c _)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (g c _)) (to (_ c _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (g g a)) (to (a g a)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (g g a)) (to (c g a)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (g g a)) (to (g a a)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (g g a)) (to (g c a)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (g g a)) (to (g g c)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (g g a)) (to (g g g)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (g g a)) (to (g g t)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (g g a)) (to (g g _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (g g a)) (to (g t a)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (g g a)) (to (g _ a)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (g g a)) (to (t g a)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (g g a)) (to (_ g a)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (g g c)) (to (a g c)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (g g c)) (to (c g c)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (g g c)) (to (g a c)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (g g c)) (to (g c c)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (g g c)) (to (g g a)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (g g c)) (to (g g g)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (g g c)) (to (g g t)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (g g c)) (to (g g _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (g g c)) (to (g t c)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (g g c)) (to (g _ c)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (g g c)) (to (t g c)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (g g c)) (to (_ g c)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (g g g)) (to (a g g)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (g g g)) (to (c g g)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (g g g)) (to (g a g)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (g g g)) (to (g c g)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (g g g)) (to (g g a)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (g g g)) (to (g g c)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (g g g)) (to (g g t)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (g g g)) (to (g g _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (g g g)) (to (g t g)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (g g g)) (to (g _ g)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (g g g)) (to (t g g)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (g g g)) (to (_ g g)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (g g t)) (to (a g t)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (g g t)) (to (c g t)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (g g t)) (to (g a t)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (g g t)) (to (g c t)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (g g t)) (to (g g a)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (g g t)) (to (g g c)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (g g t)) (to (g g g)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (g g t)) (to (g g _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (g g t)) (to (g t t)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (g g t)) (to (g _ t)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (g g t)) (to (t g t)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (g g t)) (to (_ g t)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (g g _)) (to (a g _)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (g g _)) (to (c g _)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (g g _)) (to (g a _)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (g g _)) (to (g c _)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (g g _)) (to (g g a)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (g g _)) (to (g g c)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (g g _)) (to (g g g)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (g g _)) (to (g g t)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (g g _)) (to (g t _)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (g g _)) (to (g _ _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (g g _)) (to (t g _)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (g g _)) (to (_ g _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (g t a)) (to (a t a)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (g t a)) (to (c t a)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (g t a)) (to (g a a)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (g t a)) (to (g c a)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (g t a)) (to (g g a)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (g t a)) (to (g t c)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (g t a)) (to (g t g)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (g t a)) (to (g t t)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (g t a)) (to (g t _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (g t a)) (to (g _ a)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (g t a)) (to (t t a)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (g t a)) (to (_ t a)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (g t c)) (to (a t c)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (g t c)) (to (c t c)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (g t c)) (to (g a c)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (g t c)) (to (g c c)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (g t c)) (to (g g c)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (g t c)) (to (g t a)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (g t c)) (to (g t g)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (g t c)) (to (g t t)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (g t c)) (to (g t _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (g t c)) (to (g _ c)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (g t c)) (to (t t c)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (g t c)) (to (_ t c)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (g t g)) (to (a t g)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (g t g)) (to (c t g)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (g t g)) (to (g a g)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (g t g)) (to (g c g)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (g t g)) (to (g g g)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (g t g)) (to (g t a)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (g t g)) (to (g t c)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (g t g)) (to (g t t)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (g t g)) (to (g t _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (g t g)) (to (g _ g)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (g t g)) (to (t t g)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (g t g)) (to (_ t g)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (g t t)) (to (a t t)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (g t t)) (to (c t t)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (g t t)) (to (g a t)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (g t t)) (to (g c t)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (g t t)) (to (g g t)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (g t t)) (to (g t a)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (g t t)) (to (g t c)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (g t t)) (to (g t g)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (g t t)) (to (g t _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (g t t)) (to (g _ t)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (g t t)) (to (t t t)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (g t t)) (to (_ t t)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (g t _)) (to (a t _)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (g t _)) (to (c t _)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (g t _)) (to (g a _)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (g t _)) (to (g c _)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (g t _)) (to (g g _)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (g t _)) (to (g t a)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (g t _)) (to (g t c)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (g t _)) (to (g t g)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (g t _)) (to (g t t)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (g t _)) (to (g _ _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (g t _)) (to (t t _)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (g t _)) (to (_ t _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (g _ a)) (to (a _ a)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (g _ a)) (to (c _ a)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (g _ a)) (to (g a a)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (g _ a)) (to (g c a)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (g _ a)) (to (g g a)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (g _ a)) (to (g t a)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (g _ a)) (to (g _ c)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (g _ a)) (to (g _ g)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (g _ a)) (to (g _ t)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (g _ a)) (to (g _ _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (g _ a)) (to (t _ a)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (g _ a)) (to (_ _ a)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (g _ c)) (to (a _ c)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (g _ c)) (to (c _ c)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (g _ c)) (to (g a c)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (g _ c)) (to (g c c)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (g _ c)) (to (g g c)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (g _ c)) (to (g t c)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (g _ c)) (to (g _ a)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (g _ c)) (to (g _ g)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (g _ c)) (to (g _ t)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (g _ c)) (to (g _ _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (g _ c)) (to (t _ c)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (g _ c)) (to (_ _ c)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (g _ g)) (to (a _ g)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (g _ g)) (to (c _ g)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (g _ g)) (to (g a g)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (g _ g)) (to (g c g)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (g _ g)) (to (g g g)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (g _ g)) (to (g t g)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (g _ g)) (to (g _ a)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (g _ g)) (to (g _ c)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (g _ g)) (to (g _ t)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (g _ g)) (to (g _ _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (g _ g)) (to (t _ g)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (g _ g)) (to (_ _ g)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (g _ t)) (to (a _ t)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (g _ t)) (to (c _ t)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (g _ t)) (to (g a t)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (g _ t)) (to (g c t)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (g _ t)) (to (g g t)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (g _ t)) (to (g t t)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (g _ t)) (to (g _ a)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (g _ t)) (to (g _ c)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (g _ t)) (to (g _ g)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (g _ t)) (to (g _ _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (g _ t)) (to (t _ t)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (g _ t)) (to (_ _ t)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (g _ _)) (to (a _ _)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (g _ _)) (to (c _ _)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (g _ _)) (to (g a _)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (g _ _)) (to (g c _)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (g _ _)) (to (g g _)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (g _ _)) (to (g t _)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (g _ _)) (to (g _ a)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (g _ _)) (to (g _ c)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (g _ _)) (to (g _ g)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (g _ _)) (to (g _ t)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (g _ _)) (to (t _ _)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (g _ _)) (to (_ _ _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (t a a)) (to (a a a)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (t a a)) (to (c a a)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (t a a)) (to (g a a)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (t a a)) (to (t a c)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (t a a)) (to (t a g)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (t a a)) (to (t a t)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (t a a)) (to (t a _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (t a a)) (to (t c a)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (t a a)) (to (t g a)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (t a a)) (to (t t a)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (t a a)) (to (t _ a)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (t a a)) (to (_ a a)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (t a c)) (to (a a c)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (t a c)) (to (c a c)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (t a c)) (to (g a c)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (t a c)) (to (t a a)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (t a c)) (to (t a g)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (t a c)) (to (t a t)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (t a c)) (to (t a _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (t a c)) (to (t c c)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (t a c)) (to (t g c)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (t a c)) (to (t t c)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (t a c)) (to (t _ c)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (t a c)) (to (_ a c)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (t a g)) (to (a a g)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (t a g)) (to (c a g)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (t a g)) (to (g a g)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (t a g)) (to (t a a)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (t a g)) (to (t a c)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (t a g)) (to (t a t)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (t a g)) (to (t a _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (t a g)) (to (t c g)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (t a g)) (to (t g g)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (t a g)) (to (t t g)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (t a g)) (to (t _ g)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (t a g)) (to (_ a g)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (t a t)) (to (a a t)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (t a t)) (to (c a t)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (t a t)) (to (g a t)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (t a t)) (to (t a a)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (t a t)) (to (t a c)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (t a t)) (to (t a g)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (t a t)) (to (t a _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (t a t)) (to (t c t)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (t a t)) (to (t g t)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (t a t)) (to (t t t)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (t a t)) (to (t _ t)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (t a t)) (to (_ a t)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (t a _)) (to (a a _)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (t a _)) (to (c a _)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (t a _)) (to (g a _)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (t a _)) (to (t a a)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (t a _)) (to (t a c)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (t a _)) (to (t a g)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (t a _)) (to (t a t)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (t a _)) (to (t c _)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (t a _)) (to (t g _)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (t a _)) (to (t t _)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (t a _)) (to (t _ _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (t a _)) (to (_ a _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (t c a)) (to (a c a)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (t c a)) (to (c c a)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (t c a)) (to (g c a)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (t c a)) (to (t a a)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (t c a)) (to (t c c)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (t c a)) (to (t c g)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (t c a)) (to (t c t)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (t c a)) (to (t c _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (t c a)) (to (t g a)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (t c a)) (to (t t a)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (t c a)) (to (t _ a)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (t c a)) (to (_ c a)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (t c c)) (to (a c c)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (t c c)) (to (c c c)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (t c c)) (to (g c c)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (t c c)) (to (t a c)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (t c c)) (to (t c a)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (t c c)) (to (t c g)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (t c c)) (to (t c t)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (t c c)) (to (t c _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (t c c)) (to (t g c)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (t c c)) (to (t t c)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (t c c)) (to (t _ c)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (t c c)) (to (_ c c)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (t c g)) (to (a c g)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (t c g)) (to (c c g)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (t c g)) (to (g c g)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (t c g)) (to (t a g)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (t c g)) (to (t c a)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (t c g)) (to (t c c)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (t c g)) (to (t c t)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (t c g)) (to (t c _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (t c g)) (to (t g g)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (t c g)) (to (t t g)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (t c g)) (to (t _ g)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (t c g)) (to (_ c g)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (t c t)) (to (a c t)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (t c t)) (to (c c t)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (t c t)) (to (g c t)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (t c t)) (to (t a t)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (t c t)) (to (t c a)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (t c t)) (to (t c c)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (t c t)) (to (t c g)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (t c t)) (to (t c _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (t c t)) (to (t g t)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (t c t)) (to (t t t)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (t c t)) (to (t _ t)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (t c t)) (to (_ c t)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (t c _)) (to (a c _)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (t c _)) (to (c c _)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (t c _)) (to (g c _)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (t c _)) (to (t a _)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (t c _)) (to (t c a)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (t c _)) (to (t c c)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (t c _)) (to (t c g)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (t c _)) (to (t c t)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (t c _)) (to (t g _)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (t c _)) (to (t t _)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (t c _)) (to (t _ _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (t c _)) (to (_ c _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (t g a)) (to (a g a)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (t g a)) (to (c g a)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (t g a)) (to (g g a)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (t g a)) (to (t a a)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (t g a)) (to (t c a)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (t g a)) (to (t g c)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (t g a)) (to (t g g)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (t g a)) (to (t g t)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (t g a)) (to (t g _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (t g a)) (to (t t a)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (t g a)) (to (t _ a)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (t g a)) (to (_ g a)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (t g c)) (to (a g c)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (t g c)) (to (c g c)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (t g c)) (to (g g c)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (t g c)) (to (t a c)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (t g c)) (to (t c c)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (t g c)) (to (t g a)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (t g c)) (to (t g g)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (t g c)) (to (t g t)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (t g c)) (to (t g _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (t g c)) (to (t t c)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (t g c)) (to (t _ c)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (t g c)) (to (_ g c)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (t g g)) (to (a g g)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (t g g)) (to (c g g)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (t g g)) (to (g g g)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (t g g)) (to (t a g)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (t g g)) (to (t c g)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (t g g)) (to (t g a)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (t g g)) (to (t g c)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (t g g)) (to (t g t)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (t g g)) (to (t g _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (t g g)) (to (t t g)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (t g g)) (to (t _ g)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (t g g)) (to (_ g g)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (t g t)) (to (a g t)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (t g t)) (to (c g t)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (t g t)) (to (g g t)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (t g t)) (to (t a t)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (t g t)) (to (t c t)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (t g t)) (to (t g a)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (t g t)) (to (t g c)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (t g t)) (to (t g g)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (t g t)) (to (t g _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (t g t)) (to (t t t)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (t g t)) (to (t _ t)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (t g t)) (to (_ g t)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (t g _)) (to (a g _)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (t g _)) (to (c g _)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (t g _)) (to (g g _)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (t g _)) (to (t a _)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (t g _)) (to (t c _)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (t g _)) (to (t g a)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (t g _)) (to (t g c)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (t g _)) (to (t g g)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (t g _)) (to (t g t)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (t g _)) (to (t t _)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (t g _)) (to (t _ _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (t g _)) (to (_ g _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (t t a)) (to (a t a)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (t t a)) (to (c t a)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (t t a)) (to (g t a)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (t t a)) (to (t a a)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (t t a)) (to (t c a)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (t t a)) (to (t g a)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (t t a)) (to (t t c)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (t t a)) (to (t t g)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (t t a)) (to (t t t)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (t t a)) (to (t t _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (t t a)) (to (t _ a)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (t t a)) (to (_ t a)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (t t c)) (to (a t c)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (t t c)) (to (c t c)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (t t c)) (to (g t c)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (t t c)) (to (t a c)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (t t c)) (to (t c c)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (t t c)) (to (t g c)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (t t c)) (to (t t a)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (t t c)) (to (t t g)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (t t c)) (to (t t t)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (t t c)) (to (t t _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (t t c)) (to (t _ c)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (t t c)) (to (_ t c)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (t t g)) (to (a t g)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (t t g)) (to (c t g)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (t t g)) (to (g t g)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (t t g)) (to (t a g)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (t t g)) (to (t c g)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (t t g)) (to (t g g)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (t t g)) (to (t t a)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (t t g)) (to (t t c)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (t t g)) (to (t t t)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (t t g)) (to (t t _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (t t g)) (to (t _ g)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (t t g)) (to (_ t g)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (t t t)) (to (a t t)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (t t t)) (to (c t t)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (t t t)) (to (g t t)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (t t t)) (to (t a t)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (t t t)) (to (t c t)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (t t t)) (to (t g t)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (t t t)) (to (t t a)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (t t t)) (to (t t c)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (t t t)) (to (t t g)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (t t t)) (to (t t _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (t t t)) (to (t _ t)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (t t t)) (to (_ t t)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (t t _)) (to (a t _)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (t t _)) (to (c t _)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (t t _)) (to (g t _)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (t t _)) (to (t a _)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (t t _)) (to (t c _)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (t t _)) (to (t g _)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (t t _)) (to (t t a)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (t t _)) (to (t t c)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (t t _)) (to (t t g)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (t t _)) (to (t t t)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (t t _)) (to (t _ _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (t t _)) (to (_ t _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (t _ a)) (to (a _ a)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (t _ a)) (to (c _ a)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (t _ a)) (to (g _ a)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (t _ a)) (to (t a a)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (t _ a)) (to (t c a)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (t _ a)) (to (t g a)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (t _ a)) (to (t t a)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (t _ a)) (to (t _ c)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (t _ a)) (to (t _ g)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (t _ a)) (to (t _ t)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (t _ a)) (to (t _ _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (t _ a)) (to (_ _ a)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (t _ c)) (to (a _ c)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (t _ c)) (to (c _ c)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (t _ c)) (to (g _ c)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (t _ c)) (to (t a c)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (t _ c)) (to (t c c)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (t _ c)) (to (t g c)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (t _ c)) (to (t t c)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (t _ c)) (to (t _ a)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (t _ c)) (to (t _ g)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (t _ c)) (to (t _ t)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (t _ c)) (to (t _ _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (t _ c)) (to (_ _ c)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (t _ g)) (to (a _ g)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (t _ g)) (to (c _ g)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (t _ g)) (to (g _ g)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (t _ g)) (to (t a g)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (t _ g)) (to (t c g)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (t _ g)) (to (t g g)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (t _ g)) (to (t t g)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (t _ g)) (to (t _ a)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (t _ g)) (to (t _ c)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (t _ g)) (to (t _ t)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (t _ g)) (to (t _ _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (t _ g)) (to (_ _ g)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (t _ t)) (to (a _ t)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (t _ t)) (to (c _ t)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (t _ t)) (to (g _ t)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (t _ t)) (to (t a t)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (t _ t)) (to (t c t)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (t _ t)) (to (t g t)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (t _ t)) (to (t t t)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (t _ t)) (to (t _ a)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (t _ t)) (to (t _ c)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (t _ t)) (to (t _ g)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (t _ t)) (to (t _ _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (t _ t)) (to (_ _ t)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (t _ _)) (to (a _ _)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (t _ _)) (to (c _ _)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (t _ _)) (to (g _ _)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (t _ _)) (to (t a _)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (t _ _)) (to (t c _)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (t _ _)) (to (t g _)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (t _ _)) (to (t t _)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (t _ _)) (to (t _ a)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (t _ _)) (to (t _ c)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (t _ _)) (to (t _ g)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (t _ _)) (to (t _ t)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (t _ _)) (to (_ _ _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (_ a a)) (to (a a a)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ a a)) (to (c a a)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ a a)) (to (g a a)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ a a)) (to (t a a)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ a a)) (to (_ a c)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (_ a a)) (to (_ a g)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (_ a a)) (to (_ a t)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (_ a a)) (to (_ a _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (_ a a)) (to (_ c a)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (_ a a)) (to (_ g a)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (_ a a)) (to (_ t a)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (_ a a)) (to (_ _ a)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (_ a c)) (to (a a c)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ a c)) (to (c a c)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ a c)) (to (g a c)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ a c)) (to (t a c)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ a c)) (to (_ a a)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (_ a c)) (to (_ a g)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (_ a c)) (to (_ a t)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (_ a c)) (to (_ a _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (_ a c)) (to (_ c c)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (_ a c)) (to (_ g c)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (_ a c)) (to (_ t c)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (_ a c)) (to (_ _ c)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (_ a g)) (to (a a g)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ a g)) (to (c a g)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ a g)) (to (g a g)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ a g)) (to (t a g)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ a g)) (to (_ a a)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (_ a g)) (to (_ a c)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (_ a g)) (to (_ a t)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (_ a g)) (to (_ a _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (_ a g)) (to (_ c g)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (_ a g)) (to (_ g g)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (_ a g)) (to (_ t g)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (_ a g)) (to (_ _ g)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (_ a t)) (to (a a t)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ a t)) (to (c a t)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ a t)) (to (g a t)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ a t)) (to (t a t)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ a t)) (to (_ a a)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (_ a t)) (to (_ a c)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (_ a t)) (to (_ a g)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (_ a t)) (to (_ a _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (_ a t)) (to (_ c t)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (_ a t)) (to (_ g t)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (_ a t)) (to (_ t t)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (_ a t)) (to (_ _ t)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (_ a _)) (to (a a _)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ a _)) (to (c a _)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ a _)) (to (g a _)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ a _)) (to (t a _)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ a _)) (to (_ a a)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ a _)) (to (_ a c)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ a _)) (to (_ a g)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ a _)) (to (_ a t)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ a _)) (to (_ c _)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (_ a _)) (to (_ g _)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (_ a _)) (to (_ t _)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (_ a _)) (to (_ _ _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (_ c a)) (to (a c a)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ c a)) (to (c c a)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ c a)) (to (g c a)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ c a)) (to (t c a)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ c a)) (to (_ a a)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (_ c a)) (to (_ c c)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (_ c a)) (to (_ c g)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (_ c a)) (to (_ c t)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (_ c a)) (to (_ c _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (_ c a)) (to (_ g a)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (_ c a)) (to (_ t a)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (_ c a)) (to (_ _ a)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (_ c c)) (to (a c c)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ c c)) (to (c c c)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ c c)) (to (g c c)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ c c)) (to (t c c)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ c c)) (to (_ a c)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (_ c c)) (to (_ c a)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (_ c c)) (to (_ c g)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (_ c c)) (to (_ c t)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (_ c c)) (to (_ c _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (_ c c)) (to (_ g c)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (_ c c)) (to (_ t c)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (_ c c)) (to (_ _ c)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (_ c g)) (to (a c g)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ c g)) (to (c c g)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ c g)) (to (g c g)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ c g)) (to (t c g)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ c g)) (to (_ a g)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (_ c g)) (to (_ c a)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (_ c g)) (to (_ c c)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (_ c g)) (to (_ c t)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (_ c g)) (to (_ c _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (_ c g)) (to (_ g g)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (_ c g)) (to (_ t g)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (_ c g)) (to (_ _ g)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (_ c t)) (to (a c t)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ c t)) (to (c c t)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ c t)) (to (g c t)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ c t)) (to (t c t)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ c t)) (to (_ a t)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (_ c t)) (to (_ c a)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (_ c t)) (to (_ c c)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (_ c t)) (to (_ c g)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (_ c t)) (to (_ c _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (_ c t)) (to (_ g t)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (_ c t)) (to (_ t t)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (_ c t)) (to (_ _ t)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (_ c _)) (to (a c _)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ c _)) (to (c c _)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ c _)) (to (g c _)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ c _)) (to (t c _)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ c _)) (to (_ a _)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (_ c _)) (to (_ c a)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ c _)) (to (_ c c)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ c _)) (to (_ c g)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ c _)) (to (_ c t)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ c _)) (to (_ g _)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (_ c _)) (to (_ t _)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (_ c _)) (to (_ _ _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (_ g a)) (to (a g a)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ g a)) (to (c g a)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ g a)) (to (g g a)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ g a)) (to (t g a)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ g a)) (to (_ a a)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (_ g a)) (to (_ c a)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (_ g a)) (to (_ g c)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (_ g a)) (to (_ g g)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (_ g a)) (to (_ g t)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (_ g a)) (to (_ g _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (_ g a)) (to (_ t a)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (_ g a)) (to (_ _ a)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (_ g c)) (to (a g c)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ g c)) (to (c g c)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ g c)) (to (g g c)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ g c)) (to (t g c)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ g c)) (to (_ a c)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (_ g c)) (to (_ c c)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (_ g c)) (to (_ g a)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (_ g c)) (to (_ g g)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (_ g c)) (to (_ g t)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (_ g c)) (to (_ g _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (_ g c)) (to (_ t c)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (_ g c)) (to (_ _ c)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (_ g g)) (to (a g g)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ g g)) (to (c g g)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ g g)) (to (g g g)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ g g)) (to (t g g)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ g g)) (to (_ a g)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (_ g g)) (to (_ c g)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (_ g g)) (to (_ g a)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (_ g g)) (to (_ g c)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (_ g g)) (to (_ g t)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (_ g g)) (to (_ g _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (_ g g)) (to (_ t g)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (_ g g)) (to (_ _ g)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (_ g t)) (to (a g t)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ g t)) (to (c g t)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ g t)) (to (g g t)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ g t)) (to (t g t)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ g t)) (to (_ a t)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (_ g t)) (to (_ c t)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (_ g t)) (to (_ g a)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (_ g t)) (to (_ g c)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (_ g t)) (to (_ g g)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (_ g t)) (to (_ g _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (_ g t)) (to (_ t t)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (_ g t)) (to (_ _ t)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (_ g _)) (to (a g _)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ g _)) (to (c g _)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ g _)) (to (g g _)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ g _)) (to (t g _)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ g _)) (to (_ a _)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (_ g _)) (to (_ c _)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (_ g _)) (to (_ g a)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ g _)) (to (_ g c)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ g _)) (to (_ g g)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ g _)) (to (_ g t)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ g _)) (to (_ t _)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (_ g _)) (to (_ _ _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (_ t a)) (to (a t a)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ t a)) (to (c t a)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ t a)) (to (g t a)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ t a)) (to (t t a)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ t a)) (to (_ a a)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (_ t a)) (to (_ c a)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (_ t a)) (to (_ g a)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (_ t a)) (to (_ t c)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (_ t a)) (to (_ t g)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (_ t a)) (to (_ t t)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (_ t a)) (to (_ t _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (_ t a)) (to (_ _ a)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (_ t c)) (to (a t c)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ t c)) (to (c t c)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ t c)) (to (g t c)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ t c)) (to (t t c)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ t c)) (to (_ a c)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (_ t c)) (to (_ c c)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (_ t c)) (to (_ g c)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (_ t c)) (to (_ t a)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (_ t c)) (to (_ t g)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (_ t c)) (to (_ t t)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (_ t c)) (to (_ t _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (_ t c)) (to (_ _ c)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (_ t g)) (to (a t g)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ t g)) (to (c t g)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ t g)) (to (g t g)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ t g)) (to (t t g)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ t g)) (to (_ a g)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (_ t g)) (to (_ c g)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (_ t g)) (to (_ g g)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (_ t g)) (to (_ t a)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (_ t g)) (to (_ t c)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (_ t g)) (to (_ t t)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (_ t g)) (to (_ t _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (_ t g)) (to (_ _ g)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (_ t t)) (to (a t t)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ t t)) (to (c t t)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ t t)) (to (g t t)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ t t)) (to (t t t)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ t t)) (to (_ a t)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (_ t t)) (to (_ c t)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (_ t t)) (to (_ g t)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (_ t t)) (to (_ t a)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (_ t t)) (to (_ t c)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (_ t t)) (to (_ t g)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (_ t t)) (to (_ t _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (_ t t)) (to (_ _ t)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (_ t _)) (to (a t _)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ t _)) (to (c t _)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ t _)) (to (g t _)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ t _)) (to (t t _)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ t _)) (to (_ a _)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (_ t _)) (to (_ c _)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (_ t _)) (to (_ g _)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (_ t _)) (to (_ t a)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ t _)) (to (_ t c)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ t _)) (to (_ t g)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ t _)) (to (_ t t)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ t _)) (to (_ _ _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (_ _ a)) (to (a _ a)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ _ a)) (to (c _ a)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ _ a)) (to (g _ a)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ _ a)) (to (t _ a)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ _ a)) (to (_ a a)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ _ a)) (to (_ c a)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ _ a)) (to (_ g a)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ _ a)) (to (_ t a)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ _ a)) (to (_ _ c)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (_ _ a)) (to (_ _ g)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (_ _ a)) (to (_ _ t)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (_ _ a)) (to (_ _ _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (_ _ c)) (to (a _ c)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ _ c)) (to (c _ c)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ _ c)) (to (g _ c)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ _ c)) (to (t _ c)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ _ c)) (to (_ a c)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ _ c)) (to (_ c c)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ _ c)) (to (_ g c)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ _ c)) (to (_ t c)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ _ c)) (to (_ _ a)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (_ _ c)) (to (_ _ g)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (_ _ c)) (to (_ _ t)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (_ _ c)) (to (_ _ _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (_ _ g)) (to (a _ g)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ _ g)) (to (c _ g)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ _ g)) (to (g _ g)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ _ g)) (to (t _ g)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ _ g)) (to (_ a g)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ _ g)) (to (_ c g)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ _ g)) (to (_ g g)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ _ g)) (to (_ t g)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ _ g)) (to (_ _ a)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (_ _ g)) (to (_ _ c)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (_ _ g)) (to (_ _ t)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (_ _ g)) (to (_ _ _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (_ _ t)) (to (a _ t)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ _ t)) (to (c _ t)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ _ t)) (to (g _ t)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ _ t)) (to (t _ t)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ _ t)) (to (_ a t)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ _ t)) (to (_ c t)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ _ t)) (to (_ g t)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ _ t)) (to (_ t t)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ _ t)) (to (_ _ a)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (_ _ t)) (to (_ _ c)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (_ _ t)) (to (_ _ g)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (_ _ t)) (to (_ _ _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (_ _ _)) (to (a _ _)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ _ _)) (to (c _ _)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ _ _)) (to (g _ _)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ _ _)) (to (t _ _)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ _ _)) (to (_ a _)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ _ _)) (to (_ c _)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ _ _)) (to (_ g _)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ _ _)) (to (_ t _)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ _ _)) (to (_ _ a)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ _ _)) (to (_ _ c)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ _ _)) (to (_ _ g)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ _ _)) (to (_ _ t)) (rate neutral_rate_scale * X_gap2a_gap2t))

