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The Content-Type header was not being correctly handled. If it wasn't sent exactly as "CONTENT-TYPE" it was getting recorded in the environ as "HTTP_CONTENT_TYPE" which is against PEP-333.
In the pure-Python parser, the SCRIPT_NAME field was being mishandled due to mutation of the original environ. The code was looking for HTTP_SCRIPT_NAME which was already popped from the environ dictionary if it existed.