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This error is probably caused by very long sequences in the input. In v5.1, muscle reports the mean and maximum sequence lengths when starting and issues a warning if long sequences are found.
muscle 5.0.1428_win64 34.1Gb RAM, 16 cores
(C) Copyright 2004-2021 Robert C. Edgar.
https://drive5.com
00:00 5.9Mb 0.037% Calc posteriors
D:\MyApp\bin\muscle.exe -align .\all.fasta -output all2.fasta
Elapsed time 00:04
Max memory 5.9Mb
---Fatal error---
myalloc64(4, 1.1e+09) failed
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