 )  ;; end chain PG1 PG2 PG3


;;;;;;;;;;;
;; NCRNA ;;
;;;;;;;;;;;


 ;; Basepairs
 (chain
  (update-policy parametric)
  (terminal (LNUC RNUC))

  ;; initial probability distribution
  (initial (state (a a)) (prob ~initGap * LNUC_RNUC_aa))
  (initial (state (c a)) (prob ~initGap * LNUC_RNUC_ca))
  (initial (state (g a)) (prob ~initGap * LNUC_RNUC_ga))
  (initial (state (t a)) (prob ~initGap * LNUC_RNUC_ta))
  (initial (state (_ a)) (prob initGap * LNUC_RNUC_Xa))
  (initial (state (a c)) (prob ~initGap * LNUC_RNUC_ac))
  (initial (state (c c)) (prob ~initGap * LNUC_RNUC_cc))
  (initial (state (g c)) (prob ~initGap * LNUC_RNUC_gc))
  (initial (state (t c)) (prob ~initGap * LNUC_RNUC_tc))
  (initial (state (_ c)) (prob initGap * LNUC_RNUC_Xc))
  (initial (state (a g)) (prob ~initGap * LNUC_RNUC_ag))
  (initial (state (c g)) (prob ~initGap * LNUC_RNUC_cg))
  (initial (state (g g)) (prob ~initGap * LNUC_RNUC_gg))
  (initial (state (t g)) (prob ~initGap * LNUC_RNUC_tg))
  (initial (state (_ g)) (prob initGap * LNUC_RNUC_Xg))
  (initial (state (a t)) (prob ~initGap * LNUC_RNUC_at))
  (initial (state (c t)) (prob ~initGap * LNUC_RNUC_ct))
  (initial (state (g t)) (prob ~initGap * LNUC_RNUC_gt))
  (initial (state (t t)) (prob ~initGap * LNUC_RNUC_tt))
  (initial (state (_ t)) (prob initGap * LNUC_RNUC_Xt))
  (initial (state (a _)) (prob initGap * LNUC_RNUC_aX))
  (initial (state (c _)) (prob initGap * LNUC_RNUC_cX))
  (initial (state (g _)) (prob initGap * LNUC_RNUC_gX))
  (initial (state (t _)) (prob initGap * LNUC_RNUC_tX))
  (initial (state (_ _)) (prob initGap * LNUC_RNUC_XX))

