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<document id="3A07B083E7688039E4B73B620CB04CEC" ID-DOI="10.11646/zootaxa.5351.4.2" ID-ISSN="1175-5326" ID-ZooBank="786A8C61-AC0F-475C-B185-DD614C59AFDA" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1695991472363" checkinUser="plazi" docAuthor="Shapovalov, Andrey M. & Bagaturov, Mikhail F." docDate="2023" docId="3E6A87DAFFCEFF984993FA85FC1BF93E" docLanguage="en" docName="zootaxa.5351.4.2.pdf" docOrigin="Zootaxa 5351 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5351.4.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lethrus inermis Reitter 1897" docType="treatment" docVersion="1" lastPageNumber="436" masterDocId="C253FFA2FFCFFF994904FFB0FF99FFAA" masterDocTitle="A review of the subgenus Mesolethrus Nikolajev, 2003 of the genus Lethrus Scopoli, 1777 (Coleoptera: Geotrupidae: Lethrinae) with description of a new species" masterLastPageNumber="452" masterPageNumber="435" pageNumber="436" updateTime="1695991562692" updateUser="GgImagineBatch" zenodo-license-document="CLOSED"> | ||
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<mods:title id="24181C3EA5EB1D2EEF955DC4C5F47761">A review of the subgenus Mesolethrus Nikolajev, 2003 of the genus Lethrus Scopoli, 1777 (Coleoptera: Geotrupidae: Lethrinae) with description of a new species</mods:title> | ||
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<mods:namePart id="C44EBA858D67F6056646FF0FB366BF81">Shapovalov, Andrey M.</mods:namePart> | ||
<mods:affiliation id="B79A3F04782AA2ADB4A27ED30DDAB326">Institute of Zoology RK, 93 al-Farabi Str., Almaty, 050060, Kazakhstan.</mods:affiliation> | ||
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<mods:namePart id="382C5F01BF8108D5A8A5BACA496BE008">Bagaturov, Mikhail F.</mods:namePart> | ||
<mods:affiliation id="0A3897EEC72530B69302A8EEC9198111">Athens Institute for Education & Research, 8 Valaoritou Str., Kolonaki, 10671 Athens, Greece. & State Budgetary Enterprise “ Leningrad Zoo ”, 1 Alexandrovskiy park, 197198 St. Petersburg, Russia. & The International Union for Conservation of Nature, 17 Marshal Vasilevskiy Str., 123182 Moscow, Russia.</mods:affiliation> | ||
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<mods:date id="322F2A6020EBE139E25FEC890B034E5C">2023</mods:date> | ||
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<treatment id="3E6A87DAFFCEFF984993FA85FC1BF93E" LSID="urn:lsid:plazi:treatment:3E6A87DAFFCEFF984993FA85FC1BF93E" httpUri="http://treatment.plazi.org/id/3E6A87DAFFCEFF984993FA85FC1BF93E" lastPageNumber="436" pageId="1" pageNumber="436"> | ||
<subSubSection id="FED96547FFCEFF984993FA85FD93FAFA" box="[151,522,1333,1360]" pageId="1" pageNumber="436" type="nomenclature"> | ||
<paragraph id="B67C36CCFFCEFF984993FA85FD93FAFA" blockId="1.[151,522,1333,1360]" box="[151,522,1333,1360]" pageId="1" pageNumber="436"> | ||
<heading id="ED3481A0FFCEFF984993FA85FD93FAFA" bold="true" box="[151,522,1333,1360]" fontSize="11" level="1" pageId="1" pageNumber="436" reason="1"> | ||
<emphasis id="84B7EADEFFCEFF984993FA85FD93FAFA" bold="true" box="[151,522,1333,1360]" pageId="1" pageNumber="436"> | ||
<taxonomicName id="71C34D4FFFCEFF984993FA85FEC1FAFA" ID-CoL="6Q4FF" ID-ENA="2609324" authorityName="Reitter" authorityYear="1897" box="[151,344,1333,1360]" class="Insecta" family="Geotrupidae" genus="Lethrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="436" phylum="Arthropoda" rank="species" species="inermis"> | ||
