The exclusion section:
[ exclusions ]
1 2 3 4
2 1 3 4
3 1 2 4
4 1 2 3
Is being written to DESMOND as:
ffio_exclusions[2] {
i_ffio_ai
i_ffio_aj
:::
1 1 2
2 1 3
:::
}
It is being written correctly to GROMACS, so it looks like a conversion problem to DESMOND.