  ;; mutation rates
  (mutate (from (a a)) (to (c a)) (rate ncrna_rate_scale * LNUC_RNUC_aa2ca))
  (mutate (from (a a)) (to (g a)) (rate ncrna_rate_scale * LNUC_RNUC_aa2ga))
  (mutate (from (a a)) (to (t a)) (rate ncrna_rate_scale * LNUC_RNUC_aa2ta))
  (mutate (from (a a)) (to (_ a)) (rate ncrna_rate_scale * LNUC_RNUC_aa2Xa))
  (mutate (from (a a)) (to (c c)) (rate ncrna_rate_scale * LNUC_RNUC_aa2cc))
  (mutate (from (a a)) (to (g c)) (rate ncrna_rate_scale * LNUC_RNUC_aa2gc))
  (mutate (from (a a)) (to (t c)) (rate ncrna_rate_scale * LNUC_RNUC_aa2tc))
  (mutate (from (a a)) (to (c g)) (rate ncrna_rate_scale * LNUC_RNUC_aa2cg))
  (mutate (from (a a)) (to (g g)) (rate ncrna_rate_scale * LNUC_RNUC_aa2gg))
  (mutate (from (a a)) (to (t g)) (rate ncrna_rate_scale * LNUC_RNUC_aa2tg))
  (mutate (from (a a)) (to (a t)) (rate ncrna_rate_scale * LNUC_RNUC_aa2at))
  (mutate (from (a a)) (to (c t)) (rate ncrna_rate_scale * LNUC_RNUC_aa2ct))
  (mutate (from (a a)) (to (g t)) (rate ncrna_rate_scale * LNUC_RNUC_aa2gt))
  (mutate (from (a a)) (to (t t)) (rate ncrna_rate_scale * LNUC_RNUC_aa2tt))
  (mutate (from (a a)) (to (a _)) (rate ncrna_rate_scale * LNUC_RNUC_aa2aX))
  (mutate (from (c a)) (to (a a)) (rate ncrna_rate_scale * LNUC_RNUC_ca2aa))
  (mutate (from (c a)) (to (g a)) (rate ncrna_rate_scale * LNUC_RNUC_ca2ga))
  (mutate (from (c a)) (to (t a)) (rate ncrna_rate_scale * LNUC_RNUC_ca2ta))
  (mutate (from (c a)) (to (a c)) (rate ncrna_rate_scale * LNUC_RNUC_ca2ac))
  (mutate (from (c a)) (to (c c)) (rate ncrna_rate_scale * LNUC_RNUC_ca2cc))
  (mutate (from (c a)) (to (g c)) (rate ncrna_rate_scale * LNUC_RNUC_ca2gc))
  (mutate (from (c a)) (to (t c)) (rate ncrna_rate_scale * LNUC_RNUC_ca2tc))
  (mutate (from (c a)) (to (_ c)) (rate ncrna_rate_scale * LNUC_RNUC_ca2Xc))
  (mutate (from (c a)) (to (a g)) (rate ncrna_rate_scale * LNUC_RNUC_ca2ag))
  (mutate (from (c a)) (to (c g)) (rate ncrna_rate_scale * LNUC_RNUC_ca2cg))
  (mutate (from (c a)) (to (g g)) (rate ncrna_rate_scale * LNUC_RNUC_ca2gg))
  (mutate (from (c a)) (to (t g)) (rate ncrna_rate_scale * LNUC_RNUC_ca2tg))
  (mutate (from (c a)) (to (a t)) (rate ncrna_rate_scale * LNUC_RNUC_ca2at))
  (mutate (from (c a)) (to (c t)) (rate ncrna_rate_scale * LNUC_RNUC_ca2ct))
  (mutate (from (c a)) (to (g t)) (rate ncrna_rate_scale * LNUC_RNUC_ca2gt))
  (mutate (from (c a)) (to (t t)) (rate ncrna_rate_scale * LNUC_RNUC_ca2tt))
  (mutate (from (c a)) (to (c _)) (rate ncrna_rate_scale * LNUC_RNUC_ca2cX))
  (mutate (from (g a)) (to (a a)) (rate ncrna_rate_scale * LNUC_RNUC_ga2aa))
  (mutate (from (g a)) (to (c a)) (rate ncrna_rate_scale * LNUC_RNUC_ga2ca))
  (mutate (from (g a)) (to (t a)) (rate ncrna_rate_scale * LNUC_RNUC_ga2ta))
  (mutate (from (g a)) (to (a c)) (rate ncrna_rate_scale * LNUC_RNUC_ga2ac))
  (mutate (from (g a)) (to (c c)) (rate ncrna_rate_scale * LNUC_RNUC_ga2cc))
  (mutate (from (g a)) (to (g c)) (rate ncrna_rate_scale * LNUC_RNUC_ga2gc))
  (mutate (from (g a)) (to (t c)) (rate ncrna_rate_scale * LNUC_RNUC_ga2tc))
  (mutate (from (g a)) (to (a g)) (rate ncrna_rate_scale * LNUC_RNUC_ga2ag))
  (mutate (from (g a)) (to (c g)) (rate ncrna_rate_scale * LNUC_RNUC_ga2cg))
  (mutate (from (g a)) (to (g g)) (rate ncrna_rate_scale * LNUC_RNUC_ga2gg))
  (mutate (from (g a)) (to (t g)) (rate ncrna_rate_scale * LNUC_RNUC_ga2tg))
  (mutate (from (g a)) (to (_ g)) (rate ncrna_rate_scale * LNUC_RNUC_ga2Xg))
  (mutate (from (g a)) (to (a t)) (rate ncrna_rate_scale * LNUC_RNUC_ga2at))
  (mutate (from (g a)) (to (c t)) (rate ncrna_rate_scale * LNUC_RNUC_ga2ct))
  (mutate (from (g a)) (to (g t)) (rate ncrna_rate_scale * LNUC_RNUC_ga2gt))
  (mutate (from (g a)) (to (t t)) (rate ncrna_rate_scale * LNUC_RNUC_ga2tt))
  (mutate (from (g a)) (to (g _)) (rate ncrna_rate_scale * LNUC_RNUC_ga2gX))
  (mutate (from (t a)) (to (a a)) (rate ncrna_rate_scale * LNUC_RNUC_ta2aa))
  (mutate (from (t a)) (to (c a)) (rate ncrna_rate_scale * LNUC_RNUC_ta2ca))
  (mutate (from (t a)) (to (g a)) (rate ncrna_rate_scale * LNUC_RNUC_ta2ga))
  (mutate (from (t a)) (to (a c)) (rate ncrna_rate_scale * LNUC_RNUC_ta2ac))
  (mutate (from (t a)) (to (c c)) (rate ncrna_rate_scale * LNUC_RNUC_ta2cc))
  (mutate (from (t a)) (to (g c)) (rate ncrna_rate_scale * LNUC_RNUC_ta2gc))
  (mutate (from (t a)) (to (t c)) (rate ncrna_rate_scale * LNUC_RNUC_ta2tc))
  (mutate (from (t a)) (to (a g)) (rate ncrna_rate_scale * LNUC_RNUC_ta2ag))
  (mutate (from (t a)) (to (c g)) (rate ncrna_rate_scale * LNUC_RNUC_ta2cg))
  (mutate (from (t a)) (to (g g)) (rate ncrna_rate_scale * LNUC_RNUC_ta2gg))
  (mutate (from (t a)) (to (t g)) (rate ncrna_rate_scale * LNUC_RNUC_ta2tg))
  (mutate (from (t a)) (to (a t)) (rate ncrna_rate_scale * LNUC_RNUC_ta2at))
  (mutate (from (t a)) (to (c t)) (rate ncrna_rate_scale * LNUC_RNUC_ta2ct))
  (mutate (from (t a)) (to (g t)) (rate ncrna_rate_scale * LNUC_RNUC_ta2gt))
  (mutate (from (t a)) (to (t t)) (rate ncrna_rate_scale * LNUC_RNUC_ta2tt))
  (mutate (from (t a)) (to (_ t)) (rate ncrna_rate_scale * LNUC_RNUC_ta2Xt))
  (mutate (from (t a)) (to (t _)) (rate ncrna_rate_scale * LNUC_RNUC_ta2tX))
  (mutate (from (_ a)) (to (a a)) (rate ncrna_rate_scale * LNUC_RNUC_Xa2aa))
  (mutate (from (_ a)) (to (c a)) (rate ncrna_rate_scale * LNUC_RNUC_Xa2ca))
  (mutate (from (_ a)) (to (g a)) (rate ncrna_rate_scale * LNUC_RNUC_Xa2ga))
  (mutate (from (_ a)) (to (t a)) (rate ncrna_rate_scale * LNUC_RNUC_Xa2ta))
  (mutate (from (_ a)) (to (_ c)) (rate ncrna_rate_scale * LNUC_RNUC_Xa2Xc))
  (mutate (from (_ a)) (to (_ g)) (rate ncrna_rate_scale * LNUC_RNUC_Xa2Xg))
  (mutate (from (_ a)) (to (_ t)) (rate ncrna_rate_scale * LNUC_RNUC_Xa2Xt))
  (mutate (from (_ a)) (to (_ _)) (rate ncrna_rate_scale * LNUC_RNUC_Xa2XX))
  (mutate (from (a c)) (to (a a)) (rate ncrna_rate_scale * LNUC_RNUC_ac2aa))
  (mutate (from (a c)) (to (c a)) (rate ncrna_rate_scale * LNUC_RNUC_ac2ca))
  (mutate (from (a c)) (to (g a)) (rate ncrna_rate_scale * LNUC_RNUC_ac2ga))
  (mutate (from (a c)) (to (t a)) (rate ncrna_rate_scale * LNUC_RNUC_ac2ta))
  (mutate (from (a c)) (to (_ a)) (rate ncrna_rate_scale * LNUC_RNUC_ac2Xa))
  (mutate (from (a c)) (to (c c)) (rate ncrna_rate_scale * LNUC_RNUC_ac2cc))
  (mutate (from (a c)) (to (g c)) (rate ncrna_rate_scale * LNUC_RNUC_ac2gc))
  (mutate (from (a c)) (to (t c)) (rate ncrna_rate_scale * LNUC_RNUC_ac2tc))
  (mutate (from (a c)) (to (a g)) (rate ncrna_rate_scale * LNUC_RNUC_ac2ag))
  (mutate (from (a c)) (to (c g)) (rate ncrna_rate_scale * LNUC_RNUC_ac2cg))
  (mutate (from (a c)) (to (g g)) (rate ncrna_rate_scale * LNUC_RNUC_ac2gg))
  (mutate (from (a c)) (to (t g)) (rate ncrna_rate_scale * LNUC_RNUC_ac2tg))
  (mutate (from (a c)) (to (a t)) (rate ncrna_rate_scale * LNUC_RNUC_ac2at))
  (mutate (from (a c)) (to (c t)) (rate ncrna_rate_scale * LNUC_RNUC_ac2ct))
  (mutate (from (a c)) (to (g t)) (rate ncrna_rate_scale * LNUC_RNUC_ac2gt))
  (mutate (from (a c)) (to (t t)) (rate ncrna_rate_scale * LNUC_RNUC_ac2tt))
  (mutate (from (a c)) (to (a _)) (rate ncrna_rate_scale * LNUC_RNUC_ac2aX))
  (mutate (from (c c)) (to (a a)) (rate ncrna_rate_scale * LNUC_RNUC_cc2aa))
  (mutate (from (c c)) (to (c a)) (rate ncrna_rate_scale * LNUC_RNUC_cc2ca))
  (mutate (from (c c)) (to (g a)) (rate ncrna_rate_scale * LNUC_RNUC_cc2ga))
  (mutate (from (c c)) (to (t a)) (rate ncrna_rate_scale * LNUC_RNUC_cc2ta))
  (mutate (from (c c)) (to (a c)) (rate ncrna_rate_scale * LNUC_RNUC_cc2ac))
  (mutate (from (c c)) (to (g c)) (rate ncrna_rate_scale * LNUC_RNUC_cc2gc))
  (mutate (from (c c)) (to (t c)) (rate ncrna_rate_scale * LNUC_RNUC_cc2tc))
  (mutate (from (c c)) (to (_ c)) (rate ncrna_rate_scale * LNUC_RNUC_cc2Xc))
  (mutate (from (c c)) (to (a g)) (rate ncrna_rate_scale * LNUC_RNUC_cc2ag))
  (mutate (from (c c)) (to (c g)) (rate ncrna_rate_scale * LNUC_RNUC_cc2cg))
  (mutate (from (c c)) (to (g g)) (rate ncrna_rate_scale * LNUC_RNUC_cc2gg))
  (mutate (from (c c)) (to (t g)) (rate ncrna_rate_scale * LNUC_RNUC_cc2tg))
  (mutate (from (c c)) (to (a t)) (rate ncrna_rate_scale * LNUC_RNUC_cc2at))
  (mutate (from (c c)) (to (c t)) (rate ncrna_rate_scale * LNUC_RNUC_cc2ct))
  (mutate (from (c c)) (to (g t)) (rate ncrna_rate_scale * LNUC_RNUC_cc2gt))
  (mutate (from (c c)) (to (t t)) (rate ncrna_rate_scale * LNUC_RNUC_cc2tt))
  (mutate (from (c c)) (to (c _)) (rate ncrna_rate_scale * LNUC_RNUC_cc2cX))
  (mutate (from (g c)) (to (a a)) (rate ncrna_rate_scale * LNUC_RNUC_gc2aa))
  (mutate (from (g c)) (to (c a)) (rate ncrna_rate_scale * LNUC_RNUC_gc2ca))
  (mutate (from (g c)) (to (g a)) (rate ncrna_rate_scale * LNUC_RNUC_gc2ga))
  (mutate (from (g c)) (to (t a)) (rate ncrna_rate_scale * LNUC_RNUC_gc2ta))
  (mutate (from (g c)) (to (a c)) (rate ncrna_rate_scale * LNUC_RNUC_gc2ac))
  (mutate (from (g c)) (to (c c)) (rate ncrna_rate_scale * LNUC_RNUC_gc2cc))