<emphasis id="84B7EADEFFCEFF984993FA85FEC1FAFA" bold="true" box="[151,344,1333,1360]" italics="true" pageId="1" pageNumber="436">Lethrus inermis</emphasis> | ||
</taxonomicName> | ||
-species group | ||
</emphasis> | ||
</heading> | ||
</paragraph> | ||
</subSubSection> | ||
<subSubSection id="FED96547FFCEFF984993FACDFC1BF93E" pageId="1" pageNumber="436" type="diagnosis"> | ||
<paragraph id="B67C36CCFFCEFF984993FACDFDC7F9DA" blockId="1.[151,1437,1405,1684]" pageId="1" pageNumber="436"> | ||
<emphasis id="84B7EADEFFCEFF984993FACDFE8BFA3D" bold="true" box="[151,274,1405,1431]" pageId="1" pageNumber="436">Diagnosis.</emphasis> | ||
Dorsal surface of mandibles in both sexes with sharp, elevated keels ( | ||
<figureCitation id="2EF82A49FFCEFF984D18FACEFB1EFA32" box="[1052,1159,1406,1432]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1761,1785]" captionTargetBox="[274,1320,197,1720]" captionTargetId="figure-17@3.[264,1323,179,1735]" captionTargetPageId="3" captionText="FIGURES 1–4. Lethrus (Mesolethrus) species, male, habitus. 1–2, L. inermis Reitter, 1897 (1—Dahanakiik; 2—20 km W Oqmachit); 3, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 4, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin)." pageId="1" pageNumber="436">Figs. 1–2</figureCitation> | ||
). Elytral striae definitely impressed ( | ||
<figureCitation id="2EF82A49FFCEFF98481BFA12FE08FA16" box="[287,401,1442,1468]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1761,1785]" captionTargetBox="[274,1320,197,1720]" captionTargetId="figure-17@3.[264,1323,179,1735]" captionTargetPageId="3" captionText="FIGURES 1–4. Lethrus (Mesolethrus) species, male, habitus. 1–2, L. inermis Reitter, 1897 (1—Dahanakiik; 2—20 km W Oqmachit); 3, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 4, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin)." pageId="1" pageNumber="436">Figs. 1–2</figureCitation> | ||
). Ventrites 1–5 with prominent central denticle posteriorly (rarely denticles indistinct), denticles apically with erect setae ( | ||
<figureCitation id="2EF82A49FFCEFF984B29FA76FD1DFA4A" box="[557,644,1478,1504]" captionStart="FIGURES 32–43" captionStartId="9.[151,264,1429,1453]" captionTargetBox="[154,1407,181,1392]" captionTargetId="figure-17@9.[151,1436,181,1405]" captionTargetPageId="9" captionText="FIGURES 32–43. Lethrus (Mesolethrus) species. 32, 39, L. inermis Reitter, 1897 (Dahanakiik); 33, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 34, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin); 35, 40–42, L. microbuccis Ballion, 1871 (35, 41—Golbo env., 39—Mullabulak, 40—20 km S Guldara); 36, L. elisae Nikolajev, 2001 (Shurob); 37–38, 40–41, L. dyonini A. Shapovalov & Bagaturov, new species, paratypes (environs of Tashsay). 32–37 gena, 38 left mandible, 39 abdomen (lateral view), 40–41 abdomen and apices of elytra (antero-ventral view; 40—male, 41—female), 42–43 tuft of long erect setae on subapical ventrite (lateral view; 42—male, 43—female)." pageId="1" pageNumber="436">Fig. 39</figureCitation> | ||
). Subapical ventrite with sparse semierect setae, density of setae not clearly differing between sexes. Parameres straight, narrowly rounded apically ( | ||
