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A joint trait can jump at the split as well as along branches.traits.discrete(at_speciation=…)
already worked inside simulate_joint, so a trait could drive speciation and change at each
speciation — but nothing said so, nothing tested it, and Chapter 9 documented only the along-branch
case, which made the combination look unavailable. It is now documented, with an example, and pinned
by tests: a jump at a split is logged as on_speciation, a change along a branch as on_branch,
so the two can be told apart when scoring a method. (#256)