  (mutate (from (g c)) (to (t c)) (rate ncrna_rate_scale * LNUC_RNUC_gc2tc))
  (mutate (from (g c)) (to (a g)) (rate ncrna_rate_scale * LNUC_RNUC_gc2ag))
  (mutate (from (g c)) (to (c g)) (rate ncrna_rate_scale * LNUC_RNUC_gc2cg))
  (mutate (from (g c)) (to (g g)) (rate ncrna_rate_scale * LNUC_RNUC_gc2gg))
  (mutate (from (g c)) (to (t g)) (rate ncrna_rate_scale * LNUC_RNUC_gc2tg))
  (mutate (from (g c)) (to (_ g)) (rate ncrna_rate_scale * LNUC_RNUC_gc2Xg))
  (mutate (from (g c)) (to (a t)) (rate ncrna_rate_scale * LNUC_RNUC_gc2at))
  (mutate (from (g c)) (to (c t)) (rate ncrna_rate_scale * LNUC_RNUC_gc2ct))
  (mutate (from (g c)) (to (g t)) (rate ncrna_rate_scale * LNUC_RNUC_gc2gt))
  (mutate (from (g c)) (to (t t)) (rate ncrna_rate_scale * LNUC_RNUC_gc2tt))
  (mutate (from (g c)) (to (g _)) (rate ncrna_rate_scale * LNUC_RNUC_gc2gX))
  (mutate (from (t c)) (to (a a)) (rate ncrna_rate_scale * LNUC_RNUC_tc2aa))
  (mutate (from (t c)) (to (c a)) (rate ncrna_rate_scale * LNUC_RNUC_tc2ca))
  (mutate (from (t c)) (to (g a)) (rate ncrna_rate_scale * LNUC_RNUC_tc2ga))
  (mutate (from (t c)) (to (t a)) (rate ncrna_rate_scale * LNUC_RNUC_tc2ta))
  (mutate (from (t c)) (to (a c)) (rate ncrna_rate_scale * LNUC_RNUC_tc2ac))
  (mutate (from (t c)) (to (c c)) (rate ncrna_rate_scale * LNUC_RNUC_tc2cc))
  (mutate (from (t c)) (to (g c)) (rate ncrna_rate_scale * LNUC_RNUC_tc2gc))
  (mutate (from (t c)) (to (a g)) (rate ncrna_rate_scale * LNUC_RNUC_tc2ag))
  (mutate (from (t c)) (to (c g)) (rate ncrna_rate_scale * LNUC_RNUC_tc2cg))
  (mutate (from (t c)) (to (g g)) (rate ncrna_rate_scale * LNUC_RNUC_tc2gg))
  (mutate (from (t c)) (to (t g)) (rate ncrna_rate_scale * LNUC_RNUC_tc2tg))
  (mutate (from (t c)) (to (a t)) (rate ncrna_rate_scale * LNUC_RNUC_tc2at))
  (mutate (from (t c)) (to (c t)) (rate ncrna_rate_scale * LNUC_RNUC_tc2ct))
  (mutate (from (t c)) (to (g t)) (rate ncrna_rate_scale * LNUC_RNUC_tc2gt))
  (mutate (from (t c)) (to (t t)) (rate ncrna_rate_scale * LNUC_RNUC_tc2tt))
  (mutate (from (t c)) (to (_ t)) (rate ncrna_rate_scale * LNUC_RNUC_tc2Xt))
  (mutate (from (t c)) (to (t _)) (rate ncrna_rate_scale * LNUC_RNUC_tc2tX))
  (mutate (from (_ c)) (to (_ a)) (rate ncrna_rate_scale * LNUC_RNUC_Xc2Xa))
  (mutate (from (_ c)) (to (a c)) (rate ncrna_rate_scale * LNUC_RNUC_Xc2ac))
  (mutate (from (_ c)) (to (c c)) (rate ncrna_rate_scale * LNUC_RNUC_Xc2cc))
  (mutate (from (_ c)) (to (g c)) (rate ncrna_rate_scale * LNUC_RNUC_Xc2gc))
  (mutate (from (_ c)) (to (t c)) (rate ncrna_rate_scale * LNUC_RNUC_Xc2tc))
  (mutate (from (_ c)) (to (_ g)) (rate ncrna_rate_scale * LNUC_RNUC_Xc2Xg))
  (mutate (from (_ c)) (to (_ t)) (rate ncrna_rate_scale * LNUC_RNUC_Xc2Xt))
  (mutate (from (_ c)) (to (_ _)) (rate ncrna_rate_scale * LNUC_RNUC_Xc2XX))
  (mutate (from (a g)) (to (a a)) (rate ncrna_rate_scale * LNUC_RNUC_ag2aa))
  (mutate (from (a g)) (to (c a)) (rate ncrna_rate_scale * LNUC_RNUC_ag2ca))
  (mutate (from (a g)) (to (g a)) (rate ncrna_rate_scale * LNUC_RNUC_ag2ga))
  (mutate (from (a g)) (to (t a)) (rate ncrna_rate_scale * LNUC_RNUC_ag2ta))
  (mutate (from (a g)) (to (_ a)) (rate ncrna_rate_scale * LNUC_RNUC_ag2Xa))
  (mutate (from (a g)) (to (a c)) (rate ncrna_rate_scale * LNUC_RNUC_ag2ac))
  (mutate (from (a g)) (to (c c)) (rate ncrna_rate_scale * LNUC_RNUC_ag2cc))
  (mutate (from (a g)) (to (g c)) (rate ncrna_rate_scale * LNUC_RNUC_ag2gc))
  (mutate (from (a g)) (to (t c)) (rate ncrna_rate_scale * LNUC_RNUC_ag2tc))
  (mutate (from (a g)) (to (c g)) (rate ncrna_rate_scale * LNUC_RNUC_ag2cg))
  (mutate (from (a g)) (to (g g)) (rate ncrna_rate_scale * LNUC_RNUC_ag2gg))
  (mutate (from (a g)) (to (t g)) (rate ncrna_rate_scale * LNUC_RNUC_ag2tg))
  (mutate (from (a g)) (to (a t)) (rate ncrna_rate_scale * LNUC_RNUC_ag2at))
  (mutate (from (a g)) (to (c t)) (rate ncrna_rate_scale * LNUC_RNUC_ag2ct))
  (mutate (from (a g)) (to (g t)) (rate ncrna_rate_scale * LNUC_RNUC_ag2gt))
  (mutate (from (a g)) (to (t t)) (rate ncrna_rate_scale * LNUC_RNUC_ag2tt))
  (mutate (from (a g)) (to (a _)) (rate ncrna_rate_scale * LNUC_RNUC_ag2aX))
  (mutate (from (c g)) (to (a a)) (rate ncrna_rate_scale * LNUC_RNUC_cg2aa))
  (mutate (from (c g)) (to (c a)) (rate ncrna_rate_scale * LNUC_RNUC_cg2ca))
  (mutate (from (c g)) (to (g a)) (rate ncrna_rate_scale * LNUC_RNUC_cg2ga))
  (mutate (from (c g)) (to (t a)) (rate ncrna_rate_scale * LNUC_RNUC_cg2ta))
  (mutate (from (c g)) (to (a c)) (rate ncrna_rate_scale * LNUC_RNUC_cg2ac))
  (mutate (from (c g)) (to (c c)) (rate ncrna_rate_scale * LNUC_RNUC_cg2cc))
  (mutate (from (c g)) (to (g c)) (rate ncrna_rate_scale * LNUC_RNUC_cg2gc))
  (mutate (from (c g)) (to (t c)) (rate ncrna_rate_scale * LNUC_RNUC_cg2tc))
  (mutate (from (c g)) (to (_ c)) (rate ncrna_rate_scale * LNUC_RNUC_cg2Xc))
  (mutate (from (c g)) (to (a g)) (rate ncrna_rate_scale * LNUC_RNUC_cg2ag))
  (mutate (from (c g)) (to (g g)) (rate ncrna_rate_scale * LNUC_RNUC_cg2gg))
  (mutate (from (c g)) (to (t g)) (rate ncrna_rate_scale * LNUC_RNUC_cg2tg))
  (mutate (from (c g)) (to (a t)) (rate ncrna_rate_scale * LNUC_RNUC_cg2at))
  (mutate (from (c g)) (to (c t)) (rate ncrna_rate_scale * LNUC_RNUC_cg2ct))
  (mutate (from (c g)) (to (g t)) (rate ncrna_rate_scale * LNUC_RNUC_cg2gt))
  (mutate (from (c g)) (to (t t)) (rate ncrna_rate_scale * LNUC_RNUC_cg2tt))
  (mutate (from (c g)) (to (c _)) (rate ncrna_rate_scale * LNUC_RNUC_cg2cX))
  (mutate (from (g g)) (to (a a)) (rate ncrna_rate_scale * LNUC_RNUC_gg2aa))
  (mutate (from (g g)) (to (c a)) (rate ncrna_rate_scale * LNUC_RNUC_gg2ca))
  (mutate (from (g g)) (to (g a)) (rate ncrna_rate_scale * LNUC_RNUC_gg2ga))
  (mutate (from (g g)) (to (t a)) (rate ncrna_rate_scale * LNUC_RNUC_gg2ta))
  (mutate (from (g g)) (to (a c)) (rate ncrna_rate_scale * LNUC_RNUC_gg2ac))
  (mutate (from (g g)) (to (c c)) (rate ncrna_rate_scale * LNUC_RNUC_gg2cc))
  (mutate (from (g g)) (to (g c)) (rate ncrna_rate_scale * LNUC_RNUC_gg2gc))
  (mutate (from (g g)) (to (t c)) (rate ncrna_rate_scale * LNUC_RNUC_gg2tc))
  (mutate (from (g g)) (to (a g)) (rate ncrna_rate_scale * LNUC_RNUC_gg2ag))
  (mutate (from (g g)) (to (c g)) (rate ncrna_rate_scale * LNUC_RNUC_gg2cg))
  (mutate (from (g g)) (to (t g)) (rate ncrna_rate_scale * LNUC_RNUC_gg2tg))
  (mutate (from (g g)) (to (_ g)) (rate ncrna_rate_scale * LNUC_RNUC_gg2Xg))
  (mutate (from (g g)) (to (a t)) (rate ncrna_rate_scale * LNUC_RNUC_gg2at))
  (mutate (from (g g)) (to (c t)) (rate ncrna_rate_scale * LNUC_RNUC_gg2ct))
  (mutate (from (g g)) (to (g t)) (rate ncrna_rate_scale * LNUC_RNUC_gg2gt))
  (mutate (from (g g)) (to (t t)) (rate ncrna_rate_scale * LNUC_RNUC_gg2tt))
  (mutate (from (g g)) (to (g _)) (rate ncrna_rate_scale * LNUC_RNUC_gg2gX))
  (mutate (from (t g)) (to (a a)) (rate ncrna_rate_scale * LNUC_RNUC_tg2aa))
  (mutate (from (t g)) (to (c a)) (rate ncrna_rate_scale * LNUC_RNUC_tg2ca))
  (mutate (from (t g)) (to (g a)) (rate ncrna_rate_scale * LNUC_RNUC_tg2ga))
  (mutate (from (t g)) (to (t a)) (rate ncrna_rate_scale * LNUC_RNUC_tg2ta))
  (mutate (from (t g)) (to (a c)) (rate ncrna_rate_scale * LNUC_RNUC_tg2ac))
  (mutate (from (t g)) (to (c c)) (rate ncrna_rate_scale * LNUC_RNUC_tg2cc))
  (mutate (from (t g)) (to (g c)) (rate ncrna_rate_scale * LNUC_RNUC_tg2gc))
  (mutate (from (t g)) (to (t c)) (rate ncrna_rate_scale * LNUC_RNUC_tg2tc))
  (mutate (from (t g)) (to (a g)) (rate ncrna_rate_scale * LNUC_RNUC_tg2ag))
  (mutate (from (t g)) (to (c g)) (rate ncrna_rate_scale * LNUC_RNUC_tg2cg))
  (mutate (from (t g)) (to (g g)) (rate ncrna_rate_scale * LNUC_RNUC_tg2gg))
  (mutate (from (t g)) (to (a t)) (rate ncrna_rate_scale * LNUC_RNUC_tg2at))
  (mutate (from (t g)) (to (c t)) (rate ncrna_rate_scale * LNUC_RNUC_tg2ct))
  (mutate (from (t g)) (to (g t)) (rate ncrna_rate_scale * LNUC_RNUC_tg2gt))
  (mutate (from (t g)) (to (t t)) (rate ncrna_rate_scale * LNUC_RNUC_tg2tt))
  (mutate (from (t g)) (to (_ t)) (rate ncrna_rate_scale * LNUC_RNUC_tg2Xt))
  (mutate (from (t g)) (to (t _)) (rate ncrna_rate_scale * LNUC_RNUC_tg2tX))
  (mutate (from (_ g)) (to (_ a)) (rate ncrna_rate_scale * LNUC_RNUC_Xg2Xa))
  (mutate (from (_ g)) (to (_ c)) (rate ncrna_rate_scale * LNUC_RNUC_Xg2Xc))
  (mutate (from (_ g)) (to (a g)) (rate ncrna_rate_scale * LNUC_RNUC_Xg2ag))
  (mutate (from (_ g)) (to (c g)) (rate ncrna_rate_scale * LNUC_RNUC_Xg2cg))
  (mutate (from (_ g)) (to (g g)) (rate ncrna_rate_scale * LNUC_RNUC_Xg2gg))
  (mutate (from (_ g)) (to (t g)) (rate ncrna_rate_scale * LNUC_RNUC_Xg2tg))
  (mutate (from (_ g)) (to (_ t)) (rate ncrna_rate_scale * LNUC_RNUC_Xg2Xt))
  (mutate (from (_ g)) (to (_ _)) (rate ncrna_rate_scale * LNUC_RNUC_Xg2XX))
  (mutate (from (a t)) (to (a a)) (rate ncrna_rate_scale * LNUC_RNUC_at2aa))
  (mutate (from (a t)) (to (c a)) (rate ncrna_rate_scale * LNUC_RNUC_at2ca))