<figureCitation id="2EF82A49FFCEFF984D2BFA5AFB34F9AE" box="[1071,1197,1514,1540]" captionStart="FIGURES 9–31" captionStartId="8.[151,264,1808,1832]" captionTargetBox="[265,1328,191,1774]" captionTargetId="figure-17@8.[254,1332,181,1784]" captionTargetPageId="8" captionText="FIGURES 9–31. Lethrus (Mesolethrus) species, males genitalia. 9–10, 22–23, L. inermis Reitter, 1897 (9–10—20 km W Oqmachit, 22–23—Pashtamazar); 11–12, 24, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 13–14, 25–27, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin); 15–16, 28, L. microbuccis Ballion, 1871 (Golbo env.); 15–16, 28, L. microbuccis Ballion, 1871 (Golbo env.); 17–18, 29, L. elisae Nikolajev, 2001 (Shurob); 19–21, 30–31, L. dyonini A. Shapovalov & Bagaturov, new species, paratypes (environs of Tashsay). 9–21 aedeagus (9, 11, 13, 15, 17, 19–20—ventral view; 10, 12, 14, 16, 18, 21—lateral view), 22–31 sclerotized apex of endophallus (22, 24, 25, 28–30 dorsal view, 23, 26, 31—lateral view, 27—ventral view)." pageId="1" pageNumber="436">Figs. 9–10</figureCitation> | ||
); sclerotized apex of endophallus elongate, subclavate, noticeably curved in lateral view, in dorsal view appearing trilobed anteriorly, surface sharply spiny-granular ( | ||
<figureCitation id="2EF82A49FFCEFF9848FEF982FD1AF9E6" box="[506,643,1586,1612]" captionStart="FIGURES 9–31" captionStartId="8.[151,264,1808,1832]" captionTargetBox="[265,1328,191,1774]" captionTargetId="figure-17@8.[254,1332,181,1784]" captionTargetPageId="8" captionText="FIGURES 9–31. Lethrus (Mesolethrus) species, males genitalia. 9–10, 22–23, L. inermis Reitter, 1897 (9–10—20 km W Oqmachit, 22–23—Pashtamazar); 11–12, 24, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 13–14, 25–27, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin); 15–16, 28, L. microbuccis Ballion, 1871 (Golbo env.); 15–16, 28, L. microbuccis Ballion, 1871 (Golbo env.); 17–18, 29, L. elisae Nikolajev, 2001 (Shurob); 19–21, 30–31, L. dyonini A. Shapovalov & Bagaturov, new species, paratypes (environs of Tashsay). 9–21 aedeagus (9, 11, 13, 15, 17, 19–20—ventral view; 10, 12, 14, 16, 18, 21—lateral view), 22–31 sclerotized apex of endophallus (22, 24, 25, 28–30 dorsal view, 23, 26, 31—lateral view, 27—ventral view)." pageId="1" pageNumber="436">Figs. 22–23</figureCitation> | ||
). Dorsal surface of the body shiny ( | ||
<figureCitation id="2EF82A49FFCEFF984D0AF982FBE0F9E6" box="[1038,1145,1586,1612]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1761,1785]" captionTargetBox="[274,1320,197,1720]" captionTargetId="figure-17@3.[264,1323,179,1735]" captionTargetPageId="3" captionText="FIGURES 1–4. Lethrus (Mesolethrus) species, male, habitus. 1–2, L. inermis Reitter, 1897 (1—Dahanakiik; 2—20 km W Oqmachit); 3, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 4, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin)." pageId="1" pageNumber="436">Figs. 1–2</figureCitation> | ||
), but in older and abraded specimens surface nearly black or black. | ||
</paragraph> | ||
<paragraph id="B67C36CCFFCEFF9849C3F9CAFC1BF93E" blockId="1.[151,1437,1405,1684]" box="[199,898,1658,1684]" pageId="1" pageNumber="436"> | ||
<emphasis id="84B7EADEFFCEFF9849C3F9CAFE88F93E" bold="true" box="[199,273,1658,1684]" pageId="1" pageNumber="436">Notes.</emphasis> | ||
The species group includes the only species | ||
<emphasis id="84B7EADEFFCEFF984A0CF9CBFC1BF93E" box="[776,898,1659,1684]" italics="true" pageId="1" pageNumber="436"> | ||
<taxonomicName id="71C34D4FFFCEFF984A0CF9CBFCE7F93E" authorityName="Reitter" authorityYear="1897" box="[776,894,1659,1684]" class="Insecta" family="Geotrupidae" genus="Lethrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="436" phylum="Arthropoda" rank="species" species="inermis">L. inermis</taxonomicName> | ||
. | ||
</emphasis> | ||
</paragraph> | ||
</subSubSection> | ||
</treatment> | ||
</document> |
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