  (mutate (from (a t)) (to (g a)) (rate ncrna_rate_scale * LNUC_RNUC_at2ga))
  (mutate (from (a t)) (to (t a)) (rate ncrna_rate_scale * LNUC_RNUC_at2ta))
  (mutate (from (a t)) (to (_ a)) (rate ncrna_rate_scale * LNUC_RNUC_at2Xa))
  (mutate (from (a t)) (to (a c)) (rate ncrna_rate_scale * LNUC_RNUC_at2ac))
  (mutate (from (a t)) (to (c c)) (rate ncrna_rate_scale * LNUC_RNUC_at2cc))
  (mutate (from (a t)) (to (g c)) (rate ncrna_rate_scale * LNUC_RNUC_at2gc))
  (mutate (from (a t)) (to (t c)) (rate ncrna_rate_scale * LNUC_RNUC_at2tc))
  (mutate (from (a t)) (to (a g)) (rate ncrna_rate_scale * LNUC_RNUC_at2ag))
  (mutate (from (a t)) (to (c g)) (rate ncrna_rate_scale * LNUC_RNUC_at2cg))
  (mutate (from (a t)) (to (g g)) (rate ncrna_rate_scale * LNUC_RNUC_at2gg))
  (mutate (from (a t)) (to (t g)) (rate ncrna_rate_scale * LNUC_RNUC_at2tg))
  (mutate (from (a t)) (to (c t)) (rate ncrna_rate_scale * LNUC_RNUC_at2ct))
  (mutate (from (a t)) (to (g t)) (rate ncrna_rate_scale * LNUC_RNUC_at2gt))
  (mutate (from (a t)) (to (t t)) (rate ncrna_rate_scale * LNUC_RNUC_at2tt))
  (mutate (from (a t)) (to (a _)) (rate ncrna_rate_scale * LNUC_RNUC_at2aX))
  (mutate (from (c t)) (to (a a)) (rate ncrna_rate_scale * LNUC_RNUC_ct2aa))
  (mutate (from (c t)) (to (c a)) (rate ncrna_rate_scale * LNUC_RNUC_ct2ca))
  (mutate (from (c t)) (to (g a)) (rate ncrna_rate_scale * LNUC_RNUC_ct2ga))
  (mutate (from (c t)) (to (t a)) (rate ncrna_rate_scale * LNUC_RNUC_ct2ta))
  (mutate (from (c t)) (to (a c)) (rate ncrna_rate_scale * LNUC_RNUC_ct2ac))
  (mutate (from (c t)) (to (c c)) (rate ncrna_rate_scale * LNUC_RNUC_ct2cc))
  (mutate (from (c t)) (to (g c)) (rate ncrna_rate_scale * LNUC_RNUC_ct2gc))
  (mutate (from (c t)) (to (t c)) (rate ncrna_rate_scale * LNUC_RNUC_ct2tc))
  (mutate (from (c t)) (to (_ c)) (rate ncrna_rate_scale * LNUC_RNUC_ct2Xc))
  (mutate (from (c t)) (to (a g)) (rate ncrna_rate_scale * LNUC_RNUC_ct2ag))
  (mutate (from (c t)) (to (c g)) (rate ncrna_rate_scale * LNUC_RNUC_ct2cg))
  (mutate (from (c t)) (to (g g)) (rate ncrna_rate_scale * LNUC_RNUC_ct2gg))
  (mutate (from (c t)) (to (t g)) (rate ncrna_rate_scale * LNUC_RNUC_ct2tg))
  (mutate (from (c t)) (to (a t)) (rate ncrna_rate_scale * LNUC_RNUC_ct2at))
  (mutate (from (c t)) (to (g t)) (rate ncrna_rate_scale * LNUC_RNUC_ct2gt))
  (mutate (from (c t)) (to (t t)) (rate ncrna_rate_scale * LNUC_RNUC_ct2tt))
  (mutate (from (c t)) (to (c _)) (rate ncrna_rate_scale * LNUC_RNUC_ct2cX))
  (mutate (from (g t)) (to (a a)) (rate ncrna_rate_scale * LNUC_RNUC_gt2aa))
  (mutate (from (g t)) (to (c a)) (rate ncrna_rate_scale * LNUC_RNUC_gt2ca))
  (mutate (from (g t)) (to (g a)) (rate ncrna_rate_scale * LNUC_RNUC_gt2ga))
  (mutate (from (g t)) (to (t a)) (rate ncrna_rate_scale * LNUC_RNUC_gt2ta))
  (mutate (from (g t)) (to (a c)) (rate ncrna_rate_scale * LNUC_RNUC_gt2ac))
  (mutate (from (g t)) (to (c c)) (rate ncrna_rate_scale * LNUC_RNUC_gt2cc))
  (mutate (from (g t)) (to (g c)) (rate ncrna_rate_scale * LNUC_RNUC_gt2gc))
  (mutate (from (g t)) (to (t c)) (rate ncrna_rate_scale * LNUC_RNUC_gt2tc))
  (mutate (from (g t)) (to (a g)) (rate ncrna_rate_scale * LNUC_RNUC_gt2ag))
  (mutate (from (g t)) (to (c g)) (rate ncrna_rate_scale * LNUC_RNUC_gt2cg))
  (mutate (from (g t)) (to (g g)) (rate ncrna_rate_scale * LNUC_RNUC_gt2gg))
  (mutate (from (g t)) (to (t g)) (rate ncrna_rate_scale * LNUC_RNUC_gt2tg))
  (mutate (from (g t)) (to (_ g)) (rate ncrna_rate_scale * LNUC_RNUC_gt2Xg))
  (mutate (from (g t)) (to (a t)) (rate ncrna_rate_scale * LNUC_RNUC_gt2at))
  (mutate (from (g t)) (to (c t)) (rate ncrna_rate_scale * LNUC_RNUC_gt2ct))
  (mutate (from (g t)) (to (t t)) (rate ncrna_rate_scale * LNUC_RNUC_gt2tt))
  (mutate (from (g t)) (to (g _)) (rate ncrna_rate_scale * LNUC_RNUC_gt2gX))
  (mutate (from (t t)) (to (a a)) (rate ncrna_rate_scale * LNUC_RNUC_tt2aa))
  (mutate (from (t t)) (to (c a)) (rate ncrna_rate_scale * LNUC_RNUC_tt2ca))
  (mutate (from (t t)) (to (g a)) (rate ncrna_rate_scale * LNUC_RNUC_tt2ga))
  (mutate (from (t t)) (to (t a)) (rate ncrna_rate_scale * LNUC_RNUC_tt2ta))
  (mutate (from (t t)) (to (a c)) (rate ncrna_rate_scale * LNUC_RNUC_tt2ac))
  (mutate (from (t t)) (to (c c)) (rate ncrna_rate_scale * LNUC_RNUC_tt2cc))
  (mutate (from (t t)) (to (g c)) (rate ncrna_rate_scale * LNUC_RNUC_tt2gc))
  (mutate (from (t t)) (to (t c)) (rate ncrna_rate_scale * LNUC_RNUC_tt2tc))
  (mutate (from (t t)) (to (a g)) (rate ncrna_rate_scale * LNUC_RNUC_tt2ag))
  (mutate (from (t t)) (to (c g)) (rate ncrna_rate_scale * LNUC_RNUC_tt2cg))
  (mutate (from (t t)) (to (g g)) (rate ncrna_rate_scale * LNUC_RNUC_tt2gg))
  (mutate (from (t t)) (to (t g)) (rate ncrna_rate_scale * LNUC_RNUC_tt2tg))
  (mutate (from (t t)) (to (a t)) (rate ncrna_rate_scale * LNUC_RNUC_tt2at))
  (mutate (from (t t)) (to (c t)) (rate ncrna_rate_scale * LNUC_RNUC_tt2ct))
  (mutate (from (t t)) (to (g t)) (rate ncrna_rate_scale * LNUC_RNUC_tt2gt))
  (mutate (from (t t)) (to (_ t)) (rate ncrna_rate_scale * LNUC_RNUC_tt2Xt))
  (mutate (from (t t)) (to (t _)) (rate ncrna_rate_scale * LNUC_RNUC_tt2tX))
  (mutate (from (_ t)) (to (_ a)) (rate ncrna_rate_scale * LNUC_RNUC_Xt2Xa))
  (mutate (from (_ t)) (to (_ c)) (rate ncrna_rate_scale * LNUC_RNUC_Xt2Xc))
  (mutate (from (_ t)) (to (_ g)) (rate ncrna_rate_scale * LNUC_RNUC_Xt2Xg))
  (mutate (from (_ t)) (to (a t)) (rate ncrna_rate_scale * LNUC_RNUC_Xt2at))
  (mutate (from (_ t)) (to (c t)) (rate ncrna_rate_scale * LNUC_RNUC_Xt2ct))
  (mutate (from (_ t)) (to (g t)) (rate ncrna_rate_scale * LNUC_RNUC_Xt2gt))
  (mutate (from (_ t)) (to (t t)) (rate ncrna_rate_scale * LNUC_RNUC_Xt2tt))
  (mutate (from (_ t)) (to (_ _)) (rate ncrna_rate_scale * LNUC_RNUC_Xt2XX))
  (mutate (from (a _)) (to (a a)) (rate ncrna_rate_scale * LNUC_RNUC_aX2aa))
  (mutate (from (a _)) (to (a c)) (rate ncrna_rate_scale * LNUC_RNUC_aX2ac))
  (mutate (from (a _)) (to (a g)) (rate ncrna_rate_scale * LNUC_RNUC_aX2ag))
  (mutate (from (a _)) (to (a t)) (rate ncrna_rate_scale * LNUC_RNUC_aX2at))
  (mutate (from (a _)) (to (c _)) (rate ncrna_rate_scale * LNUC_RNUC_aX2cX))
  (mutate (from (a _)) (to (g _)) (rate ncrna_rate_scale * LNUC_RNUC_aX2gX))
  (mutate (from (a _)) (to (t _)) (rate ncrna_rate_scale * LNUC_RNUC_aX2tX))
  (mutate (from (a _)) (to (_ _)) (rate ncrna_rate_scale * LNUC_RNUC_aX2XX))
  (mutate (from (c _)) (to (c a)) (rate ncrna_rate_scale * LNUC_RNUC_cX2ca))
  (mutate (from (c _)) (to (c c)) (rate ncrna_rate_scale * LNUC_RNUC_cX2cc))
  (mutate (from (c _)) (to (c g)) (rate ncrna_rate_scale * LNUC_RNUC_cX2cg))
  (mutate (from (c _)) (to (c t)) (rate ncrna_rate_scale * LNUC_RNUC_cX2ct))
  (mutate (from (c _)) (to (a _)) (rate ncrna_rate_scale * LNUC_RNUC_cX2aX))
  (mutate (from (c _)) (to (g _)) (rate ncrna_rate_scale * LNUC_RNUC_cX2gX))
  (mutate (from (c _)) (to (t _)) (rate ncrna_rate_scale * LNUC_RNUC_cX2tX))
  (mutate (from (c _)) (to (_ _)) (rate ncrna_rate_scale * LNUC_RNUC_cX2XX))
  (mutate (from (g _)) (to (g a)) (rate ncrna_rate_scale * LNUC_RNUC_gX2ga))
  (mutate (from (g _)) (to (g c)) (rate ncrna_rate_scale * LNUC_RNUC_gX2gc))
  (mutate (from (g _)) (to (g g)) (rate ncrna_rate_scale * LNUC_RNUC_gX2gg))
  (mutate (from (g _)) (to (g t)) (rate ncrna_rate_scale * LNUC_RNUC_gX2gt))
  (mutate (from (g _)) (to (a _)) (rate ncrna_rate_scale * LNUC_RNUC_gX2aX))
  (mutate (from (g _)) (to (c _)) (rate ncrna_rate_scale * LNUC_RNUC_gX2cX))
  (mutate (from (g _)) (to (t _)) (rate ncrna_rate_scale * LNUC_RNUC_gX2tX))
  (mutate (from (g _)) (to (_ _)) (rate ncrna_rate_scale * LNUC_RNUC_gX2XX))
  (mutate (from (t _)) (to (t a)) (rate ncrna_rate_scale * LNUC_RNUC_tX2ta))
  (mutate (from (t _)) (to (t c)) (rate ncrna_rate_scale * LNUC_RNUC_tX2tc))
  (mutate (from (t _)) (to (t g)) (rate ncrna_rate_scale * LNUC_RNUC_tX2tg))
  (mutate (from (t _)) (to (t t)) (rate ncrna_rate_scale * LNUC_RNUC_tX2tt))
  (mutate (from (t _)) (to (a _)) (rate ncrna_rate_scale * LNUC_RNUC_tX2aX))
  (mutate (from (t _)) (to (c _)) (rate ncrna_rate_scale * LNUC_RNUC_tX2cX))
  (mutate (from (t _)) (to (g _)) (rate ncrna_rate_scale * LNUC_RNUC_tX2gX))
  (mutate (from (t _)) (to (_ _)) (rate ncrna_rate_scale * LNUC_RNUC_tX2XX))
  (mutate (from (_ _)) (to (_ a)) (rate ncrna_rate_scale * LNUC_RNUC_XX2Xa))
  (mutate (from (_ _)) (to (_ c)) (rate ncrna_rate_scale * LNUC_RNUC_XX2Xc))
  (mutate (from (_ _)) (to (_ g)) (rate ncrna_rate_scale * LNUC_RNUC_XX2Xg))
  (mutate (from (_ _)) (to (_ t)) (rate ncrna_rate_scale * LNUC_RNUC_XX2Xt))
  (mutate (from (_ _)) (to (a _)) (rate ncrna_rate_scale * LNUC_RNUC_XX2aX))
  (mutate (from (_ _)) (to (c _)) (rate ncrna_rate_scale * LNUC_RNUC_XX2cX))
  (mutate (from (_ _)) (to (g _)) (rate ncrna_rate_scale * LNUC_RNUC_XX2gX))
  (mutate (from (_ _)) (to (t _)) (rate ncrna_rate_scale * LNUC_RNUC_XX2tX))
 )  ;; end chain LNUC RNUC

 ;; Unpaired bases
 (chain
  (update-policy parametric)
  (terminal (NUC))

  ;; initial probability distribution
  (initial (state (a)) (prob NUC_a))
  (initial (state (c)) (prob NUC_c))
  (initial (state (g)) (prob NUC_g))
  (initial (state (t)) (prob NUC_t))
  (initial (state (_)) (prob NUC_X))

  ;; mutation rates
  (mutate (from (a)) (to (c)) (rate ncrna_rate_scale * NUC_a2c))
  (mutate (from (a)) (to (g)) (rate ncrna_rate_scale * NUC_a2g))
  (mutate (from (a)) (to (t)) (rate ncrna_rate_scale * NUC_a2t))
  (mutate (from (a)) (to (_)) (rate ncrna_rate_scale * NUC_a2X))
  (mutate (from (c)) (to (a)) (rate ncrna_rate_scale * NUC_c2a))
  (mutate (from (c)) (to (g)) (rate ncrna_rate_scale * NUC_c2g))
  (mutate (from (c)) (to (t)) (rate ncrna_rate_scale * NUC_c2t))
  (mutate (from (c)) (to (_)) (rate ncrna_rate_scale * NUC_c2X))
  (mutate (from (g)) (to (a)) (rate ncrna_rate_scale * NUC_g2a))
  (mutate (from (g)) (to (c)) (rate ncrna_rate_scale * NUC_g2c))
  (mutate (from (g)) (to (t)) (rate ncrna_rate_scale * NUC_g2t))
  (mutate (from (g)) (to (_)) (rate ncrna_rate_scale * NUC_g2X))
  (mutate (from (t)) (to (a)) (rate ncrna_rate_scale * NUC_t2a))
  (mutate (from (t)) (to (c)) (rate ncrna_rate_scale * NUC_t2c))
  (mutate (from (t)) (to (g)) (rate ncrna_rate_scale * NUC_t2g))
  (mutate (from (t)) (to (_)) (rate ncrna_rate_scale * NUC_t2X))
  (mutate (from (_)) (to (a)) (rate ncrna_rate_scale * NUC_X2a))
  (mutate (from (_)) (to (c)) (rate ncrna_rate_scale * NUC_X2c))
  (mutate (from (_)) (to (g)) (rate ncrna_rate_scale * NUC_X2g))
  (mutate (from (_)) (to (t)) (rate ncrna_rate_scale * NUC_X2t))
 )  ;; end chain NUC


;;;;;;;;;;;;;;;;
;; TRANSPOSON ;;
;;;;;;;;;;;;;;;;

;; (LTIR RTIR) initial distribution is identical to (LNUC RNUC).
;; Subsequent evolution (rates) are according to neutral (cf pseudogene gene).

;; Basepairs
 (chain
  (update-policy parametric)
  (terminal (LTIR RTIR))

  ;; initial probability distribution
  (initial (state (a a)) (prob ~initGap * LNUC_RNUC_aa))
  (initial (state (c a)) (prob ~initGap * LNUC_RNUC_ca))
  (initial (state (g a)) (prob ~initGap * LNUC_RNUC_ga))
  (initial (state (t a)) (prob ~initGap * LNUC_RNUC_ta))
  (initial (state (_ a)) (prob initGap * LNUC_RNUC_Xa))
  (initial (state (a c)) (prob ~initGap * LNUC_RNUC_ac))
  (initial (state (c c)) (prob ~initGap * LNUC_RNUC_cc))
  (initial (state (g c)) (prob ~initGap * LNUC_RNUC_gc))
  (initial (state (t c)) (prob ~initGap * LNUC_RNUC_tc))
  (initial (state (_ c)) (prob initGap * LNUC_RNUC_Xc))
  (initial (state (a g)) (prob ~initGap * LNUC_RNUC_ag))
  (initial (state (c g)) (prob ~initGap * LNUC_RNUC_cg))
  (initial (state (g g)) (prob ~initGap * LNUC_RNUC_gg))
  (initial (state (t g)) (prob ~initGap * LNUC_RNUC_tg))
  (initial (state (_ g)) (prob initGap * LNUC_RNUC_Xg))
  (initial (state (a t)) (prob ~initGap * LNUC_RNUC_at))
  (initial (state (c t)) (prob ~initGap * LNUC_RNUC_ct))
  (initial (state (g t)) (prob ~initGap * LNUC_RNUC_gt))
  (initial (state (t t)) (prob ~initGap * LNUC_RNUC_tt))
  (initial (state (_ t)) (prob initGap * LNUC_RNUC_Xt))
  (initial (state (a _)) (prob initGap * LNUC_RNUC_aX))
  (initial (state (c _)) (prob initGap * LNUC_RNUC_cX))
  (initial (state (g _)) (prob initGap * LNUC_RNUC_gX))
  (initial (state (t _)) (prob initGap * LNUC_RNUC_tX))
  (initial (state (_ _)) (prob initGap * LNUC_RNUC_XX))

  ;; mutation rates
  (mutate (from (a a)) (to (a c)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (a a)) (to (a g)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (a a)) (to (a t)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (a a)) (to (a _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (a a)) (to (c a)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (a a)) (to (g a)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (a a)) (to (t a)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (a a)) (to (_ a)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (a c)) (to (a a)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (a c)) (to (a g)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (a c)) (to (a t)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (a c)) (to (a _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (a c)) (to (c c)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (a c)) (to (g c)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (a c)) (to (t c)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (a c)) (to (_ c)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (a g)) (to (a a)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (a g)) (to (a c)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (a g)) (to (a t)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (a g)) (to (a _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (a g)) (to (c g)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (a g)) (to (g g)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (a g)) (to (t g)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (a g)) (to (_ g)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (a t)) (to (a a)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (a t)) (to (a c)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (a t)) (to (a g)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (a t)) (to (a _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (a t)) (to (c t)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (a t)) (to (g t)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (a t)) (to (t t)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (a t)) (to (_ t)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (a _)) (to (a a)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (a _)) (to (a c)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (a _)) (to (a g)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (a _)) (to (a t)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (a _)) (to (c _)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (a _)) (to (g _)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (a _)) (to (t _)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (a _)) (to (_ _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (c a)) (to (a a)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (c a)) (to (c c)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (c a)) (to (c g)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (c a)) (to (c t)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (c a)) (to (c _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (c a)) (to (g a)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (c a)) (to (t a)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (c a)) (to (_ a)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (c c)) (to (a c)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (c c)) (to (c a)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (c c)) (to (c g)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (c c)) (to (c t)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (c c)) (to (c _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (c c)) (to (g c)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (c c)) (to (t c)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (c c)) (to (_ c)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (c g)) (to (a g)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (c g)) (to (c a)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (c g)) (to (c c)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (c g)) (to (c t)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (c g)) (to (c _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (c g)) (to (g g)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (c g)) (to (t g)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (c g)) (to (_ g)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (c t)) (to (a t)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (c t)) (to (c a)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (c t)) (to (c c)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (c t)) (to (c g)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (c t)) (to (c _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (c t)) (to (g t)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (c t)) (to (t t)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (c t)) (to (_ t)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (c _)) (to (a _)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (c _)) (to (c a)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (c _)) (to (c c)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (c _)) (to (c g)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (c _)) (to (c t)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (c _)) (to (g _)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (c _)) (to (t _)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (c _)) (to (_ _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (g a)) (to (a a)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (g a)) (to (c a)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (g a)) (to (g c)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (g a)) (to (g g)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (g a)) (to (g t)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (g a)) (to (g _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (g a)) (to (t a)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (g a)) (to (_ a)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (g c)) (to (a c)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (g c)) (to (c c)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (g c)) (to (g a)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (g c)) (to (g g)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (g c)) (to (g t)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (g c)) (to (g _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (g c)) (to (t c)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (g c)) (to (_ c)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (g g)) (to (a g)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (g g)) (to (c g)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (g g)) (to (g a)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (g g)) (to (g c)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (g g)) (to (g t)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (g g)) (to (g _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (g g)) (to (t g)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (g g)) (to (_ g)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (g t)) (to (a t)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (g t)) (to (c t)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (g t)) (to (g a)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (g t)) (to (g c)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (g t)) (to (g g)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (g t)) (to (g _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (g t)) (to (t t)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (g t)) (to (_ t)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (g _)) (to (a _)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (g _)) (to (c _)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (g _)) (to (g a)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (g _)) (to (g c)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (g _)) (to (g g)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (g _)) (to (g t)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (g _)) (to (t _)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (g _)) (to (_ _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (t a)) (to (a a)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (t a)) (to (c a)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (t a)) (to (g a)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (t a)) (to (t c)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (t a)) (to (t g)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (t a)) (to (t t)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (t a)) (to (t _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (t a)) (to (_ a)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (t c)) (to (a c)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (t c)) (to (c c)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (t c)) (to (g c)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (t c)) (to (t a)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (t c)) (to (t g)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (t c)) (to (t t)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (t c)) (to (t _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (t c)) (to (_ c)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (t g)) (to (a g)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (t g)) (to (c g)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (t g)) (to (g g)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (t g)) (to (t a)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (t g)) (to (t c)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (t g)) (to (t t)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (t g)) (to (t _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (t g)) (to (_ g)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (t t)) (to (a t)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (t t)) (to (c t)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (t t)) (to (g t)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (t t)) (to (t a)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (t t)) (to (t c)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (t t)) (to (t g)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (t t)) (to (t _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (t t)) (to (_ t)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (t _)) (to (a _)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (t _)) (to (c _)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (t _)) (to (g _)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (t _)) (to (t a)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (t _)) (to (t c)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (t _)) (to (t g)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (t _)) (to (t t)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (t _)) (to (_ _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (_ a)) (to (a a)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ a)) (to (c a)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ a)) (to (g a)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ a)) (to (t a)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ a)) (to (_ c)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (_ a)) (to (_ g)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (_ a)) (to (_ t)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (_ a)) (to (_ _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (_ c)) (to (a c)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ c)) (to (c c)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ c)) (to (g c)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ c)) (to (t c)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ c)) (to (_ a)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (_ c)) (to (_ g)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (_ c)) (to (_ t)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (_ c)) (to (_ _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (_ g)) (to (a g)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ g)) (to (c g)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ g)) (to (g g)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ g)) (to (t g)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ g)) (to (_ a)) (rate neutral_rate_scale * X_c2t_g2a))
  (mutate (from (_ g)) (to (_ c)) (rate neutral_rate_scale * X_c2g_g2c))
  (mutate (from (_ g)) (to (_ t)) (rate neutral_rate_scale * X_c2a_g2t))
  (mutate (from (_ g)) (to (_ _)) (rate neutral_rate_scale * X_c2gap_g2gap))
  (mutate (from (_ t)) (to (a t)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ t)) (to (c t)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ t)) (to (g t)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ t)) (to (t t)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ t)) (to (_ a)) (rate neutral_rate_scale * X_a2t_t2a))
  (mutate (from (_ t)) (to (_ c)) (rate neutral_rate_scale * X_a2g_t2c))
  (mutate (from (_ t)) (to (_ g)) (rate neutral_rate_scale * X_a2c_t2g))
  (mutate (from (_ t)) (to (_ _)) (rate neutral_rate_scale * X_a2gap_t2gap))
  (mutate (from (_ _)) (to (a _)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ _)) (to (c _)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ _)) (to (g _)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ _)) (to (t _)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ _)) (to (_ a)) (rate neutral_rate_scale * X_gap2a_gap2t))
  (mutate (from (_ _)) (to (_ c)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ _)) (to (_ g)) (rate neutral_rate_scale * X_gap2c_gap2g))
  (mutate (from (_ _)) (to (_ t)) (rate neutral_rate_scale * X_gap2a_gap2t))

 )  ;; end chain LTIR RTIR

)  ;; end grammar

;; Standard DNA alphabet + gaps as a 5th character
(alphabet
 (name GappedDNA)
 (token (a c g t _))
 (complement (t g c a _))
 (extend (to n) (from a) (from c) (from g) (from t))
 (extend (to x) (from a) (from c) (from g) (from t))
 (extend (to u) (from t))
 (extend (to r) (from a) (from g))
 (extend (to y) (from c) (from t))
 (extend (to m) (from a) (from c))
 (extend (to k) (from g) (from t))
 (extend (to s) (from c) (from g))
 (extend (to w) (from a) (from t))
 (extend (to h) (from a) (from c) (from t))
 (extend (to b) (from c) (from g) (from t))
 (extend (to v) (from a) (from c) (from g))
 (extend (to d) (from a) (from g) (from t))
 (wildcard *)
)  ;; end alphabet GappedDNA